Tille trevally

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Tille trevally
Tille trevally Honiara.jpeg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Carangiformes
Family: Carangidae
Genus: Caranx
Species:
C. tille
Binomial name
Caranx tille
G. Cuvier, 1833
Caranx tille distribution.png
Approximate range of the tille trevally
Synonyms [2]

Caranx cynodon Bleeker, 1851

The tille trevally (Caranx tille), also known as the tille kingfish, is a species of large marine fish classified in the jack family, Carangidae. The tille trevally is distributed through the tropical and subtropical waters of the Indo-Pacific region, ranging from South Africa in the west to Fiji, Japan and Australia in the east. The species is best distinguished by its rounded, strongly convex anterior profile, with other detailed anatomical features also useful. The tille trevally reaches a maximum length of 80 cm and a weight of 7.2 kg. It is predominantly an inshore species, inhabiting coastal reef and lagoon environments, although has been recorded on deep offshore seamounts. It is a predatory fish, taking various species of fish and crustaceans as prey, with little known of its reproductive cycle. It is of minor importance to fisheries throughout its range, taken by hook and line, gill nets and purse seines. The tille trevally is also considered a good game fish, and an excellent table fish. The species acquired its scientific and common names from a local name used by Pondicherry fishermen, koton tille, which Georges Cuvier then used when he named the species in 1833.

Contents

Taxonomy and naming

The tille trevally is classified within the genus Caranx , one of a number of groups known as the jacks or trevallies. Caranx itself is part of the larger jack and horse mackerel family Carangidae, which in turn is part of the order Carangiformes. [3]

The species was first scientifically described by Georges Cuvier in 1833 based on the holotype specimen collected from Pondicherry, India. [4] He named the species Caranx tille, with the specific epithet derived from a local name used by fishermen for the species in Pondicherry, koton tille. [5] Cuvier noted that Jean Baptiste Leschenault, who collected the species, expressed doubt over the validity of the new species despite local fishermen identifying it as separate to other trevallies. [5] The species was independently redescribed three times; the first was by Pieter Bleeker in 1851 as Caranx cynodon, the second time by Henry Weed Fowler in 1904 as Caranx semisomnus, and lastly as Caranx auriga by Alvin Seale in 1910. [6] The latter name caused a specific homonym with Caranx auriga, a name proposed for a different species by Charles Walter De Vis in 1884. To rectify this, William Ogilby proposed the name Citula virga as a replacement, while Roxas and Martin proposed Caranx manilensis. All these names were soon recognized as synonymous with Caranx tille, and are now considered invalid junior synonyms under ICZN rules. [6]

Description

The tille trevally is a large species, growing to a maximum known length of 80 cm and a recorded weight of 7.3 kg. [2] It is similar in general appearance to most jacks in the genus, having a compressed, oblong body, with the dorsal profile more convex than the ventral profile, particularly anteriorly. This curvature gives the fish a discernibly 'blunt' head and steep snout, which is the most obvious identifying feature of the species. [7] The dorsal fin is in two distinct sections; the first consists of eight spine and the second of one spine and 20 to 22 soft rays. The anal fin consists of two anteriorly detached spines followed by one spine and 16 to 18 soft rays, [8] while the pelvic fins have one spine followed by 18 soft rays. The lateral line is moderately arched anteriorly, with 53 to 54 scales in this section, while the straight section contains none to two scales and 33 to 42 strong scutes. The breast is completely scaled. [9] The species has well-developed adipose eyelids, while its dentition consists of an outer row of widely spaced canines and an inner band of villiform teeth in the upper jaw, with a row of widely spaced conical teeth on the lower jaw. The tille trevally has 22 to 25 gill rakers and 24 vertebrae. [7]

