Incurvarioidea | |
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A longhorn moth, Nemophora degeerella | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Superorder: | Amphiesmenoptera |
Order: | Lepidoptera |
Suborder: | Glossata |
Infraorder: | Heteroneura |
Superfamily: | Incurvarioidea |
Families | |
Adelidae Contents | |
Diversity | |
About 600 species |
Incurvarioidea is a superfamily of primitive monotrysian moths in the order Lepidoptera which consists of Leafcutters, yucca moths and relatives. This superfamily is characterised by a piercing, extensible ovipositor used for laying eggs in plants (Davis, 1999). Many species are day-flying with metallic patterns.
The Monotrysia are a group of insects in the lepidopteran order which are not currently considered to be a natural group or clade. The group consists only of moths and most of these are small and are relatively understudied in many regions of the world. The group is so named because the female has a single genital opening for mating and laying eggs, in contrast to the rest of the Lepidoptera (Ditrysia), which have two female reproductive openings. They comprise all of the group Heteroneura but the Ditrysia.
Lepidoptera is an order of insects that includes butterflies and moths. About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 per cent of the total described species of living organisms. It is one of the most widespread and widely recognizable insect orders in the world. The Lepidoptera show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, and is among the four most speciose orders, along with the Hymenoptera, Diptera, and Coleoptera.
The Prodoxidae are a family of moths, generally small in size and nondescript in appearance. They include species of moderate pest status, such as the currant shoot borer, and others of considerable ecological and evolutionary interest, such as various species of "yucca moths".
Schreckensteinioidea is a superfamily in the insect order Lepidoptera containing a single family, Schreckensteiniidae, or "bristle-legged moths", because of the stout spines on the hindlegs. The superfamily and family were both described by Thomas Bainbrigge Fletcher in 1929. The relationships of this family within the group apoditrysia are currently uncertain. One of the species, the blackberry skeletoniser, is widespread and common across Europe and has been introduced as a biological control to Hawaii, whilst three species of Corsocasis occur in South East Asia.
Neopseustidae is a small family of day and night-flying "archaic bell moths" in the order Lepidoptera. They are classified into their own superfamily Neopseustoidea and infraorder Neopseustina. Four genera are known. These primitive moths are restricted to South America and South east Asia and their biology is unknown. Nematocentropus appears to be the most primitive genus occurring in Assam, Myanmar and Sichuan, China, three species of Neopseustis are distributed from Assam to Taiwan, whilst Synempora andesae and three species of Apoplania occur in southern South America. The morphology of the antennae and the proboscis has been studied in detail.
Immoidea is a superfamily of pantropical moths containing only the family Immidae comprising ten genera with around 250 species, over half of them in the genus Imma. Many are brightly coloured and diurnal. The position of this group is currently uncertain within the group Obtectomera. The larvae feed on the leaves of dicotyledons and conifers including Podocarpus.
A superfamily of usually very small monotrysian moths that are characterised by small or large eyecaps over the compound eyes. Nepticuloidea comprises two families, the "pigmy moths" (Nepticulidae), with 12 genera which are very diverse worldwide and are usually leaf miners, and the "white eyecap moths" (Opostegidae), also worldwide but with five genera and about a ninth as many species, whose biology is less well known.
Palaephatoidea is a superfamily of insects in the order Lepidoptera with a single family, Palaephatidae with seven known genera. These "Gondwanaland moths" exhibit a disjunct distribution occurring mainly in South America, with four species in eastern Australia and Tasmania and one in South Africa. The larvae spin together leaves of Proteaceae (Ptyssoptera) or Verbenaceae (Azaleodes) . Palaephatoidea, a typical monotrysian group, is one two main candidates as the sister group of most of the Lepidoptera, the Ditrysia.
Tischerioidea is the superfamily of "trumpet" leaf miner moths. The superfamily contains just one family, Tischeriidae, and traditionally one genus, Tischeria, but currently three genera are recognised, widespread across the world including South America, except for Australasia. This is one candidate as the sister group of the bulk of Lepidoptera, the Ditrysia, and they have a monotrysian type of female reproductive system. These small moths are leaf-miners in the caterpillar stage, feeding mainly on Fagaceae, Asteraceae, and Malvaceae (Astrotischeria), and some also on Rhamnaceae, Tiliaceae, and Rosaceae.
Nepticulidae is a family of very small moths with a worldwide distribution. They are characterised by eyecaps over the eyes. These pigmy moths or midget moths, as they are commonly known, include the smallest of all living moths, with a wingspan that can be as little as 3 mm in the case of the European pigmy sorrel moth, but more usually 3.5–10 mm. The wings of adult moths are narrow and lanceolate, sometimes with metallic markings, and with the venation very simplified compared to most other moths.
