Aulacomnium palustre

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Aulacomnium palustre
Aulacomnium palustre.jpeg
Scientific classification Red Pencil Icon.png
Kingdom: Plantae
Division: Bryophyta
Class: Bryopsida
Subclass: Bryidae
Order: Rhizogoniales
Family: Aulacomniaceae
Genus: Aulacomnium
Species:
A. palustre
Binomial name
Aulacomnium palustre

Aulacomnium palustre, the bog groove-moss [1] or ribbed bog moss, is a moss that is nearly cosmopolitan in distribution. It occurs in North America, Hispaniola, Venezuela, Eurasia, and New Zealand. In North America, it occurs across southern arctic, [2] subboreal, [3] and boreal [4] regions from Alaska and British Columbia to Greenland and Quebec. Documentation of ribbed bog moss's distribution in the contiguous United States is probably incomplete. It is reported sporadically south to Washington, Wyoming, Georgia, and Virginia.

Contents

Habitat types and plant communities

Ribbed bog moss is frequent in arctic to subboreal wetlands. Moss assemblages are typically diverse in northern (arctic, subarctic, and boreal) plant communities, and individual moss species often have low cover and/or frequency. Moss species with coverages of 2% to 4% can be common to dominant in boreal communities, although ribbed bog moss attains coverages as great as 40% in some boreal communities.

Ribbed bog moss grows in open and forested wetland communities. In unforested northern communities, ribbed bog moss is found in sedge (Carex spp.) meadows, sphagnum (Sphagnum spp.) peatlands, heath-sedge fens, and willow (Salix spp.)-dominated fens. In forests, ribbed bog moss grows in the ground layer of boreal and subboreal white spruce (Picea glauca), black spruce (P. mariana), mixed spruce-tamarack (Larix laricina), and jack pine (Pinus banksiana) fens and bogs of Alaska, Minnesota, and Canada and in boreal spruce-birch (Betula spp.) forests of Alaska and northwestern Canada. Mosses are abundant in taiga forests of interior Alaska and Canada, forming characteristic strata in nearly every taiga forest type.

Less is known of ribbed bog moss associations south and east of Minnesota, although ribbed bog moss has been noted in some swamp, coniferous and/or hardwood bog, and grassland communities. Ribbed bog moss grows in red maple (Acer rubrum) swamps of Long Island, New York, and Little listed ribbed bog moss as common (1-4% frequency) in Atlantic white-cedar (Chamaecyparis thyoides) swamps of southern New Jersey. [5] Ribbed bog moss is also common in jack pine, aspen (Populus spp.), and mixed-hardwood forests of the Great Lakes states and southern Canada. Ribbed bog moss grows on tallgrass prairie in Kansas and Arkansas. In the Pacific Northwest ribbed bog moss occurs in alpine, subalpine, wet and dry coniferous forest, and open peatland communities. In a survey of alpine and unforested subalpine communities of the North Cascade Range in Washington and British Columbia, ribbed bog moss occurred in graminoid, forb, heath, and willow communities.

General botanical characteristics

Mosses have 2 phases in their life cycle: the gametophyte (n) and sporophyte (2n) generations. Each generation is morphologically distinct.

Gametophytes

Ribbed bog moss stems comprise most of ribbed bog moss's biomass and are easily visible. Stems are erect and spreading in habit, forming clumps or lawns. They range from 1 to 4 inches (3–9 cm) long; most stems are vegetative but some bear reproductive organs. Short vegetative stems may end in a stalk-bearing clusters of gemmae. Ribbed bog moss is heterothallic, with male and female reproductive organs borne on separate reproductive shoots. Male and female stems develop antheridia and archegonia, respectively, at their tips. Approximately the top 0.6 inch (1.5 cm) of both vegetative and reproductive stems is alive; lower stem tissue is usually dead. Ribbed bog moss leaves are bright yellowish-green to green; their bright color sometimes gives ribbed bog moss an incandescent appearance ("glow moss"). Bright leaves that contrast starkly with the reddish-brown stems typically make ribbed bog moss the most conspicuous species in moss assemblages. The leaves are lanceolate in shape and often tomentose, becoming twisted and brown when dry. They range from 3 to 5 mm long. Ribbed bog moss anchors to the substrate with rhizoids. Because ribbed bog moss lacks vascular tissue, water uptake occurs by osmosis and capillary action. A network of capillary spaces between stems and rhizoids enhances water uptake; ribbed bog moss usually absorbs water more efficiently than associated sphagnum mosses.