The tille trevally is generally a uniformly pale olive to silvery grey colour all over as a juvenile, with the fins being pale to dusky. As the fish becomes larger, the head and upper body become a darker olive green to bluish-grey colour, with the underside fading to a silvery white. The second dorsal fin is olive to black and lacks the white tip shown by Caranx papuensis , while the caudal and anal fins are yellow-olive to black. The species has a black spot on the upper opercular margin. [7] [8]

Distribution and habitat

The smaller eye and colourless fins help distinguish the tille trevally from the bigeye trevally Tille trevally Palau.jpg
The smaller eye and colourless fins help distinguish the tille trevally from the bigeye trevally

The tille trevally is distributed through the tropical and subtropical waters of the Indian and west Pacific Oceans. [7] In the western part of its range, the species distributed throughout South Africa and Madagascar waters north along the east African coast up to Tanzania, with an apparent break in its range from Tanzania to India. Its range continues from India east to South East Asia and the Indonesian Archipelago. The distribution extends south to northern Australia, north to Japan, and to Fiji in the east. [2]

The tille trevally is typically an inshore species, preferentially inhabiting coastal coral and rocky reefs, [7] as well as moving through sandy bays and lagoons. The species is known to move with the tide in the Solomon Islands, entering inner lagoon regions through tidal corridors as the tide rises, and exiting back to the outer reef as the tide falls. This is thought to be due to the fish following small prey fish into the lagoon, where these small species spawn. [10] The species is also known to enter estuarine waters and the lower reaches of rivers. [11] Despite being more prevalent in inshore waters, the tille trevally has also been recorded in far offshore waters, living on seamounts in West Mariana Ridge of the Pacific, where seamount peaks reach to 50 m from the surface. Here it occurs with other normally inshore fishes, and is one of the top predators in this ecosystem. [12]

Biology and fishery

The tille trevally's biology is poorly understood, with only basic aspects of its diet known. It is a predatory species which takes other fish and various crustaceans as prey. [2] Known hunting movements have been described above, with it moving into shallow lagoons to hunt small baitfish which shelter and spawn in the mangroves at high tide. [10] Reproduction and growth parameters are entirely unknown.[ citation needed ]

The tille trevally is of minor importance in most fisheries throughout its range, although it is locally important in some areas. One such fishery is in Wewak, Papua New Guinea, where it and three lutjanids made up 50% of the catch, with C. tille the predominant carangid in the fishery. [13] It is taken by hook and line, gill nets, purse seines and various artisanal gear. [7] According to Cuvier, the species is an excellent table fish, [5] and is also considered a good gamefish in larger sizes. [7]

Related Research Articles

<span class="mw-page-title-main">Crevalle jack</span> Species of fish

The crevalle jack, also known as the common jack, black-tailed trevally, couvalli jack, black cavalli, jack crevale, or yellow cavalli is a common species of large marine fish classified within the jack family, Carangidae. The crevalle jack is distributed across the tropical and temperate waters of the Atlantic Ocean, ranging from Nova Scotia, Canada to Uruguay in the western Atlantic and Portugal to Angola in the eastern Atlantic, including the Mediterranean Sea. It is distinguishable from similar species by its deep body, fin colouration and a host of more detailed anatomical features, including fin ray and lateral line scale counts. It is one of the largest fish in the genus Caranx, growing to a maximum known length of 124 cm and a weight of 32 kg, although is rare at lengths greater than 60 cm. The crevalle jack inhabits both inshore and offshore waters to depths of around 350 m, predominantly over reefs, bays, lagoons and occasionally estuaries. Young fish dispersed north by currents in the eastern Atlantic are known to migrate back to more tropical waters before the onset of winter; however, if the fish fail to migrate, mass mortalities occur as the temperature falls below the species' tolerance.

