Dipterostemon | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Order: | Asparagales |
Family: | Asparagaceae |
Subfamily: | Brodiaeoideae |
Genus: | Dipterostemon Rydb. [1] |
Species: | D. capitatus |
Binomial name | |
Dipterostemon capitatus | |
Synonyms [2] | |
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Dipterostemon is a monotypic genus of flowering plants in the family Asparagaceae. Its only species is Dipterostemon capitatus, synonym Dichelostemma capitatum, [1] known by the common names blue dicks, wild hyacinth, [3] purplehead and brodiaea (alternately spelled brodiea and brodeia [4] ), native to the Western United States (particularly Arizona, California, Oregon, Utah, and New Mexico) and northwest Mexico. [5]
Dipterostemon capitatus is an herbaceous perennial growing from an underground corm to a height of as much as 60 cm. It has 2–3 leaves which are 10–40 cm long. The inflorescence is head- or umbel-like, and dense. It usually contains 2 to 15 flowers, which have a blue, blue-purple, pink-purple, or white perianth. The flower tube is 3–12 mm and is narrowly cylindrical to campanulate. Flowers have six fertile stamens, deeply notched, lanceolate, white, angled inward, slightly reflexed at tip, with outer filaments wider at the base. It has a twisted and fleshy peduncle, a set of membranous, petal-like stamen appendages around the anthers, and angular black seeds. It reproduces from seed and vegetative means in the form of cormlets. The cormlets are attached to the parent corm by stolons and are sessile, produced in the axils of the old leaf bases on the mature corm. [6] Plants thrive in open disturbed environments, and are a common post-fire succession species in chaparral. Flowering peaks in March. [7]
D. capitatus occurs from sea level up to 2,300 meters. It inhabits a wide variety of plant communities, including vernal pools, valley grassland, scrub, coniferous forests, and open woodlands. It seems not to colonize after fire by seed, but rather by cormlets. After fire, plants are exposed to unshaded environments with little brush competition, and vigorously flower in open environments with increased soil nutrients.
Grasslands that have been burned may exhibit thousands of plants where none have appeared in recent years. Corms may sit for a decade or more and wait for fire or other favorable environmental conditions before breaking ground. Suppression of fire may cause increased shade and plant competition and decrease population numbers of D. capitatus.
The species was first described by George Bentham in 1857 as Brodiaea capitata. [8] In 1868, Alphonso Wood moved it to the genus Dichelostemma as Dichelostemma capitatum. [9] In 1912, Per Axel Rydberg moved it to Dipterostemon as Dipterostemon capitatus. [10] Most sources continued to use Wood's placement in Dichelostemma, including the Flora of North America, which however noted that it was "quite distinct from other species in the genus" and that it was perhaps "best treated as its own genus". [11] In 2017, restoration of Dipterostemon as a monotypic genus was proposed. [12] This has been accepted by sources such as Plants of the World Online [1] and the Jepson Flora Project. [13]
Three subspecies are accepted: [2]
Corms have been gathered by Native Americans in California, parts of the Great Basin, and the Southwest. These corms were an important starch source in their diet. California tribes dug and continue to dig the corms before flowering, during flowering, or after seeding depending on the tribe and individual family. Traditional gathering sites were visited annually, and there were gathering tracts with different kinds of corms and bulbs owned and maintained by particular families.
Corms are eaten by animals such as black bears, mule deer, non-native wild pigs, rabbits, and pocket gophers. As some of the corms are eaten, others are dispersed. The animals detach the cormlets, which aerates the soil, prepares the seedbed, thins the plant population, and leaves some cormlets behind. Once separated, corms may take less time to reach flowering size.
Although slow to flowering starting from seeds, it is sometimes used in horticulture. The flowers mix well in native beds, especially when contrasted against other California native species such as California poppies ( Eschscholzia californica ). With patience and proper care, these plants can become quite dramatic come spring.
Indigenous people had several different types of management activities to ensure future corm production at gathering sites:
Periodic digging and thinning of the corms or separating the cormlets, and replanting them may have enhanced plant numbers and densities. Digging corms acts as a form of tillage, which will increase the size of the gathering tract, aerate the soil, lower weed competition, and prepare the seedbed to increase seed germination rates. Dipterostemon capitatus populations require periodic disturbance to maintain and increase their populations; therefore, indigenous harvesting regimes may help maintain populations. Populations that become overcrowded and show reduced vigor can be divided and separated.
Seeds sown in the fall usually readily germinate and do not need special treatment. If planting seeds, they will take several years to reach flowering size. Propagation by corms is much easier.
Corm, bulbo-tuber, or bulbotuber is a short, vertical, swollen underground plant stem that serves as a storage organ that some plants use to survive winter or other adverse conditions such as summer drought and heat (perennation).
Brodiaea, also known by the common name cluster-lilies, is a monocot genus of flowering plants.
Triteleia is a genus of monocotyledon flowering plants also known as triplet lilies. The 16 species are native to western North America, from British Columbia south to California and east to Wyoming and Arizona, with one species in northwestern Mexico. However, they are most common in California. They are perennial plants growing from a fibrous corm roughly spherical in shape. They get their name from the fact that all parts of their flowers come in threes.
