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Eucladoceros Temporal range: | |
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Skull of Eucladoceros dicranios | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Family: | Cervidae |
Subfamily: | Cervinae |
Genus: | † Eucladoceros Falconer, 1868 |
Species | |
| |
Synonyms | |
PolycladusPomel, 1854 [1] |
Eucladoceros (Greek for "well-branched antler") is an extinct genus of large deer whose fossils have been discovered across Eurasia, from Europe to China, spanning from the Late Pliocene-Early Pleistocene. [2] It is noted for its unusual comb-like or branching antlers.
Species of Eucladoceros were large-sized deer. European species lie E. dicranios and E. ctenoides are suggested to have reached a body mass of 250–300 kilograms (550–660 lb), [3] while East Asian E. boulei is suggested to have reached body masses of 350 kilograms (770 lb). [2] Body size of European species increased over time. [4] E. giulii has been estimated to have had a shoulder height of 1.55 metres (5.1 ft) while E. senezensis has an estimated shoulder height of 1.7 metres (5.6 ft). [5] Species of Eucladoceros are noted for their branching antlers, with a large number of tines projecting from the front part of the main antler beam. [6] In many species like Eucladoceros ctenoides the antlers have a comb-like branching pattern, while those of the type species Eucladoceros dicranios has a more complex dichotomous branching pattern. [4] The antlers of E. dicranios are proporotionally large relative to body size, and are among the largest antlers known among deer. [7] The teeth of Eucladoceros species are similar in some aspects to those of the genus Cervus , but lack certain derived characters typical of the teeth of that genus. [6]
The first find (E. dicranios) was in 1841 by Florentine naturalist Filippo Nesti, director of the "Museo di Storia Naturale di Firenze" ("Museum of Natural History of Florence"). The earliest species of Eucladoceros was described from the Early Pliocene of China. The most abundant fossil remains of Eucladoceros have come from the Early Pleistocene of Europe and China.
The systematics of European forms is confused and up to twelve poorly defined species are reported. The majority of those species names are synonymous, and at present only two or three good species are recognized: E. dicranios from England, Italy and Azov Sea Area in South Russia; E. ctenoides from Greece, Italy, France, Spain, the Netherlands, and England; and E. teguliensis (a senior synonym of E. senezensis) from France, the Netherlands, and Britain. The latter species is regarded by some authors as a subspecies of E. ctenoides, since there are some finds (for instance, from Ceyssaguet, France) that show a transitional character between E. ctenoides and E. teguliensis.
Some poor remains of Eucladoceros are found also in Tajikistan, Pakistan, and India.
Analysis of the limbs of Eucladoceros suggest that they are most similar to living deer that occupy open habitats. [8] Dental microwear analysis of Eucladoceros ctenoides suggests that its diet was largely plastic and widely varied according to local conditions. [9]
The genus was formally described by Hugh Falconer in 1868. [10]
Eucladoceros is widely agreed to a be member of the tribe Cervini, though its placement within this group has been debated. A 2017 study analysing the bony labyrinth suggested that Eucladoceros was most closely related to the living genera Rusa (sambar) and Cervus (red deer, elk, sika deer). [11] Some authors have proposed a close relationship to the extinct "giant deer" genus Praemegaceros . [12]
A deer or true deer is a hoofed ruminant ungulate of the family Cervidae. Cervidae is divided into subfamilies Cervinae and Capreolinae. Male deer of almost all species, as well as female reindeer, grow and shed new antlers each year. These antlers are bony extensions of the skull and are often used for combat between males.
The Irish elk, also called the giant deer or Irish deer, is an extinct species of deer in the genus Megaloceros and is one of the largest deer that ever lived. Its range extended across Eurasia during the Pleistocene, from Ireland to Lake Baikal in Siberia. The most recent remains of the species have been radiocarbon dated to about 7,700 years ago in western Russia. Its antlers, which can span 3.5 metres (11 ft) across are the largest known of any deer. It is not closely related to either living species called the elk, with it being widely agreed that its closest living relatives are fallow deer (Dama).
Megaloceros is an extinct genus of deer whose members lived throughout Eurasia from the Pleistocene to the early Holocene. The type and only undisputed member of the genus, Megaloceros giganteus, vernacularly known as the "Irish elk" or "giant deer", is also the best known. Fallow deer are thought to be their closest living relatives. Megaloceros has been suggested to be closely related to other genera of "giant deer", like the East Asian genus Sinomegaceros, and the European Praemegaceros.
The Cervinae or the Old World deer, are a subfamily of deer. Alternatively, they are known as the plesiometacarpal deer, due to having lost the parts of the second and fifth metacarpal bones closest to the foot, distinct from the telemetacarpal deer of the Capreolinae.
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The Capreolinae, Odocoileinae, or the New World deer are a subfamily of deer. Alternatively, they are known as the telemetacarpal deer, due to their bone structure being different from the plesiometacarpal deer subfamily Cervinae. The telemetacarpal deer maintain their distal lateral metacarpals, while the plesiometacarpal deer maintain only their proximal lateral metacarpals. The Capreolinae are believed to have originated in the Middle Miocene, between 7.7 and 11.5 million years ago, in Central Asia.
Xenocyon is an extinct group of canids, either considered a distinct genus or a subgenus of Canis. The group includes Canis (Xenocyon) africanus, Canis (Xenocyon) antonii and Canis (Xenocyon) falconeri that gave rise to Canis (Xenocyon) lycanoides. The hypercarnivorous Xenocyon is thought to be closely related and possibly ancestral to modern dhole and the African wild dog, as well as the insular Sardinian dhole.
Odocoileus lucasi, known commonly as the American mountain deer, is an extinct species of North American deer.
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The Finnish forest reindeer(Rangifer fennicus fennicus, also known as Eurasian or European forest reindeer is a rare subspecies of the reindeer native to Finland and northwestern Russia. They are found primarily in Russian Karelia and the provinces of North Karelia, Savonia and Kainuu in Finland, though some range into central south Finland. They are distinct from the semi-domesticated mountain reindeer in their larger size, longer legs and preference for dense boreal forest habitat, where they are rarely seen by humans, over the open tundra. The Finnish herd migrates seasonally back and forth across the long Russo-Finnish border.
Rucervus is a genus of deer from India, Nepal, Indochina, and the Chinese island of Hainan. The only extant representatives, the barasingha or swamp deer and Eld's deer, are threatened by habitat loss and hunting; another species, Schomburgk’s deer, went extinct in 1938. Deer species found within the genus Rucervus are characterized by a specific antler structure, where the basal ramification is often supplemented with an additional small prong, and the middle tine is never present. The crown tines are inserted on the posterior side of the beam and may be bifurcated or fused into a small palmation.
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Cervavitus is a genus of prehistoric deer that lived from the late Miocene to the Early Pleistocene in parts of Western and Eastern Europe, Central Asia and China.
Praemegaceros is an extinct genus of deer, known from the Pleistocene and Holocene of Western Eurasia. Praemegaceros is considered to be a genus of "giant deer", with many species having an estimated body mass of around 400 kilograms (880 lb), considerably larger than most living deer.
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Sinomegaceros is an extinct genus of deer known from the Late Pliocene/Early Pleistocene to Late Pleistocene of Central and East Asia. It is considered to be part of the group of "giant deer", with a close relationship to Megaloceros. Many members of the genus are noted for their distinctive palmate antler brow tines.
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