Sporophytes

Sporophytes grow out of archegonia. The sporophyte consists of a foot that anchors the sporophyte to the archegonia, a stalk, and a spore capsule. Ribbed bog moss stalks are vertically straight and about 1.8 inches (4.5 cm) long. Ribbed bog moss is named for its distinct spore capsule, which is strongly ribbed, cylindrical, and about 4 mm long. The capsule is capped with a calyptra.

Regeneration processes

Ribbed bog moss regenerates sexually and vegetatively.

Gametophyte dispersal and establishment

A spore is the first growth stage of a developing gametophyte. When the spore capsule matures, ribbed bog moss's calyptra splits along the side, exposing spores. Release of the exposed spores requires dry weather and is governed by a row of "teeth" that ring the capsule's top. The capsule teeth are hygroscopic, bending outward when air is dry and permitting spores to fall. Wind disperses ribbed bog moss spores long distances by shaking the capsule. When air is moist, the teeth bend inward, holding the spores within the capsule. The spores require a moist substrate to germinate. A germinated spore develops into a protonema (a branched, threadlike structure). Rhizoids grow down from the protonema and penetrate the substrate. Stems arise from buds that develop on the protonema surface. As stems grow, they develop their own rhizoids and become independent of the protonema. Mature male and female stems develop antheridia and archegonia, which produce sperm and eggs, respectively. Ribbed bog moss antheridia do not develop synchronously on the same stem. Mature and immature antheridia are intermixed on individual male shoots; therefore, sperm cells on the same stem do not all develop at the same time. However, sperm cells within a single antheridium have synchronous development. Fertilization requires a moist or saturated environment. Before fertilization, the antheridium absorbs water and swells, forcing the spore cap off. Rain may splash sperm into the archegonium, or sperm may swim to the archegonium.

Spore banking

Ribbed bog moss may germinate from spores stored in the substrate, but banked spores are probably less important to ribbed bog moss regeneration than freshly dispersed spores. Ross-Davis and Frego found that annual dispersal deposited a far greater number of moss spores on boreal substrates compared to the number of spores buried in the spore bank. [6]

Breeding system

Ribbed bog moss is dioecious.

Sporophyte development

Ribbed bog moss's sporophyte generation develops from the fertilized egg. Eggs are fertilized within the archegonium. The sporophyte embryo grows rapidly, differentiating into foot, stalk, and capsule tissue. Spores develop within the capsule.

Vegetative regeneration

Ribbed bog moss reproduces asexually from specialized gametophyte tissues and from plant breakage. It reproduces frequently from gemmae. Ribbed bog moss may also regenerate when paraphyses (minute filaments arising from ribbed bog moss's antheridia) detach. In the laboratory, 12.5% of detached ribbed bog moss paraphyses developed into propagules. Ribbed bog moss establishes readily when chunks of ribbed bog moss shoots are moved to new sites by soil movement or transplanting. Ribbed bog moss is apparently competitive in its ability to establish from stem chunks. In a laboratory experiment, ribbed bog moss that was collected from an Alberta peatland, shredded, and placed on a peat substrate showed greatest frequency (100%) of 4 moss species so treated. Ribbed bog moss also showed fastest growth relative to the other mosses throughout the 125-day experiment.