<span class="mw-page-title-main">Bigeye trevally</span> Species of fish

The bigeye trevally, also known as the bigeye jack, great trevally, six-banded trevally and dusky jack, is a species of widespread large marine fish classified in the jack family Carangidae. The bigeye trevally is distributed throughout the tropical waters of the Indian and Pacific Oceans, ranging from South Africa in the west to California and Ecuador in the east, including Australia to the south and Japan in the north. The bigeye trevally is best distinguished by its colouration, having a dark second dorsal fin with a white tip on the lobe, and also possessing a small dark spot on the operculum. Other more detailed anatomical features also set the species apart from other members of Caranx. The species is known to grow to a length of 120 cm and 18 kg.

<span class="mw-page-title-main">Bluefin trevally</span> Species of fish

The bluefin trevally, also known as the bluefin jack, bluefin kingfish, bluefinned crevalle, blue ulua, omilu, and spotted trevally), is a species of large, widely distributed marine fish classified in the jack family, Carangidae. The bluefin trevally is distributed throughout the tropical waters of the Indian and Pacific Oceans, ranging from Eastern Africa in the west to Central America in the east, including Japan in the north and Australia in the south. The species grows to a maximum known length of 117 centimetres (46 in) and a weight of 43.5 kilograms (96 lb), however is rare above 80 centimetres (31 in). Bluefin trevally are easily recognised by their electric blue fins, tapered snout and numerous blue and black spots on their sides. Juveniles lack these obvious colours, and must be identified by more detailed anatomical features such as fin ray and scute counts. The bluefin trevally inhabits both inshore environments such as bays, lagoons and shallow reefs, as well as deeper offshore reefs, atolls and bomboras. Juveniles prefer shallower, protected waters, even entering estuaries for short periods in some locations.

<span class="mw-page-title-main">Orange-spotted trevally</span> Species of fish

The orange-spotted trevally, Carangoides bajad is a species of inshore marine fish in the jack family, Carangidae. The species is fairly common in tropical to subtropical waters of the Indo-Pacific, ranging from Madagascar in the west to Japan in the east, typically inhabiting inshore reefs. The species has characteristic orange-yellow spots on its sides, although counts of fin rays and scutes are needed to distinguish it from related species with similar colouring. Orange-spotted trevallies are powerful predators, taking a variety of small fish, nekton, and crustaceans, and reach sexual maturity around 25 cm long. It is a moderately large fish, reaching a maximum known length of 55 cm. The species is occasionally taken by fishermen throughout its range, and is generally considered to be bycatch. The exception to this is in the southern Persian Gulf, where it makes up a large proportion of the fishery.

<span class="mw-page-title-main">Shrimp scad</span> Species of fish

The shrimp scad, is a species of widespread tropical marine fish of the jack family, Carangidae. The shrimp scad is widely distributed in the tropical and subtropical western Indian Ocean and areas of the eastern Pacific Ocean, ranging from South Africa in the west to Hawaii in the east, including Japan and Australia to the north and south. The species is commonly found on inshore reefs and sandy substrates. It has the common body profile of a scad, and may be difficult to differentiate from others in the genus Alepes. It is one of the larger scads, growing to 40 cm, but often is encountered at much smaller sizes. The shrimp scad often forms large schools, and is carnivorous, consuming a variety of crustaceans and small fish. It is of moderate importance to fisheries throughout its range.

<i>Caranx lugubris</i> Species of fish

Caranx lugubris, the black jack, black trevally, black kingfish, coal fish or black ulua, is a species of large ocean fish in the jack family Carangidae. The species has a circumtropical distribution, found in oceanic, offshore waters of the tropical zones of the Pacific, Atlantic and Indian Oceans. The species is particularly prevalent around offshore islands such as the Caribbean islands in the Atlantic, Hawaii and French Polynesia in the Pacific and the Seychelles and Maldives in the Indian Ocean. Black jack are rare in shallow waters, preferring deep reefs, ledges and seamounts in clear waters. The species is easily distinguished by its black to grey fins and jet black scutes, with the head having a steep profile near the snout. The largest recorded length is 1 m and weight of 17.9 kg. The black jack lives either individually or in small schools, and is known to school with other species. It is a predatory fish, taking a variety of fish, crustaceans and molluscs as prey. Sexual maturity is reached at 34.6 cm in females and 38.2 cm in males, with spawning taking place between February and September in the Caribbean. The early life history of the species is very poorly understood. Black jack are of high importance to many island fisheries, but are rarely encountered in most continental fisheries. The species has a reputation as a gamefish, and is variably considered a terrible or excellent food fish, although several cases of ciguatera poisoning have been attributed to the species. The species was initially named Caranx ascensionis by Georges Cuvier, however several issues with the use of this name have seen Felipe Poey's name Caranx lugubris become the valid scientific name.

