Hordeum murinum | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Clade: | Commelinids |
Order: | Poales |
Family: | Poaceae |
Subfamily: | Pooideae |
Genus: | Hordeum |
Species: | H. murinum |
Binomial name | |
Hordeum murinum | |
Subspecies | |
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Hordeum murinum is a species of flowering plant in the grass family Poaceae, commonly known as wall barley or false barley. It is a close relative of cultivated barley (H. vulgare).
Hordeum murinum complex is the most widespread of all Hordeum species. [1] The center of distribution of H. murinum is in the Mediterranean area, Central Europe, Western Asia, and North Africa. [1] Hordeum murinum is quite widespread and common. It flowers during May through July in mainly coastal areas. It is an ancient introduction in the British Isles, common in England and Wales but less common in Scotland and Ireland . [2]
It can grow to 30 cm in height, and its unbranched spikes can reach 10 cm long. It produces small, dry nutlets and its leaves can be 8 millimetres (5⁄16 in) wide with short, blunt ligules. It is an annual winter species whose seeds germinate and develop in the spring. [3] It is also referred to as wall barley and are tetraploids. [4] It is distinct from other species of the genus because of its morphology and molecular genetics. [1] It is also distinct because of the barriers it has with the Hordeum taxa when it comes to its ability to cross with other species. [1]
Among its subspecies is included H. m. ssp. leporinum, known as hare barley.
Precipitation is the most important factor in the production of seeds for this species. A greater quantity of dry material is produced with medium precipitation and better distribution. [3] In drier years with early or late rainfalls, there is no chance of re-seeding for this species. [4] The species uses a greater part of its reproductive resources for seed production, allowing it to adapt to different water conditions. [3] Controlling the seeding rate favors high-quality strand of barley. The sowing rate for wall barley increases when seed production and forage increases. [5] This helps to obtain ideal and sustainable forage and seed yield in rangelands of Jordan. The height of the plant and protein content does not respond to seeding rates, but the height of the plant and protein content does vary with years. [5] Anatomical characteristics of leaf blades differ between the taxa.[ citation needed ] This weedy species along with hare barley and smooth barley can be hard to control. [4]
There are 3 subspecies:
Hordeum murinum ssp. leporinum, known as hare barley, [6] mouse barley, [7] and barley grass. [8] This subspecies grows in tufts from 10 to 40 cm (4 to 16 in) in height, [8] [9] and its flowers are attached to branches rather than to the main axis.[ citation needed ] It is native to the Mediterranean region near continental, oceanic, and colder climates, [1] as well as northern Africa and temperate Asia, and it is widely naturalised elsewhere. [7] It was first published as the full species H. leporinum by Johann Heinrich Friedrich Link in 1834. In 1882 it was redescribed as a subspecies of H. murinum by Giovanni Arcangeli, though today some authorities maintain it at the species level. [8] [9] Jakob & Blattner 2009 find cytotypes of both tetraploid and hexaploid. [10]
Another subspecies is 'Hordeum murinum ssp. glaucum. [1] Jakob & Blattner 2009 find it is diploid. [10] It appears in warmer climates of the Mediterranean region.
The above subspecies differ primarily because of their chromosome numbers, spikelet morphology, and geographical distribution. [1] H. leporinum is more dominant in areas where the rainfall is greater than 425 mm (17 in). [4] H. glaucum is more dominant in semiarid regions where rainfall is less than that. [4]
The last is the tetraploid Hordeum murinum ssp. murinum. [10]
Jakob & Blattner 2009 find the diploid subspecies H. m. ssp. glaucum is the only extant parent contributing to contemporary H. m. populations. [10] They find that another species of the same group – an extinct member of the Xu group – combined with it to form the tetraploids, and another unknown extinct species then contributed to the hexaploids. [10]
Although H. murinum is considered a difficult weed in cereal crop fields, [4] it is useful for feeding grazing animals. [4] It is also the main source of forage for cattle production in areas with water deficits. [3]
In England in the late 20th century among children the plant was referred to colloquially as the 'Flea Dart', from the aerodynamic shape of its seedhead, and the aphids that are often present within it in its immature state. [11] In China the subspecies leporinum is a common ingredient in the spring Qingming Festival snack qīngtuán .