Growth

Ribbed bog moss growth is robust. It showed a "tall and dense growth habit" in a greenhouse common garden; ribbed bog moss, juniper hair cap moss (Polytrichum juniperinum), and papillose sphagnum (S. papillosum) crowded out 3 other moss species. Dry climate slows or stops ribbed bog moss growth. On the Boreal Ecosystem Research and Monitoring Sites study area in Saskatchewan, ribbed bog moss had a negative mean annual growth rate in a drought year (2003). Mean annual growth rate in a wet year (2004) was 2.7 mm. Ribbed bog moss was sensitive to saturated conditions in the wet year; stem lengths were greatest on relatively drier microsites, and ribbed bog moss growth rate increased slightly with increasing depth to the water table.

Site characteristics

Ribbed bog moss is a habitat generalist. It was, for example, 1 of 6 mosses having broad ecological amplitude in a survey of bryophyte habitats on peatlands across Alberta's Mackenzie River basin. Ribbed bog moss tolerates a wide range of moisture levels, substrates, nutrient loads, terrain, and climates.

Moisture regime

Ribbed bog moss generally grows on wetlands including fens, bogs, marshes, pond margins, streambanks, wet meadows, and riparian shrublands. In subalpine fir forests of central Idaho, ribbed bog moss occurs on seeps and springs that remain moist throughout the fire season. Ribbed bog moss is an indicator species of wet to very wet soils in Canada. In northern British Columbia, ribbed bog moss is an indicator species of undisturbed wet conifer sites. The white spruce/field horsetail/ribbed bog moss association occurs on the wettest white spruce forests in subboreal British Columbia; the water table is near the soil surface for most of the growing season. In a geothermal meadow in Haida Gwaii, British Columbia, ribbed bog moss occurred on sites with high local humidity (31-66%) due to nearby thermal pools. Ribbed bog moss did not grow on dry sites, although drained microsites may favor ribbed bog moss growth on otherwise saturated substrates. Ribbed bog moss does not tolerate salt spray, which prevents its establishment on coastal dunelands.

Ribbed bog moss is not confined to wet sites in all areas. Some forests with ribbed bog moss dry in late summer, and ribbed bog moss grows on relatively xeric hummock mounds on bogs in Birds Hill Provincial Park, Manitoba. Ribbed bog moss occupies both dry and wet sites on the Boreal Ecosystem Research and Monitoring Sites study area. It dominates relatively dry, shaded microsites in the area; the water table is 20 to 26 inches (50–65 cm) below ground, and there is 30% to 60% black spruce and/or tamarack cover.

Substrates

Ribbed bog moss is primarily a groundlayer species, but it does not require a particular substrate to establish and grow. It is most common on peat but also grows on thinner organic soils and other substrates. Ribbed bog moss frequently grows on peat overlying permafrost in Alaska and northern Canada. On northern peatlands, the peat layer generally ranges from 38 to 102 inches (15–40 cm) thick, and organic content of the soil layer is high. Studies on peatlands in Quebec showed ribbed bog moss "preferred sites with high organic matter depth" (P<0.01). Ribbed bog moss grows on organic surface layers overlying varying soil textures. Ribbed bog moss also grows on burned substrates including ash, mineral soil, scorched organic soil, scorched peat, and scorched downed woody debris.

Ribbed bog moss grows on peat and other organic soil layers more often than on downed bark or wood, but is reported growing on woody debris or other dead wood in a few locations. In northern British Columbia, ribbed bog moss substrates included disturbed forest floors, logs, and stumps at 44%, 13%, and 3% frequencies, respectively. Ribbed bog moss was found on downed woody debris in a mixed quaking aspen-paper birch-balsam fir (Populus tremuloides-Betula papyrifera-Abies balsamifera) forest in east-central Alberta and on stumps in a mixed-hardwood forest in Wisconsin. Ribbed bog moss rarely grows on standing live or dead wood.

