Balsam fir | |
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Tree with cones | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Gymnospermae |
Division: | Pinophyta |
Class: | Pinopsida |
Order: | Pinales |
Family: | Pinaceae |
Genus: | Abies |
Species: | A. balsamea |
Binomial name | |
Abies balsamea | |
Range | |
Synonyms [2] | |
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Abies balsamea or balsam fir is a North American fir, native to most of eastern and central Canada (Newfoundland west to central Alberta) and the northeastern United States (Minnesota east to Maine, and south in the Appalachian Mountains to West Virginia). [3]
Balsam fir is a small to medium-size evergreen tree typically 14–20 metres (46–66 ft) tall, occasionally reaching a height of 27 metres (89 ft). The narrow conic crown consists of dense, dark-green leaves. The bark on young trees is smooth, grey, and with resin blisters (which tend to spray when ruptured), becoming rough and fissured or scaly on old trees. The leaves are flat and needle-like, 15 to 30 mm (5⁄8 to 1+1⁄8 in) long, dark green above often with a small patch of stomata near the tip, and two white stomatal bands below, and a slightly notched tip. They are arranged spirally on the shoot, but with the leaf bases twisted so that the leaves appear to be in two more-or-less horizontal rows on either side of the shoot. The needles become shorter and thicker the higher they are on the tree. The seed cones are erect, 40 to 80 mm (1+1⁄2 to 3+1⁄4 in) long, dark purple, ripening brown and disintegrating to release the winged seeds in September.
For thousands of years Native Americans used balsam fir for medicinal and therapeutic purposes. The needles are eaten directly off the tree by many animals and humans. Higher content dosage is ingested in tea. Balsam fir contains vitamin C, which has been studied for its effects on bacterial and viral infections. [4]
Balsam fir's essential oil and some of its compounds have shown efficacy against ticks. [5]
The male reproductive organs generally develop more rapidly and appear sooner than the female organs. The male organs contain microsporangia which divide to form sporogenous tissue, composed of cells which become archesporial cells. These develop into microspores, or pollen-mother cells, once they are rounded and filled with starch grains. When the microspores undergo meiosis in the spring, four haploid microspores are produced which eventually become pollen grains. Once the male strobilus has matured the microsporangia are exposed at which point the pollen is released.
The female megasporangiate is larger than the male. It contains bracts and megasporophylls, each of which contains two ovules, arranged in a spiral. These then develop a nucellus in which a mother cell is formed. Meiosis occurs and a megaspore is produced as the first cell of the megagametophyte. As cell division takes place the nucleus of the megaspore thickens, and cell differentiation occurs to produce prothallial tissue containing an ovum. The remaining undifferentiated cells then form the endosperm.
When the male structure releases its pollen grains, some fall onto the female strobilus and reach the ovule. At this point the pollen tube begins to generate, and eventually the sperm and egg meet at which point fertilization occurs. [6]
There are two varieties:
Balsam firs are very shade tolerant, and tend to grow in cool climates, ideally with a mean annual temperature of 40 °F (4 °C), with consistent moisture at their roots. [7] They typically grow in the following four forest types:
The foliage is browsed by moose and deer. [9] The seeds are eaten by American red squirrels, grouse, and pine mice; [10] the tree also provides food for crossbills and chickadees, as well as shelter for moose, snowshoe hares, white-tailed deer, ruffed grouse, and other small mammals and songbirds. The needles are eaten by some lepidopteran caterpillars, for example the Io moth (Automeris io).
Abies balsamea is one of the most cold-hardy trees known, surviving at temperatures as low as −45 °C (−49 °F) (USDA Hardiness Zone 2). Specimens even showed no ill effects when immersed in liquid nitrogen at −196 °C (−320.8 °F). [11]
It is listed as endangered in Connecticut. This status applies to native populations only. [12]
The balsam fir is the preferred main host of the eastern spruce budworm, which is a major destructive pest throughout the eastern United States and Canada. [13] During cyclical population outbreaks, major defoliation of the balsam fir can occur, which may significantly reduce radial growth. [14] This can kill the tree. An outbreak in Quebec in 1957 killed over 75% of balsam fir in some stands. [15]
The needles of balsam fir can be infected by the fungus Delphinella balsameae . [16]
Both varieties of the species are very popular as Christmas trees, particularly in the northeastern United States. Balsam firs cut for Christmas are not taken from the forest, but are grown on large plantations. The balsam fir is one of the greatest exports of Quebec and New England. It is celebrated for its rich green needles, natural conical shape, and needle retention after being cut, and it is notably the most fragrant of all Christmas tree varieties. [17]
The balsam fir was used six times for the US Capitol Christmas Tree between 1964 and 2019. [11]
Abies balsamea is also grown as an ornamental tree for parks and gardens. Very hardy down to −20 °C (−4 °F) or below, it requires a sheltered spot in full sun. The dwarf cultivar A. balsamea 'Hudson' (Hudson fir), grows to only 1 m (3.3 ft) tall by 1.5 m (4.9 ft) broad, and has distinctive blue-green foliage with pale undersides. It does not bear cones. It has gained the Royal Horticultural Society's Award of Garden Merit. [18] [19]
