Cycloneuralia

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Cycloneuralia
Priapulus caudatus.jpg
Priapulus caudatus
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Kingdom: Animalia
Superphylum: Ecdysozoa
Clade: Cycloneuralia
Subclades

Cycloneuralia is a clade of ecdysozoan animals including the Scalidophora (Kinorhynchans, Loriciferans, Priapulids) and the Nematoida (nematodes, Nematomorphs). [1] [2] It may be paraphyletic, or may be a sister group to Panarthropoda. Or perhaps Panarthropoda is paraphyletic with respect to Cycloneuralia. [3] The group has also been considered a single phylum, sometimes given the old name Nemathelminthes. [4] The uniting character is the nervous system organization with a circumpharyngeal brain and somata–neuropil–somata pattern. [5] The name derives from the position of the brain around the pharynx. [6]

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Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians. Primates arose 85–55 million years ago first from small terrestrial mammals, which adapted to living in the trees of tropical forests: many primate characteristics represent adaptations to life in this challenging environment, including large brains, visual acuity, color vision, a shoulder girdle allowing a large degree of movement in the shoulder joint, and dextrous hands. Primates range in size from Madame Berthe's mouse lemur, which weighs 30 g (1 oz), to the eastern gorilla, weighing over 200 kg (440 lb). There are 376–524 species of living primates, depending on which classification is used. New primate species continue to be discovered: over 25 species were described in the 2000s, 36 in the 2010s, and three in the 2020s.

<span class="mw-page-title-main">Paraphyly</span> Type of taxonomic group

In taxonomy, a grouping is paraphyletic if it consists of the grouping's last common ancestor and most of its descendants, but excludes a few monophyletic subgroups. The grouping is said to be paraphyletic with respect to the excluded subgroups. In contrast, a monophyletic grouping includes a common ancestor and all of its descendants. The terms are commonly used in phylogenetics and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic.

<span class="mw-page-title-main">Vertebrate</span> Subphylum of chordates with backbones

Vertebrates comprise all animal taxa within the subphylum Vertebrata, including all mammals, birds, reptiles, amphibians and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following:

<span class="mw-page-title-main">Loricifera</span> Phylum of tiny marine invertebrates

Loricifera is a phylum of very small to microscopic marine cycloneuralian sediment-dwelling animals with 43 described species. and approximately 100 more that have been collected and not yet described. Their sizes range from 100 μm to ca. 1 mm.

<span class="mw-page-title-main">Lobopodia</span> Group of extinct worm-like animals with legs

The lobopodians, members of the informal group Lobopodia, or the formally erected phylum Lobopoda Cavalier-Smith (1998), are panarthropods with stubby legs called lobopods, a term which may also be used as a common name of this group as well. While the definition of lobopodians may differ between literatures, it usually refers to a group of soft-bodied, worm-like fossil panarthropods such as Aysheaia and Hallucigenia.

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<span class="mw-page-title-main">Crab</span> Infraorder of decapod crustaceans

Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen), usually hidden entirely under the thorax. They live in all the world's oceans, in freshwater, and on land, are generally covered with a thick exoskeleton, and have a single pair of pincers. They first appeared during the Jurassic Period.

<span class="mw-page-title-main">Chinchilla</span> Rodent genus

Chinchillas are either of two species of crepuscular rodents of the parvorder Caviomorpha. They are slightly larger and more robust than ground squirrels, and are native to the Andes mountains in South America. They live in colonies called "herds" at high elevations up to 4,270 m (14,000 ft). Historically, chinchillas lived in an area that included parts of Bolivia, Peru and Chile, but today, colonies in the wild are known only in Chile. Along with their relatives, viscachas, they make up the family Chinchillidae. They are also related to the chinchilla rat.

<span class="mw-page-title-main">Ecdysozoa</span> Superphylum of protostomes including arthropods, nematodes and others

Ecdysozoa is a group of protostome animals, including Arthropoda, Nematoda, and several smaller phyla. The grouping of these animal phyla into a single clade was first proposed by Eernisse et al. (1992) based on a phylogenetic analysis of 141 morphological characters of ultrastructural and embryological phenotypes. This clade, that is, a group consisting of a common ancestor and all its descendants, was formally named by Aguinaldo et al. in 1997, based mainly on phylogenetic trees constructed using 18S ribosomal RNA genes.

<span class="mw-page-title-main">Panarthropoda</span> Animal taxon

Panarthropoda is a proposed animal clade containing the extant phyla Arthropoda, Tardigrada and Onychophora. Panarthropods also include extinct marine legged worms known as lobopodians ("Lobopodia"), a paraphyletic group where the last common ancestor and basal members (stem-group) of each extant panarthropod phylum are thought to have risen. However the term "Lobopodia" is sometimes expanded to include tardigrades and onychophorans as well.

<span class="mw-page-title-main">Concussion</span> Type of traumatic brain injury

A concussion, also known as a mild traumatic brain injury (mTBI), is a head injury that temporarily affects brain functioning. Symptoms may include loss of consciousness (LOC); memory loss; headaches; difficulty with thinking, concentration, or balance; nausea; blurred vision; sleep disturbances, and mood changes. Any of these symptoms may begin immediately, or appear days after the injury. Concussion should be suspected if a person indirectly or directly hits their head and experiences any of the symptoms of concussion. It is not unusual for symptoms to last 2 weeks in adults and 4 weeks in children. Fewer than 10% of sports-related concussions among children are associated with loss of consciousness.