Ribbed bog moss showed broad substrate tolerances in a greenhouse common pot study in Scotland. Ribbed bog moss vegetative propagules were sown with propagules of 6 other mosses to test substrate preferences. After 1 year, ribbed bog moss abundance was similar on heather (Calluna vulgaris) litter, European white birch (Betula pendula) litter, dead shrub litter, Scots pine (Pinus sylvestris) needles, sand, and sphagnum peat substrates. To test particle-size microsite preferences on peat substrates, the peat was broken into various fragment sizes from minute to large (<0.25 inch to >2 inches (0.63–5 cm)). Ribbed bog moss grew on peat of all particle sizes but was most frequent on small (0.5-1 inch (1.25-2.5 cm)) peat particles.

Water and substrate chemistry

The pH of water, peat, and/or soil is usually acidic to neutral in mires with ribbed bog moss, although ribbed bog moss tolerates mildly alkaline conditions. For example, ribbed bog moss grows in extremely acidic peatlands overlying permafrost in spruce taiga of Alaska but also grows in calcareous bogs in Birds Hill Provincial Park. In Minnesota, ribbed bog moss is reported from bogs ranging from 5.0 to 7.3 in pH. A survey of bryophytes on peatlands across Alberta's Mackenzie River basin found ribbed bog moss was abundant on sites ranging from 4.5 to 7.5 in pH.

Mires are classified on pH and mineral gradients from extreme-poor (very strongly acid and low in calcium and magnesium) to extreme-rich (neutral to alkaline and high in calcium and magnesium). Fens are richer than bogs. Ribbed bog moss occurs in poor and rich mires. On mires across British Columbia and Alberta, ribbed bog moss grows in mires with broad ranges of water pH and electrical conductivity, but is most common on strongly acidic peatlands (water pH <5.5) with moderate calcium and magnesium levels (<200 μS/cm**). Ribbed bog moss also grows in moderate-rich fens with higher pH and electrical conductivity values. In a peat-core study in central Alberta, macrofossil ribbed bog moss was an indicator species of moderate-rich fens; ribbed bog moss occurred most often on mires that were moderately acidic (x pH=6.0) and had low electrical conductivity (x=125 μS/cm) and moderately high water tables (x= 8.7 inches (22 cm) deep). In a study of alpine mires in Italy, ribbed bog moss was intermediate in mire pH and mineral content compared to associated nonvascular and vascular plant species, occurring on both poor and rich mires. All associated mosses had narrower pH and mineral tolerances than ribbed bog moss. Gignac provides information on ribbed bog moss habitats in British Columbia and Alberta including ranges in pH, electrical conductivity, relative depth to the water table, and relative overstory cover.

Nutrients

Field observations and laboratory experiments suggest that ribbed bog moss has broad tolerance and may be relatively insensitive to macronutrient concentrations. On the Svalbard archipelago in Norway, ribbed bog moss grows on small "bird islands" where eiders, Arctic terns, and other migratory birds nest offshore of the main island, Spitsbergen. Nitrogen levels on the bird islands are very high. On Spitsbergen Island, however, ribbed bog moss grows on dry hummocks and moist hummock edges, both of which have low nitrogen levels but probably provide moisture levels that favor ribbed bog moss. In a laboratory experiment, nitrogen fertilizer initially slowed ribbed bog moss growth rate, but growth rates of ribbed bog moss with and without nitrogen were similar at the end of 125 days. Addition of phosphorus had no effect on ribbed bog moss growth.

Ribbed bog moss is listed as an indicator species of nitrogen-medium soils in British Columbia.

Landscape

Tundra and taiga areas where ribbed bog moss grows are generally flat to gently sloped, but local relief creates drainage patterns that often result in distinct moss assemblages. Site geology, including topography, bedrock composition, catchment hydrology, and basin bathymetry, affects wetland drainage and partially controls rates of transition from open water to bog. Ribbed bog moss is common on hummocks, which tend to dry out faster than adjacent lowlands. In British Columbia and Alberta, ribbed bog moss often dominates hummock tops that are surrounded by sphagnum peatlands. Although ribbed bog moss generally attains greatest coverage on hummock tops, it sometimes forms lawns and strings in low areas. In Labrador, tufted bulrush-mountain fly honeysuckle/ribbed bog moss associations occur on low strings of consolidated peat and on elevated peatlands. The low strings lie 38 to 76 inches (15–30 cm) above the water table and lack erosion patterns.