Other cultivars include:
The resin is used to produce Canada balsam, and was traditionally used as a cold remedy and as a glue for glasses, optical instrument components, and for preparing permanent mounts of microscope specimens. Given its use as a traditional remedy and the relatively high ascorbic acid content of its needles, historian Jacques Mathieu has argued that the balsam fir was the "aneda" that cured scurvy during the second expedition into Canada of Jacques Cartier. [20] The wood is milled for framing lumber (part of SPF lumber), siding and pulped for paper manufacture. Balsam fir oil is an EPA approved nontoxic rodent repellent. The balsam fir is also used as an air freshener and as incense. [21]
Prior to the availability of foam rubber and air mattresses, balsam fir boughs were a preferred mattress in places where trees greatly outnumbered campers. Many fir limbs are vertically bowed from alternating periods of downward deformation from snow loading and new growth reaching upward for sunlight. Layers of inverted freshly cut limbs from small trees created a pleasantly fragrant mattress lifting bedding off the wet ground; and the bowed green limbs were springs beneath the soft needles. Upper layers of limbs were placed with the cut ends of the limbs touching the earth to avoid uncomfortably sharp spots and sap. [22]
Native Americans use it for a variety of medicinal purposes. [23]
The Abenaki use the gum for slight itches and as an antiseptic ointment. [24] They stuff the leaves, [25] needles, and wood into pillows as a panacea. [26]
The Algonquin people of Quebec apply a poultice of the gum to open sores, insect bites, boils and infections, use the needles as a sudatory for women after childbirth and for other purposes, use the roots for heart disease, use the needles to make a laxative tea, and use the needles for making poultices. [27]
The Atikamekw chew the sap as a cold remedy, and use the boughs as mats for the tent floor. [28]
The Cree use the pitch for menstrual irregularity, and take an infusion of the bark and sometimes the wood for coughs. They use the pitch and grease used as an ointment for scabies and boils. They apply a poultice of pitch applied to cuts. They also use a decoction of pitch and sturgeon oil used for tuberculosis, and take an infusion of bark for tuberculosis. They also use the boughs to make brush shelters and use the wood to make paddles. [29]
The Innu people grate the inner bark and eat it to benefit their diet. [30]
The Iroquois use a steam from a decoction of branches as a bath for rheumatism and parturition, and ingest a decoction of the plant for rheumatism. They take a compound decoction for colds and coughs, sometimes mixing it with alcohol. They apply a compound decoction of the plant for cuts, sprains, bruises and sores. [31] They apply a poultice of the gum and dried beaver kidneys for cancer. [32] They also take a compound decoction in the early stages of tuberculosis, and they use the plant for bedwetting and gonorrhea. [33]
The Maliseet use the juice of the plant as a laxative, [34] use the pitch in medicines, [35] and use an infusion of the bark, sometimes mixed with spruce and tamarack bark, for gonorrhea. [36] They use the needles and branches as pillows and bedding, the roots as thread, and use the pitch to waterproof seams in canoes. [35]
The Menominee use the inner bark as a seasoner for medicines, take an infusion of the inner bark for chest pain, and use the liquid balsam pressed from the trunk for colds and pulmonary troubles. They also use the inner bark as a poultice for unspecified illnesses. [37] They also apply gum from plant blisters to sores. [38]
The Miꞌkmaq use a poultice of inner bark for an unspecified purpose, [37] use the buds, cones and inner bark for diarrhea, use the gum for burns, colds, fractures, sores and wounds, use the cones for colic, and use the buds as a laxative. They also use the bark used for gonorrhea and buds used as a laxative. [39] They use the boughs to make beds, use the bark to make a beverage, and use the wood for kindling and fuel. [40]
The Ojibwe melt the gum on warm stones and inhale the fumes for headache. [41] They also use a decoction of the root as an herbal steam for rheumatic joints. [41] They also combine the gum with bear's grease and use it as an ointment for hair. [42] They use the needle-like leaves in as part of ceremony involving the sweat bath, and use the gum for colds and inhale the leaf smoke for colds. [43] They use the plant as a cough medicine. [44] The gum is used for sores and a compound containing leaves is used as wash. The liquid balsam from bark blisters is used for sore eyes. [43] They boil the resin twice and add it to suet or fat to make a canoe pitch. [45] The bark gum is taken for chest soreness from colds, applied to cuts and sores, and decoction of the bark is used to induce sweating. The bark gum is also taken for gonorrhea. [46]
The Penobscot smear the sap over sores, burns, and cuts. [47]
The Potawatomi use the needles to make pillows, believing that the aroma prevented one from getting a cold. [48] They also use the balsam gum as a salve for sores, and take an infusion of the bark for tuberculosis and other internal afflictions. [48]
Balsam fir is the provincial tree of New Brunswick.
Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta, also known as Coniferophyta or Coniferae. The division contains a single extant class, Pinopsida. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas-firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. As of 2002, Pinophyta contained seven families, 60 to 65 genera, and more than 600 living species.