<span class="mw-page-title-main">Turbellaria</span> Class of flatworms

The Turbellaria are one of the traditional sub-divisions of the phylum Platyhelminthes (flatworms), and include all the sub-groups that are not exclusively parasitic. There are about 4,500 species, which range from 1 mm (0.039 in) to large freshwater forms more than 500 mm (20 in) long or terrestrial species like Bipalium kewense which can reach 600 mm (24 in) in length. All the larger forms are flat with ribbon-like or leaf-like shapes, since their lack of respiratory and circulatory systems means that they have to rely on diffusion for internal transport of metabolites. However, many of the smaller forms are round in cross section. Most are predators, and all live in water or in moist terrestrial environments. Most forms reproduce sexually and with few exceptions all are simultaneous hermaphrodites.

<span class="mw-page-title-main">Platyzoa</span> Phylum of unsegmented animals

The paraphyletic "Platyzoa" are a group of protostome unsegmented animals proposed by Thomas Cavalier-Smith in 1998. Cavalier-Smith included in Platyzoa the phylum Platyhelminthes, and a new phylum, the Acanthognatha, into which he gathered several previously described phyla of microscopic animals. Later it has been described as paraphyletic, containing the Rouphozoa and the Gnathifera.

<span class="mw-page-title-main">Scalidophora</span> Proposed taxonomic clade

Scalidophora is a group of marine pseudocoelomate protostomes that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous cuticle that is moulted, rings of scalids on the introvert, flosculi, and two rings of introvert retracts. However, the monophyly of the Scalidophora was not supported by two molecular studies, where the position of the Loricifera was uncertain or as sister to the Panarthropoda. Both studies supported a reduced Scalidophora comprising the Kinorhyncha and Priapulida as sister phyla. Their closest relatives are the Panarthropoda, Nematoda and Nematomorpha; thus, they are placed in the group Ecdysozoa.

<span class="mw-page-title-main">Viridiplantae</span> Clade of archaeplastids including green algae and the land plants

Viridiplantae are a clade of eukaryotic organisms that comprise approximately 450,000–500,000 species and play important roles in both terrestrial and aquatic ecosystems. They are made up of the green algae, which are primarily aquatic, and the land plants (embryophytes), which emerged from within them. Green algae traditionally excludes the land plants, rendering them a paraphyletic group. However it is accurate to think of land plants as a kind of algae. Since the realization that the embryophytes emerged from within the green algae, some authors are starting to include them. They have cells with cellulose in their cell walls, and primary chloroplasts derived from endosymbiosis with cyanobacteria that contain chlorophylls a and b and lack phycobilins. Corroborating this, a basal phagotroph archaeplastida group has been found in the Rhodelphydia.

<span class="mw-page-title-main">Pancrustacea</span> Clade comprising all crustaceans and hexapods

Pancrustacea is the clade that comprises all crustaceans, including the cladistically included hexapods. This grouping is contrary to the Atelocerata hypothesis, in which Hexapoda and Myriapoda are sister taxa, and Crustacea are only more distantly related. As of 2010, the Pancrustacea taxon is considered well accepted, with most studies recovering Hexapoda within Crustacea. The clade has also been called Tetraconata, referring to having four cone cells in the ommatidia. This name is preferred by some scientists as a means of avoiding confusion with the use of "pan-" to indicate a clade that includes a crown group and all of its stem group representatives.

<span class="mw-page-title-main">History of life</span> Processes by which organisms evolved on Earth

The history of life on Earth traces the processes by which living and fossil organisms evolved, from the earliest emergence of life to present day. Earth formed about 4.5 billion years ago and evidence suggests that life emerged prior to 3.7 Ga. Although there is some evidence of life as early as 4.1 to 4.28 Ga, it remains controversial due to the possible non-biological formation of the purported fossils.

<span class="mw-page-title-main">Arthropod</span> Phylum of invertebrates with jointed exoskeletons

Arthropods are invertebrate animals in the phylum Arthropoda. They possess an exoskeleton with a cuticle made of chitin, often mineralised with calcium carbonate, a segmented body, and paired jointed appendages. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. They are an extremely diverse group, with up to 10 million species.

<span class="mw-page-title-main">Nematode</span> Phylum of worms with tubular digestive systems with openings at both ends

The nematodes or roundworms constitute the phylum Nematoda, with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broad range of environments. Less formally, they are categorized as helminths, but are taxonomically classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa, and unlike flatworms, have tubular digestive systems with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum.

<span class="mw-page-title-main">Crustacean</span> Subphylum of arthropods

Crustaceans belong to the subphylum Crustacea,, and form a large, diverse group of arthropods including decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. Some crustaceans are more closely related to insects and the other hexapods than they are to certain other crustaceans.

References

  1. Schmidt-Rhaesa, A.; Bartolomaeus, T.; Lemburg, C.; Ehlers, U.; Garey, J.R. (1998). "The position of the Arthropoda in the phylogenetic system - Articulata". J. Morphol. (238): 263–285. Archived from the original on 10 May 2010.
  2. Telford MJ, Bourlat SJ, Economou A, Papillon D, Rota-Stabelli O (April 2008). "The evolution of the Ecdysozoa". Philos. Trans. R. Soc. Lond. B Biol. Sci. 363 (1496): 1529–37. doi:10.1098/rstb.2007.2243. PMC   2614232 . PMID   18192181.
  3. Dunn, et al., 2008; Dunn, CW; Hejnol, A; Matus, DQ; Pang, K; Browne, WE; Smith, SA; Seaver, E; Rouse, GW; et al. (2008). "Broad phylogenomic sampling improves resolution of the animal tree of life". Nature 452 (7188): 745–749.
  4. "A revised six-kingdom system of life" (PDF). Archived from the original (PDF) on 2016-03-04. Retrieved 2016-02-27.
  5. The neuromuscular system of Pycnophyes kielensis (Kinorhyncha: Allomalorhagida) investigated by confocal laser scanning microscopy - EvoDevo
  6. "Explanations.html". Archived from the original on 2013-02-07. Retrieved 2009-06-28.