In Kluane National Park, Yukon, the white spruce/ribbed bog moss forest community occurs on poorly drained east- and north-facing slopes.

Elevation

There are few reports of ribbed bog moss's elevational tolerances. Ribbed bog moss is reported from 1,600 to 5,700 feet (500-1,750 m) in west-central Alberta and at 827 feet (252 m) in Kosciucko County, Indiana. In subalpine and alpine communities of the North Cascade Range in Washington and British Columbia, ribbed bog moss occurs on high-elevation (>7,380 feet (2,250 m)) sites that remain snow-free most of the year.

Climate

Ribbed bog moss occurs in arctic, subarctic, boreal, and subboreal zones with cold meso-thermal, oceanic, continental, or cold-humid climates. It is more common in arctic, subarctic, and boreal than subboreal zones. A survey of bryophytes on peatlands across Alberta's Mackenzie River basin found optimal ribbed bog moss growth occurred on sites with annual temperatures ranging from 20 to 32 °F (-4 to 0 °C). In interior arctic, subarctic, and boreal zones, climate is strongly continental in the west, becoming more humid to the east. Climate shift from continental to humid generally occurs near Hudson Bay and Lake Superior. In a study of moss habitats across British Columbia, Alberta, and Manitoba, ribbed bog moss was an indicator species for continentality of climate: it was the only moss common to all continental peatlands surveyed. On sites from coastal British Columbia to central Alberta, ribbed bog moss was most common on subcontinental sites (intermediate between coastal and continental climates). Ribbed bog moss was intermediate on gradients ranking breadth of moss habitat niches. Climate factors evaluated included length of growing season, amount of precipitation during the growing season, temperature, and aridity. Feather mosses (Hylocomiaceae) and sphagnum mosses generally had wider niches and dominated more sites than ribbed bog moss. Ribbed bog moss's rarity on all but cold sites in the lower 48 states suggests that ribbed bog moss does not tolerate long periods of warm weather. In a geothermal meadow on Queen Charlotte Island, British Columbia, ribbed bog moss was absent from sites where nearby thermal pools raised local soil temperatures above 86 °F (30 °C).

Uses

Importance to wildlife and livestock

Ribbed bog moss provides few known direct benefits to wildlife or livestock. Mosses in general are low in carbohydrates, proteins, and fats compared to vascular plants, and animals seldom graze them. Caribou may eat mosses when little other forage is available. Wildlife seeking cover probably avoid open areas dominated by ribbed bog moss or other low vegetation. In Wood Buffalo National Park, Alberta, aerial photos identified areas dominated by ribbed bog moss, bog Labrador tea, and/or Bebb willow as indicators of nonnesting habitat for whooping cranes.

American robins at the Mountain Lake Biological Station, Virginia, used ribbed bog moss as nest material.

Northern mires provide habitat for a variety of invertebrates including worms, crustaceans, arachnids, and insects, particularly mosquitoes, midges, and other flies.

Value for rehabilitation of disturbed sites

Chunks from ribbed bog moss lawns can be transplanted onto disturbed sites. In Minnesota, moss plugs used in restoration projects on old peat mines are harvested from nearby unmined sites in spring, before the ground thaws. Although ribbed bog moss is not often used for restoration, it could be a valuable addition to restoration projects. Foote classified ribbed bog moss communities along the Alaskan Highway in Yukon as having "high site sensitivity", with moderate potential for erosion. [7] Black spruce/ribbed bog moss communities on the Alaskan Pipeline are rated "sensitive to highly sensitive" to erosion and disturbance.

