Tethysaurus Temporal range: Late Cretaceous, | |
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Tethysaurus skeleton | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Squamata |
Superfamily: | † Mosasauroidea |
Family: | † Mosasauridae |
Subfamily: | † Tethysaurinae |
Genus: | † Tethysaurus Bardet, Pereda-Suberbiola & Jalil, 2003 |
Type species | |
†Tethysaurus nopcsai Bardet, Pereda-Suberbiola & Jalil, 2003 |
Tethysaurus is an extinct genus of tethysaurine mosasauroid from the Early Turonian (Late Cretaceous) period. The only species is Tethysaurus nopcsai.
The name means "Tethys' lizard of Nopcsa", a reference to the Greek goddess of the sea Tethys (also the name of the Tethys Ocean, an ancient sea between southern Europe and northern Africa) and to the Hungarian paleontologist Baron Ferenc Nopcsa, who made pioneering studies on Adriatic aquatic squamates. It was found in the Akrabou Formation, near the villages of Tadirhourst and Asfla in the region of Goulmima, Errachidia Province, in Morocco, with three referred specimens that included a nearly complete articulated skull, mandible, vertebrae and portions of the appendicular skeleton. The diagnosis after Bardet et al. is "(...) prefrontal strongly vaulted in anterior view; parietal exhibits a triangular table ending posteriorly in two pointed pegs overlying the supraoccipital; jugal with a large and wide ascending ramus; medullar floor of the basioccipital pierced by three foramina; splenial with a large notched dorsomedial process; surangular exposed medially ventral to the coronoid; dental formula: 19-20 maxillary, 15-19 pterygoid and at least 19 dentary teeth; large paracotylar and parazygosphenal foramina on vertebrae." [1]
A mosasaur measuring under 3 m (9.8 ft) long, [1] Tethysaurus displays a number of basal and derived features that led to an initial classification as an intermediate stage between primitive aigialosaurids of the Cenomanian and derived mosasaurids from the Turonian to the Maastrichtian. [1] More recent analysis put Tethysaurus in a clade along Russellosaurus and Yaguarasaurus called the parafamily Russellosaurina, as a basal Turonian clade of Mosasauridae. [2]
Cladogram based in the analysis by Makádi et al. in 2012: [3]
Tethysaurinae |
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Mosasaurs comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes.
The Mosasaurinae are a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "mosasaurines" and their fossils have been recovered from every continent except for South America.
Plioplatecarpinae is a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "plioplatecarpines" and have been recovered from all continents, though the occurrences in Australia remain questionable. The subfamily includes the genera Latoplatecarpus, Platecarpus, Plioplatecarpus and Plesioplatecarpus.
The Halisaurinae are a subfamily of mosasaurs, a group of Late Cretaceous marine lizards. They were small to medium-sized, ranging from just under 3 meters in Eonatator sternbergi to as much as 8 or 9 meters in Pluridens serpentis. They tended to have relatively slender jaws and small, numerous teeth, suggesting a diet of small fish and other prey. Although the skeleton is primitive compared to other Mosasauridae in many respects, halisaurines had the distinctive hypocercal tail of other mosasaurids suggesting good swimming ability, and they persisted alongside other mosasaurs until the end of the Cretaceous. The earliest known remains of halisaurines occur in rocks of Santonian age and the subfamily persists until the latest Maastrichtian. Halisaurines are known from North and South America, Europe, Asia and Africa, indicating a more or less global distribution in the Late Cretaceous. Four genera are currently recognized: Eonatator, Halisaurus, Phosphorosaurus and Pluridens.
Eonatator is an extinct genus of marine lizard belonging to the mosasaur family. It is a close relative of Halisaurus, and part of the same subfamily, the Halisaurinae. It is known from the Late Cretaceous of North America, Colombia and Sweden. Originally, this taxon was included within Halisaurus, but was placed in its own genus, which also led to the subfamily Halisaurinae being created for the two genera.
Russellosaurus is an extinct genus of tethysaurine mosasauroid from the Late Cretaceous of North America. The genus was described from a skull discovered in an exposure of the Arcadia Park Shale at Cedar Hill, Dallas County in the south-central part of the DFW Metroplex in Texas, United States. The skull was found in 1992 by a member of the Dallas Paleontological Society, who then donated to the museum. Other fragmentary specimens of Russelosaurus have been recovered from the slightly older Kamp Ranch Limestone at two other localities in the Dallas area.
Aigialosauridae is a family of Late Cretaceous semiaquatic pythonomorph lizards closely related to the mosasaurs. Regarded by some paleontologists as a distinct monophyletic group and by others as an adaptive grade within the basal mosasauroids, recent molecular and morphological data suggests that they are the oldest known members of the lineage leading to the mosasaurs.