<span class="mw-page-title-main">Longfin trevally</span> Species of fish

The longfin trevally, also known as the longfin kingfish, longfin cavalla or armed trevally, is a species of inshore marine fish in the jack family, Carangidae. The species is common in tropical to subtropical waters of the Indo-Pacific, ranging from South Africa in the west to Japan in the east, typically inhabiting inshore reefs and bays. The species is easily distinguished by its elongate dorsal and anal fin lobes and filamentous dorsal rays, as well as its scaleless breast. Longfin trevally are pelagic predators, taking a variety of small fish, cephalopods and crustaceans, and reach sexual maturity at around 21 cm. The species has a maximum known length of 57 cm and weight of 3.5 kg. The longfin trevally has a very complex taxonomic history which is closely intertwined with another currently valid species, Carangoides ciliarius, which may yet prove to be synonymous. Longfin trevally are of minor importance to fisheries throughout their range and are considered good table fish, and are occasionally taken by anglers.

<span class="mw-page-title-main">Longnose trevally</span> Species of fish

The longnose trevally, also known as the tea-leaf trevally, club-nosed trevally, grunting trevally or dusky trevally, is a species of inshore marine fish in the jack family, Carangidae. The species is distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans from South Africa to New Zealand and Japan, inhabiting coastal waters, especially reefs, to a depth of 90 m. The longnose trevally is distinguished from similar species by a combination of a scaleless breast and the number of gill rakers and fin rays. It is a moderately large fish, growing to a maximum known length of 72 cm and 4.35 kg. The longnose trevally is a predatory fish, consuming small fish, crustaceans and molluscs. The species is of minor commercial importance throughout its range, and is considered to be a good table fish.

<span class="mw-page-title-main">Coastal trevally</span> Species of fish

The coastal trevally, also known as the onion trevally, Japanese trevally or bluefin kingfish, is a species of inshore marine fish in the jack family Carangidae. The species is distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans, from South Africa in the west to Japan and New Caledonia in the east, reaching as far south as Australia. The species is found on deep coastal reefs, both in schools and as solitary individuals, where they prey on small midwater organisms including crustaceans, small fish and cephalopods. The species is taken as bycatch in a number of fisheries throughout its range by a number of fishing methods and is of little commercial value, but is considered to be a good table fish. A mistype in the original volume in which Eduard Rüppell named the species led to the combination Carangoides caeruleopinnatus, which has incorrectly spread through the literature.

<span class="mw-page-title-main">Green jack</span> Species of fish

The green jack, also known as the horse jack, is an abundant species of coastal marine fish in the jack family Carangidae. The species is distributed in the eastern Pacific Ocean along the American coastline from Santa Cruz Island off California in the north to Peru in the south, as well as a number of islands including the Galapagos and recently, Hawaii. The green jack is distinguished from other similar carangid species by a number of features including gill raker and lateral line scale counts, and the presence of an adipose eyelid. It is a moderately large species, growing to at least 55 cm in length and 2.81 kg, although unconfirmed reports suggest a much larger maximum size. It lives in a wide range of continental shelf environments including estuaries, bays, reefs and offshore seamounts, living both pelagically and demersally. The green jack is a predatory species, preying on a variety of fish, crustaceans and cephalopods, as well as zooplankton. Most fish are sexually mature by the time they attain 38 cm, with spawning occurring between May and October. The species is of high importance to fisheries throughout its range, caught by pelagic trawls, a variety of netting methods and hook and line. The green jack is also of interest to anglers, taken by bait and lures, although is considered to only be fair in eating quality.