Notes

  1. Edwards, Sean R. (2012). English Names for British Bryophytes. British Bryological Society Special Volume. 5 (4 ed.). Wootton, Northampton: British Bryological Society. ISBN   978-0-9561310-2-7. ISSN   0268-8034.
  2. "Arctic" refers to treeless areas underlain with continuous permafrost; "subarctic" refers to mixed tundra and open forest underlain with discontinuous permafrost.
  3. "Subboreal" refers to areas in southern Canada, the Great Lakes states, and the Northeast that were once underlain with permafrost that has since melted.
  4. "boreal" refers to northern coniferous forests where permafrost occurs only sporadically.
  5. Little, S. 1951. Observations on the minor vegetation of the pine barren swamps in southern New Jersey. Bulletin of the Torrey Botanical Club. 78(2): 153-160.
  6. Ross-Davis, Amy L. and Katherine A. Frego. 2004. Propagule sources of forest floor bryophytes: spatiotemporal compositional patterns. The Bryologist. 107(1): 88-97.
  7. Foote, M. Joan. 1983. Classification, description, and dynamics of plant communities after fire in the taiga of interior Alaska. Res. Pap. PNW-307. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 108 p.

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Peat, also known as turf, is an accumulation of partially decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem is the most efficient carbon sink on the planet, because peatland plants capture CO2 naturally released from the peat, maintaining an equilibrium. In natural peatlands, the "annual rate of biomass production is greater than the rate of decomposition", but it takes "thousands of years for peatlands to develop the deposits of 1.5 to 2.3 m [4.9 to 7.5 ft], which is the average depth of the boreal [northern] peatlands". Sphagnum moss, also called peat moss, is one of the most common components in peat, although many other plants can contribute. The biological features of Sphagnum mosses act to create a habitat aiding peat formation, a phenomenon termed 'habitat manipulation'. Soils consisting primarily of peat are known as histosols. Peat forms in wetland conditions, where flooding or stagnant water obstructs the flow of oxygen from the atmosphere, slowing the rate of decomposition.

Bog Type of wetland that accumulates peat due to incomplete decomposition of plant matter

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Moss Division of plants

Mosses, or the taxonomic division Bryophyta, are small, non-vascular flowerless plants that typically form dense green clumps or mats, often in damp or shady locations. The individual plants are usually composed of simple leaves that are generally only one cell thick, attached to a stem that may be branched or unbranched and has only a limited role in conducting water and nutrients. Although some species have conducting tissues, these are generally poorly developed and structurally different from similar tissue found in vascular plants. Mosses do not have seeds and after fertilisation develop sporophytes with unbranched stalks topped with single capsules containing spores. They are typically 0.2–10 cm (0.1–3.9 in) tall, though some species are much larger. Dawsonia, the tallest moss in the world, can grow to 50 cm (20 in) in height.

Bryophyte terrestrial plants that lack vascular tissue

Bryophytes are an informal group consisting of three divisions of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. They are characteristically limited in size and prefer moist habitats although they can survive in drier environments. The bryophytes consist of about 20,000 plant species. Bryophytes produce enclosed reproductive structures, but they do not produce flowers or seeds. They reproduce via spores. Bryophytes are usually considered to be a paraphyletic group and not a monophyletic group, although some studies have produced contrary results. Regardless of their status, the name is convenient and remains in use as an informal collective term. The term "bryophyte" comes from Greek βρύον, bryon "tree-moss, oyster-green" and φυτόν, phyton "plant".

<i>Picea mariana</i> species of plant

Picea mariana, the black spruce, is a North American species of spruce tree in the pine family. It is widespread across Canada, found in all 10 provinces and all 3 Arctic territories. Its range extends into northern parts of the United States: in Alaska, the Great Lakes region, and the upper Northeast. It is a frequent part of the biome known as taiga or boreal forest.

<i>Sphagnum</i> genus of mosses, peat moss

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Meesia uliginosa, the broad-nerved hump-moss, is a rare moss of the Western U.S. It occurs all around the northern hemisphere in higher latitudes, and in some places is not as rare as in the Western U.S.