Brodiaeoideae are a monocot subfamily of flowering plants in the family Asparagaceae, order Asparagales. They have been treated as a separate family, Themidaceae. They are native to Central America and western North America, from British Columbia to Guatemala. The name of the subfamily is based on the type genus Brodiaea.
Dichelostemma is a genus of North American plants closely related to the genus Brodiaea and sometimes regarded as part of that group.
Frangula californica is a species of flowering plant in the buckthorn family native to western North America. It produces edible fruits and seeds. It is commonly known as California coffeeberry and California buckthorn.
Dichelostemma congestum is a species of flowering plant known by the common name ookow or fork-toothed ookow. It is native to California, Oregon and Washington.
Dichelostemma volubile is a species of flowering plant known by the common names twining snakelily and twining brodiaea. This wildflower is endemic to the mountain foothills of California, where it grows in scrub and woodland.
Perityle is a genus of flowering plants in the daisy family. They are known generally as rock daisies.
Triteleia ixioides, known as prettyface or golden star, is a monocotyledon flowering plant in the genus Triteleia. It is native to northern and central California and southwestern Oregon, where it can be found in coastal and inland coniferous forests and other habitat. It is a perennial wildflower growing from a corm. It produces one to two basal leaves up to 50 centimeters long by 1.5 wide. The inflorescence arises on an erect stem up to 80 centimeters tall. It is an umbel-like cluster of several flowers each borne on a pedicel up to 7 centimeters long. The flowers are variable in size, measuring one to nearly three centimeters in length. They are pale to bright yellow, or sometimes purple-tinged white. There are six tepals with darker midveins in shades of green, brown, or purple. The lobes are funnel-shaped and may open flat or somewhat reflexed. The six stamens form a fused tube that protrudes from the corolla; they have broad, flat filaments and whitish, yellowish, or blue anthers.
Hooveria purpurea is a species of flowering plant related to the agaves known by the common name purple amole. This species of soap plant is endemic to California, where it grows in the Santa Lucia Range, in the Central Coast region. There are two varieties of this plant, and both are believed to be quite rare. It is a federally listed threatened species.
Dudleya brevifolia, is a rare succulent plant known by the common name short-leaved liveforever, short-leaved dudleya or rarely the Del Mar Hasseanthus. It is an edaphic endemic that only grows on Lindavista formation marine terraces, on surfaces with ironstone nodules. The leaves are deciduous, and disappear after the inflorescence develops. The small white flowers are star-shaped with a yellow center. After flowering, any above ground trace of the plant will disappear, and it survives under the earth with a starch-rich subterranean caudex. Dudleya brevifolia is only found on coastal mesas along a small strip of coast in San Diego County, California.
Sambucus racemosa is a species of elderberry known by the common names red elderberry and red-berried elder.
Brodiaea coronaria is the type species of Brodiaea and also known by the common names harvest brodiaea and crown brodiaea. It is native to western North America from British Columbia to northern California, where it grows in mountains and grasslands.
Brodiaea filifolia, known by the common name threadleaf brodiaea, is a rare species of flowering plant in the cluster-lily genus. It is endemic to southern California, mostly in the region around the junction of Orange, Riverside, and San Diego Counties.
Brodiaea orcuttii is a species of flowering plant in the family Asparagaceae, subfamily Brodiaeoideae. It is a cluster-lily known by the common name Orcutt's brodiaea. The bulb is native to Southern California, mainly San Diego County, where it is an uncommon species. Its range probably extends into Baja California.
Brodiaea stellaris is a species of flowering plant in the cluster-lily genus known by the common name starflower brodiaea.
Triteleia hyacinthina is a species of flowering plant known by the common names white brodiaea, white tripletlily, hyacinth brodiaea, and fool's onion. It is native to western North America from British Columbia to Idaho to central California. Its habitat includes grassland and vernally moist areas such as meadows and vernal pools. It is a perennial herb growing from a corm. It produces two or three basal leaves up to 40 centimeters (16 in) long by 2 centimeters (0.79 in) wide. The inflorescence arises on an erect stem up to 60 centimeters (24 in) tall and bears an umbel-like cluster of many flowers. Each flower is a funnel-shaped bloom borne on a pedicel up to 5 centimeters (2.0 in) long. The flower is white, often tinged purple along the tubular throat, with six green-veined tepals. There are six stamens with white, yellow, or occasionally blue anthers.
Watsonia meriana is a species of flowering plant in the family Iridaceae known by the common name bulbil bugle-lily. It is one of several Watsonia species known as wild watsonia. It is native to the Cape Provinces of South Africa, but it is well known as an ornamental plant grown in gardens for its showy spikes of flowers, and is an invasive species in areas where it has escaped cultivation.
Acmispon is a genus of flowering plant in the family Fabaceae (legumes), native to North America and the west coast of Chile in South America. It includes several species of American bird's-foot trefoils and deervetches formerly contained in the globally distributed genus Lotus. The former genus Syrmatium is included in Acmispon. The Jepson eFlora accepts only Acmispon.