Dallasaurus is a basal mosasauroid from the Upper Cretaceous of North America. Along with Russellosaurus, Dallasaurus is one of the two oldest mosasauroid taxa currently known from North America. It is also one of the smallest known mosasaurines, measuring approximately 1 m (3.3 ft) in length.
Goronyosaurus is an extinct genus of marine lizard belonging to the mosasaur family. Fossils of Goronyosaurus are exclusively known from the Late Maastrichtian of the Iullemmeden Basin in West Africa, specifically the Dukamaje Formation of Niger and Nigeria and Farin Doutchi Formation of Niger. The type specimen was first described in 1930 as Mosasaurus nigeriensis, but subsequent remains revealed a highly unique set of adaptations that prompted the species to be reclassified as the only species of the new genus Goronyosaurus in 1972. These unique adaptations have made Goronyosaurus notoriously difficult to classify within the Mosasauridae and it is often left out of phylogenetic analyses, although most authors agree that Goronyosaurus belonged to Mosasauridae.
Yaguarasaurus is an extinct genus of mosasauroid from the Late Cretaceous (Turonian) period of Colombia, South America. The remains discovered were defined as a new genus and species of mosasaurid, Yaguarasaurus columbianus, by the Colombian paleontologist María Páramo, former director of the Museo de Geología José Royo y Gómez of INGEOMINAS in Bogotá. The first fossils remains of this animal suggested a cranial length of 47 centimetres (19 in) and a total length of 5 metres (16 ft); an additional skull that measures 87 centimetres (34 in) long implies a larger size.
Angolasaurus is an extinct genus of mosasaur. Definite remains from this genus have been recovered from the Turonian and Coniacian of Angola, and possibly the Coniacian of the United States, the Turonian of Brazil, and the Maastrichtian of Niger. While at one point considered a species of Platecarpus, recent phylogenetic analyses have placed it between the (then) plioplatecarpines Ectenosaurus and Selmasaurus, maintaining a basal position within the plioplatecarpinae.
Acteosaurus is an extinct genus of aquatic lizard that lived in the upper Cretaceous period. Its species, A. tommasinii and A. crassicostatus, were described in 1860 and 1993. Though A. crassicostatus is probably a junior synonym for Adriosaurus suessi, A. tommasinii was found to be similar to coniasaurs, mosasauroids, and a sister taxon to modern snakes in 2010.
Eremiasaurus is a genus of mosasaurs, an extinct group of marine reptiles. It lived during the Maastrichtian stage of the Late Cretaceous in what is now North Africa. Only one species is known, E. heterodontus, described in 2012 from two remarkably complete fossil specimens discovered in the Ouled Abdoun Basin, Morocco. This site is known to have delivered a significant number of other related mosasaurs.
Pannoniasaurus is an extinct genus of tethysaurine mosasauroid known from the Late Cretaceous Csehbánya Formation of Hungary. It contains a single species, Pannoniasaurus inexpectatus, dubbed "unexpected" because it was discovered in freshwater sediments, unlike other mosasaurs, which were marine predators. It was a medium-sized mosasaur, reaching up to 6 metres (20 ft) in length.
The Tethysaurinae are a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "tethysaurines" and have been recovered from North America and Africa. Only two tethysaurine genera are known, Pannoniasaurus and Tethysaurus. The genera Yaguarasaurus and Russellosaurus were previously considered tethysaurines until they were grouped with Romeosaurus in the new subfamily Yaguarasaurinae. A possible member of this clade (subfamily) is a mosasaur specimen known from a maxilla fragment, found in 1960 in the Czech Republic, in Dolní Újezd near Litomyšl.
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Goulmimichthys is an extinct genus of ray-finned fishes in the family Pachyrhizodontidae. The genus, first described by Cavin in 1995, is known from various Turonian age formations. The type species G. arambourgi from the Akrabou Formation in the El Rachidia Province of Morocco, and other fossils described are G. gasparini of the La Frontera Formation, Colombia, and G. roberti from the Agua Nueva Formation of Mexico.
Thalassotitan is an extinct genus of large mosasaurs that lived during the late Maastrichtian of the Cretaceous period in what is now Morocco, around 66 million years ago. The only known species is T. atrox, described in 2022 from fossils discovered in the Ouled Abdoun Basin, where many other mosasaurs have been found. It was assigned to the tribe Prognathodontini alongside other mosasaurs like Prognathodon and Gnathomortis. The prognathodontines are separated from other mosasaurs based on their massive jaws and robust teeth.