Ploidy is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively, in each homologous chromosome pair, which chromosomes naturally exist as. Somatic cells, tissues, and individual organisms can be described according to the number of sets of chromosomes present : monoploid, diploid, triploid, tetraploid, pentaploid, hexaploid, heptaploid or septaploid, etc. The generic term polyploid is often used to describe cells with three or more sets of chromosomes.
Hordeum is a genus of annual and perennial plants in the grass family. They are native throughout the temperate regions of Africa, Eurasia, and the Americas.
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Spelt, also known as dinkel wheat or hulled wheat, is a species of wheat that has been cultivated since approximately 5000 BCE.
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During 10,000 years of cultivation, numerous forms of wheat, many of them hybrids, have developed under a combination of artificial and natural selection. This diversity has led to much confusion in the naming of wheats. Genetic and morphological characteristics of wheat influence its classification; many common and botanical names of wheat are in current use.
Dactylis glomerata is a species of flowering plant in the grass family Poaceae, commonly known as cock's-foot, orchard grass, or cat grass (due to its popularity for use with domestic cats). It is a cool-season perennial C3 bunchgrass native throughout most of Europe, temperate Asia, and northern Africa.
Triticeae is a botanical tribe within the subfamily Pooideae of grasses that includes genera with many domesticated species. Major crop genera found in this tribe include wheat, barley, and rye; crops in other genera include some for human consumption, and others used for animal feed or rangeland protection. Among the world's cultivated species, this tribe has some of the most complex genetic histories. An example is bread wheat, which contains the genomes of three species with only one being a wheat Triticum species. Seed storage proteins in the Triticeae are implicated in various food allergies and intolerances.
Hordeum jubatum, with common names foxtail barley, bobtail barley, squirreltail barley, and intermediate barley, is a perennial plant species in the grass family Poaceae. It occurs wild mainly in northern North America and adjacent northeastern Siberia. However, as it escaped often from gardens it can be found worldwide in areas with temperate to warm climates, and is considered a weed in many countries. The species is a polyploid and originated via hybridization of an East Asian Hordeum species with a close but extinct relative of Californian H. brachyantherum. It is grown as an ornamental plant for its attractive inflorescences and when done flowering for its inflorescence.
Hordeum pusillum, also known as little barley, is an annual grass native to most of the United States and southwestern Canada. It arrived via multiple long-distance dispersals of a southern South American species of Hordeum about one million years ago. Its closest relatives are therefore not the other North American taxa like meadow barley or foxtail barley, but rather Hordeum species of the Pampas of central Argentina and Uruguay. It is less closely related to the Old World domesticated barley, from which it diverged about 12 million years ago. It is diploid.
Hordeum brachyantherum, known by the common name meadow barley, is a species of barley. It is native to western North America from Alaska to northern Mexico, coastal areas of easternmost Russia (Kamchatka), and a small area of coastal Newfoundland.
Barley, a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikelets and making it much easier to harvest. Its use then spread throughout Eurasia by 2000 BC. Barley prefers relatively low temperatures to grow, and well-drained soil. It is relatively tolerant of drought and soil salinity, but is less winter-hardy than wheat or rye.
Chloris gayana is a species of grass known by the common name Rhodes grass. It is native to Africa but it can be found throughout the tropical and subtropical world as a naturalized species.
Hordeum intercedens is a diploid, annual species of wild barley known by the common names bobtail barley and vernal barley. It is native to southern California and northern Baja California, where it is an increasingly rare member of the flora in saline and alkaline soils near seasonal waterflows and vernal pool habitats. Today most occurrences are located on the Channel Islands of California; many of the occurrences known from the mainland have been extirpated in the process of land development. This is an annual grass growing erect to bent in small tufts with stems up to 40 centimeters long. The inflorescence is a green spike up to 6.5 centimeters long made up of awned spikelets between 1 and 2 centimeters long.
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