Poor fen

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<i>Ceratodon purpureus</i> species of plant

Ceratodon purpureus is a dioicous moss with a color ranging from yellow-green to red. The height amounts to 3 centimeters. It is found worldwide, mainly in urban areas and next to roads on dry sand soils. It can grow in a very wide variety of habitats, from polluted highway shoulders and mine tailings to areas recently denuded by wildfire to the bright slopes of Antarctica. Its common names include redshank, purple forkmoss, ceratodon moss, fire moss, and purple horn toothed moss.

Raised bog acidic, wet habitats that are poor in mineral salts

Raised bogs, also called ombrotrophic bogs, are acidic, wet habitats that are poor in mineral salts and are home to flora and fauna that can cope with such extreme conditions. Raised bogs, unlike fens, are exclusively fed by precipitation (ombrotrophy) and from mineral salts introduced from the air. They thus represent a special type of bog, hydrologically, ecologically and in terms of their development history, in which the growth of peat mosses over centuries or millennia plays a decisive role. They also differ in character from blanket bogs which are much thinner and occur in wetter, cloudier climatic zones.

Mire Wetland terrain without forest cover, dominated by living, peat-forming plants

A mire, peatland or quagmire is a wetland type, dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types of mires share the common characteristic of being saturated with water at least seasonally with actively forming peat, while having its own set of vegetation and organisms. Like coral reefs, mires are unusual landforms in that they derive mostly from biological rather than physical processes, and can take on characteristic shapes and surface patterning.

<i>Sphagnum fuscum</i> peat moss

Sphagnum fuscum, the rusty bogmoss or rusty peat moss, is a peat moss found commonly in Norway and Sweden, and can be found scattered across North America, the United Kingdom, and in southern to eastern Europe.

<i>Sphagnum cuspidatum</i> species of moss

Sphagnum cuspidatum, the feathery bogmoss, toothed sphagnum, or toothed peat moss, is a peat moss found commonly in Great Britain, Norway, Sweden, the eastern coast of the United States, and in Colombia.

<i>Trichophorum cespitosum</i> species of plant

Trichophorum cespitosum, commonly known as deergrass or tufted bulrush, is a species of flowering plant in the sedge family. It was originally described by the Swedish naturalist Carl Linnaeus in 1753 as Scirpus cespitosus, but was transferred to the genus Trichophorum by the Swedish botanist Carl Johan Hartman in 1849, becoming Trichophorum cespitosum.

<i>Climacium dendroides</i> species of plant

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<i>Polytrichum strictum</i> species of plant

Polytrichum strictum, commonly known as bog haircap moss or strict haircap, is an evergreen and perennial species of moss native to Sphagnum bogs and other moist habitats in temperate climates. It has a circumboreal distribution, and is also found in South America and Antarctica.

<i>Sphagnum papillosum</i> species of plant

Sphagnum papillosum, the papillose peatmoss, is a species of peat moss distributed throughout the northern hemisphere. Although sometimes confused with Sphagnum imbricatum and Sphagnum palustre, it is distinguished by its yellow-green to brown short, blunt branches and papillose chlorophyllose cells.

<i>Buxbaumia viridis</i> species of plant

Buxbuamia viridis, also known as the green shield moss, is a rare bryophyte found sporadically throughout the northern hemisphere. The gametophyte of this moss is not macroscopically visible; the large, distinct sporophyte of B. viridis is the only identifying structure of this moss. This moss can be found singularly or in small groups on decaying wood, mostly in humid, sub-alpine to alpine Picea abies, Abies alba, or mixed tree forests. This moss is rare and conservation efforts are being made in most countries B. viridis is found in.

<i>Plagiomnium venustum</i> species of plant

Plagiomnium venustum, also known as magnificent leafy moss, is a species of moss belonging to the family Mniaceae. It is found mainly in western North America along the coastal region. This moss can be identified from other members of the Plagiomnium genus by dark coloured stomata guide cells and the absence of sterile stems. It is most commonly found growing as a mat on a variety of substrate, but mainly on humus and moist soil.

References