Firs are evergreen coniferous trees belonging to the genus Abies in the family Pinaceae. There are approximately 48–65 extant species, found on mountains throughout much of North and Central America, Eurasia, and North Africa. The genus is most closely related to Keteleeria, a small genus confined to eastern Asia.
Picea mariana, the black spruce, is a North American species of spruce tree in the pine family. It is widespread across Canada, found in all 10 provinces and all 3 territories. It is the official tree of Newfoundland and Labrador and is that province's most abundant tree. Its range extends into northern parts of the United States: in Alaska, the Great Lakes region, and the upper Northeast. It is a frequent part of the biome known as taiga or boreal forest.
Pinus nigra, the Austrian pine or black pine, is a moderately variable species of pine, occurring across Southern Europe from the Iberian Peninsula to the eastern Mediterranean, on the Anatolian peninsula of Turkey, Corsica and Cyprus, as well as Crimea and in the high mountains of Northwest Africa. The world's oldest black pine, located in the Banaz district of Uşak, Türkiye, is estimated to be 1000 years old. This makes it significant in the country, which is known for a very dry climate, inhospitable for most trees. It has a length of 11 meters, a diameter of 3 meters and a circumference of 9.60 meters.
Abies alba, the European silver fir or silver fir, is a fir native to the mountains of Europe, from the Pyrenees north to Normandy, east to the Alps and the Carpathians, Slovakia, Slovenia, Croatia, Bosnia and Herzegovina, Montenegro, Serbia, and south to Italy, Bulgaria, Kosovo, Albania and northern Greece.
Larix laricina, commonly known as the tamarack, hackmatack, eastern larch, black larch, red larch, or American larch, is a species of larch native to Canada, from eastern Yukon and Inuvik, Northwest Territories east to Newfoundland, and also south into the upper northeastern United States from Minnesota to Cranesville Swamp, West Virginia; there is also an isolated population in central Alaska.
Abies grandis is a fir native to northwestern North America, occurring at altitudes of sea level to 1,700 metres (5,600 ft). It is a major constituent of the Grand Fir/Douglas Fir Ecoregion of the Cascade Range.
The blue spruce, also commonly known as Colorado spruce or Colorado blue spruce, is a species of spruce tree native to North America in Arizona, Colorado, Idaho, New Mexico, Utah and Wyoming. It is noted for its blue-green colored needles, and has therefore been used as an ornamental tree in many places far beyond its native range.
Abies magnifica, the red fir or silvertip fir, is a western North American fir, native to the mountains of southwest Oregon and California in the United States. It is a high-elevation tree, typically occurring at 1,400–2,700 metres (4,600–8,900 ft) elevation, though only rarely reaching tree line. The name red fir derives from the bark color of old trees.
The Fraser fir, sometimes spelled Frasier fir, is an endangered species of fir native to the Appalachian Mountains of the southeastern United States. They are endemic to only seven montane regions in the Appalachian Mountains.
Abies amabilis, commonly known as the Pacific silver fir, is a fir native to the Pacific Northwest of North America, occurring in the Pacific Coast Ranges and the Cascade Range. It is also commonly referred to in English as the white fir, red fir, lovely fir, amabilis fir, Cascades fir, or silver fir. The species name is Latin for 'lovely'.
Juniperus monosperma is a species of juniper native to western North America, in the United States in Arizona, New Mexico, southern Colorado, western Oklahoma (Panhandle), and western Texas, and in Mexico in the extreme north of Chihuahua. It grows at 970–2300 m altitude.
The Great Balsam Mountains, or Balsam Mountains, are in the mountain region of western North Carolina, United States. The Great Balsams are a subrange of the Blue Ridge Mountains, which in turn are a part of the Appalachian Mountains. The most famous peak in the Great Balsam range is Cold Mountain, which is the centerpiece of author Charles Frazier's bestselling novel Cold Mountain. Other notable peaks include Richland Balsam, which is the highest peak in the range, Black Balsam Knob, and Mount Pisgah.
Spruce-pine-fir (SPF) is a classification of lumber that can be traded on commodities exchanges.
Pseudognaphalium obtusifolium, synonyms including Gnaphalium obtusifolium and Pseudognaphalium saxicola, is a member of the family Asteraceae. It is found on open dry sandy habitat throughout eastern North America. Common names include old field balsam, rabbit tobacco and sweet everlasting. When crushed, the plant exudes a characteristic maple-syrup scent.
This is a list of plants used by the indigenous people of North America. For lists pertaining specifically to the Cherokee, Iroquois, Navajo, and Zuni, see Cherokee ethnobotany, Iroquois ethnobotany, Navajo ethnobotany, and Zuni ethnobotany.
See also Zuni ethnobotany, and Native American ethnobotany.
This is a list of plants and how they are used in Zuni culture.
This is a list of plants documented to have been traditionally used by the Cherokee, and how they are used.
The Iroquois use a wide variety of medicinal plants, including quinine, chamomile, ipecac, and a form of penicillin.