<i>Caranx</i> Genus of fishes

Caranx is a genus of tropical to subtropical marine fishes in the jack family Carangidae, commonly known as jacks, trevallies and kingfishes. They are moderate- to large-sized, deep-bodied fishes which are distinguished from other carangid genera by specific gill raker, fin ray and dentition characteristics. The genus is represented in the Pacific, Indian and Atlantic Oceans, inhabiting both inshore and offshore regions, ranging from estuaries and bays to deep reefs and offshore islands. All species are powerful predators, taking a variety of fish, crustaceans and cephalopods, while they in turn are prey to larger pelagic fishes and sharks. A number of fish in the genus have a reputation as powerful gamefish and are highly sought by anglers. They often make up high amounts of the catch in various fisheries, but are generally considered poor to fair table fishes.

<span class="mw-page-title-main">Yellowspotted trevally</span> Species of fish

The yellowspotted trevally, also known as the yellowspotted kingfish, goldspotted trevally, tarrum, or yellowspot, is a widespread species of large inshore marine fish in the jack family Carangidae. The yellowspotted trevally inhabits the tropical and subtropical waters of the western Indo-Pacific region, from South Africa in the west to Japan and Australia in the east. The species is known to grow to a maximum length of at least 1.2 m, and is distinguished by gill raker and fin morphology, as well as the distinctive golden spots which give the fish its name. The yellowspotted trevally generally prefers inshore rocky and coral reefs, but is occasionally found over deep offshore sand banks to a depth of 100 m. It is a predatory fish, taking fish, cephalopods, and crustaceans, and shows diet partitioning with other trevallies in studies conducted in Australian waters. Reproduction is poorly studied, although observational evidence suggests spawning occurs in aggregations, probably during summer in South Africa. It is generally of minor importance to commercial fisheries throughout its range, but is considered an excellent sportfish by anglers and spearfishermen, and a good table fish.

<span class="mw-page-title-main">Bumpnose trevally</span> Species of fish

The bumpnose trevally, also known as the bumpnose kingfish or onion kingfish, is a species of relatively small inshore marine fish classified in the jack family Carangidae. The bumpnose trevally is fairly common in the tropical and subtropical waters of the Indo-west Pacific region, ranging from South Africa in the west to Japan and Samoa in the east. It is a small species by carangid standards, reaching a maximum known length of 32 cm, and can be distinguished from the similarly shaped Carangoides armatus by a distinct 'bump' on the snout, which gives the fish its common name. The species inhabits coastal waters, often living along bays and beaches, where it takes shrimp, small crabs, and juvenile fish as prey. The bumpnose trevally is of minor importance to fisheries throughout its range, taken by hook and line, trawls, and seine nets. It is also of minor importance to anglers, taken by baits from beaches and piers, and is considered a modest table fish.

<span class="mw-page-title-main">Coachwhip trevally</span> Species of fish

The coachwhip trevally, also known as the oblong trevally or oblique-banded trevally, is a species of inshore marine fish classified in the jack family Carangidae. The coachwhip trevally is distributed through the Indo-west Pacific region, ranging from South Africa in the west to Fiji and Japan in the east. It is a moderately large fish, growing to a known maximum length of 46 cm and can be distinguished from similar species by an array of detailed morphological features including dentition, fin ray counts and scale patterns. The coachwhip trevally inhabits coastal waters throughout its range, known to prefer estuarine waters in a number of localities. Nothing is known of its diet or reproductive biology, and is of little importance to fisheries, occasionally taken as bycatch in trawl and hook and line fisheries.

<span class="mw-page-title-main">Barcheek trevally</span> Species of fish

The barcheek trevally, also known as the barcheek kingfish, shortridge trevally or oblique-banded trevally, is a species of moderately large marine fish of the jack family Carangidae. The barcheek trevally is distributed throughout the tropical waters of the Indo-west Pacific region, ranging from South Africa in the west to Japan, Australia and a number of small central Pacific islands in the east. The species inhabits inshore and offshore waters, found along the slopes of lagoons and out to deeper reefs on the continental shelf, where it preys on small fish and benthic crustaceans. It is a moderately large fish, growing to a maximum recorded length of 50 cm, and can be distinguished from similar species by its somewhat protruding lower jaw and the dark banding on its operculum. It is of minor importance to fisheries throughout its range, taken by trawling, hook and line methods and various inshore fish netting methods.

<span class="mw-page-title-main">Brownback trevally</span> Species of fish

The brownback trevally, also known as the brown-backed trevally, is a species of small inshore marine fish classified in the jack family, Carangidae. The brownback trevally is distributed in two populations through the tropical waters of the Indo-west Pacific region, ranging from the Persian Gulf east to India, South East Asia and the Indonesian islands. The species is distinguished from similar species by its completely scaled breast and black-tipped second dorsal fin, and is known to reach a maximum length of 25 cm. The brownback trevally inhabits inshore waters including bays and estuaries, where it preys on demersal crustaceans and small fish. Other aspects of its biology are poorly known, and it is of minor importance to fisheries, occasionally caught by hook and line or trawls. William Smith-Vaniz has recently suggested the two distinct populations may actually represent two distinct species.

<span class="mw-page-title-main">Blacktip trevally</span> Species of fish

The blacktip trevally, also known as the blacktip kingfish or yellowtail kingfish, is a species of large marine fish classified in the jack family Carangidae. The blacktip trevally is distributed throughout the tropical to subtropical Indian and West Pacific Oceans, ranging from South Africa in the west to Fiji, Japan and northern Australia in the east. It inhabits coastal waters throughout its range, preferring moderately deep clear waters over rocky and coral reefs. The blacktip trevally is easily distinguished by its yellow fins and a dark upper caudal fin lobe which gives the species its common name, as well as a host of other anatomical features. The species is known to reach a maximum size of 1 m. It is a benthopelagic predator, commonly forming small shoals where it takes a variety of fishes, cephalopods and crustaceans as prey. Little is known of reproduction in the species, and spawning is assumed to take place in more tropical regions of its range, with juveniles known to inhabit bays and large estuaries. Blacktip trevally are often caught using hook and line and various nets in commercial fisheries although don't make up a large part of the market. They are also popular with anglers due to their fighting ability and decent table qualities.

<span class="mw-page-title-main">Brassy trevally</span> Species of fish

The brassy trevally, Caranx papuensis is a species of large marine fish classified in the jack family, Carangidae.

<span class="mw-page-title-main">Fringefin trevally</span> Species of fish

The fringefin trevally, also called fringe-finned trevally, round-finned trevally or reef herring, is a species of inshore marine fish classified in the jack and horse mackerel family Carangidae. A relatively small fish, the fringefin trevally is known to reach 40 cm (16 in), but is mostly encountered at lengths less than 25 cm (9.8 in). The fringefin trevally has an ovate body, with distinctive orange-yellow fins and a black opercular spot. The dental patterns of the species distinguish it from the closely related scads of the genus Alepes. Males have characteristic elongated dorsal and anal fins which produce a series of trailing filaments. The fringefin trevally is restricted to the waters of the Indo-Pacific, ranging from northern Australia to Papua New Guinea and eastern Indonesia. An inshore species, it is found in coastal and estuarine environments and exhibits daily and seasonal movements. The fringefin trevally is predatory, taking crustaceans as prey. The species is often taken as bycatch in prawn trawls and occasionally taken by anglers.

<span class="mw-page-title-main">Longrakered trevally</span> Species of fish

The longrakered trevally, also known as the cale cale trevally and heavyjawed kingfish, is a species of marine fish in the jack and horse mackerel family Carangidae. The longrakered trevally is distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans, from Mozambique and Madagascar in the west, to Japan and northern Australia in the east. A large species growing to a recorded length of 1 m, the longrakered trevally is distinguished by is protruding lower jaw, elongated gill rakers and lack of villiform teeth on its tongue. It is an inshore species, restricted to coastal and estuarine regions, where it preys on fishes and crustaceans. Little is known of the species reproductive cycle or growth. The longrakered trevally is of minor importance to fisheries and is often taken as bycatch in finfish and prawn trawls, as well as by recreational fishermen.

References

  1. Smith-Vaniz, W.F. & Williams I. (2017) [errata version of 2016 assessment]. "Caranx tille". IUCN Red List of Threatened Species . 2016: e.T20430800A115378876. doi: 10.2305/IUCN.UK.2016-3.RLTS.T20430800A65927858.en .
  2. 1 2 3 4 Froese, Rainer; Pauly, Daniel (eds.) (2019). "Caranx tille" in FishBase . August 2019 version.
  3. J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016). Fishes of the World (5th ed.). Wiley. pp. 380–387. ISBN   978-1-118-34233-6.
  4. Hosese, D.F.; Bray, D.J.; Paxton, J.R.; Alen, G.R. (2007). Zoological Catalogue of Australia Vol. 35 (2) Fishes. Sydney: CSIRO. p. 1150. ISBN   978-0-643-09334-8.
  5. 1 2 3 Cuvier, G.; A. Valenciennes (1849). Histoire naturelle des poissons. F.G. Levrault. pp. IX, 93.
  6. 1 2 California Academy of Sciences: Ichthyology (April 2009). "Caranx tille". Catalog of Fishes. CAS. Retrieved 2009-05-16.
  7. 1 2 3 4 5 6 7 Smith-Vaniz, W. (1999). "Carangidae" (PDF). In Carpenter, K.E.; Niem, V.H. (eds.). The living marine resources of the Western Central Pacific Vol 4. Bony fishes part 2 (Mugilidae to Carangidae). FAO species identification guide for fishery purposes. Rome: FAO. pp. 2659–2757. ISBN   92-5-104301-9.
  8. 1 2 Randall, John Ernest; Roger C. Steene; Gerald R. Allen (1997). Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press. p. 161. ISBN   0-8248-1895-4.
  9. Lin, Pai-Lei; Shao, Kwang-Tsao (1999). "A Review of the Carangid Fishes (Family Carangidae) From Taiwan with Descriptions of Four New Records". Zoological Studies. 38 (1): 33–68.
  10. 1 2 Hamilton, R.; R. Walter (1999). "Indigenous ecological knowledge and its role in fisheries research design: A case study from Roviana Lagoon, Western Province, Solomon Islands" (PDF). SPC Traditional Marine Resource Management and Knowledge Information Bulletin. 11: 13–25. Archived from the original (PDF) on 2011-06-11. Retrieved 2009-05-21.
  11. Boseto, D.; A.P. Jenkins (2006). A checklist of freshwater and brackish water fishes of the Fiji Islands (PDF). University of the South Pacific, Institute of Applied Sciences. pp. 1–9. Archived from the original (PDF) on 2016-03-03.
  12. Miller, M.J.; T. Inagaki; J. Aoyama; K. Yoshizumi; T. Ajinomoto; K. Tsukamoto (2004). "Ecology of Ichthyofauna at Seamounts: Unseen Islands of Biodiversity in the Deep Ocean" (PDF). Ocean Research Institute, The University of Tokyo. Retrieved 2009-05-21.[ dead link ]
  13. Chapau, M.R. (1988). "A review of deepwater handline fishing in Papua New Guinea" (PDF). SPC Inshore Fisheries Research. BP-82: 1–7. Retrieved 2009-05-21.