Incurvariidae is a family of small primitive monotrysian moths in the order Lepidoptera. There are twelve genera recognised. Many species are leaf miners and much is known of their host plants, excluding Paraclemensia acerifoliella. The most familiar species in Europe are perhaps Incurvaria masculella and Phylloporia bistrigella. The narrow wings are held tightly along the body at rest and some species have very long antennae.
The Adelidae or fairy longhorn moths are a family of monotrysian moths in the lepidopteran infraorder Heteroneura. Most species have at least partially metallic patterns coloration and are diurnal, sometimes swarming around the tips of branches with an undulating flight. Others are crepuscular and have a drab coloration. Fairy longhorn moths have a wingspan of 4–28 millimeters, and males often have especially long antennae, 1–3 times as long as the forewing.
Cecidosidae is a little-known family of primitive monotrysian moths in the order Lepidoptera which have a piercing ovipositor used for laying eggs in plant tissue in which they induce galls, or they mine in bark. Nine species occur in southern Africa, five species in South America and Xanadoses nielseni was recently described from New Zealand. Some minute parasitoid wasps are known.
Carposinidae, the "fruitworm moths", is a family of insects in the order Lepidoptera. These moths are narrower winged than Copromorphidae, with less rounded forewing tips. Males often have conspicuous patches of scales on either surface. The mouthparts are quite diagnostic, usually with prominent, upcurved "labial palps", the third segment long, and the second segment covered in large scales. Unlike Copromorphidae, the "M2" and sometimes "M1" vein on the hindwings is absent. The relationship of Carposinidae relative to Copromorphidae needs further investigation. It is considered possible that the family is artificial, being nested within Copromorphidae. The Palearctic species have been revised by Alexey Diakonoff (1989).
Crinopteryx is a monotypic genus of moths and the sole genus of the also monotypic family Crinopterygidae. Its sole species, Crinopteryx familiella is a primitive monotrysian moth endemic to Europe, where it is restricted to the Mediterranean region of France, Italy, the Iberian Peninsula and Sicily. Crinopterygidae is not the only moth family restricted to this region.
Copromorphoidea, the "fruitworm moths" is a superfamily of insects in the lepidopteran order. These moths are small to medium-sized and are broad-winged bearing some resemblance to the superfamilies Tortricoidea and Immoidea. The antennae are often "pectinate" especially in males, and many species of these well camouflaged moths bear raised tufts of scales on the wings and a specialised fringe of scales at the base of the hindwing sometimes in females only; there are a number of other structural characteristics. The position of this superfamily is not certain, but it has been placed in the natural group of "Apoditrysia" "Obtectomera", rather than with the superfamilies Alucitoidea or Epermenioidea within which it has sometimes previously been placed, on the grounds that shared larval and pupal characteristics of these groups have probably evolved independently. It has been suggested that the division into two families should be abandoned.
A family of primitive monotrysian moths in the order Lepidoptera, Heliozelidae are small, metallic day-flying moths with shiny smooth heads. In Europe the small adult moths are seldom noticed as they fly quite early in the spring. The larvae are leaf miners and the vacated leaf mines are distinctive because the larva leaves a large hole at the end.
Opostegidae or "white eyecap moths" is a family of insects in the order Lepidoptera that is characterised by particularly large eyecaps over the compound eyes. Opostegidae are most diverse in the New World tropics.
Acanthoctesia or "archaic sun moths" is an infraorder of insects in the lepidopteran order, containing a single superfamily, Acanthopteroctetoidea, and a single family, Acanthopteroctetidae. They are currently considered the fifth group up on the comb of branching events in the extant lepidopteran phylogeny. They also represent the most basal lineage in the lepidopteran group Coelolepida characterised in part by its scale morphology. Moths in this superfamily are usually small and iridescent. Like other "homoneurous" Coelolepida and non-ditrysian Heteroneura, the ocelli are lost. There are variety of unique structural characteristics. There are two described genera of these primitive moths. Catapterix was originally described within its own family but Acanthopteroctetes shares with it a number of specialised structural features including similar wing morphology.
Andesiana is a genus representing its own family Andesianidae and superfamily Andesianoidea, the "Andean endemic moths". It contains three species with a wingspan up to 5.4 cm. in female A. similis and 3.5 cm. in males. This far surpasses in size any previously known monotrysian moth. These large Microlepidoptera are restricted to Andean South America, from where they were described originally in 1989 in the family Cossidae by their discoverer Patricia Gentili.
Cimeliidae, the gold moths, is a family of moths whose precise relationships within the Macrolepidoptera are currently uncertain, but they currently represent the only family in a recently recognized superfamily whose nearest relatives include the butterflies, Calliduloidea, Drepanoidea, Geometroidea, Bombycoidea, Mimallonoidea, Lasiocampoidea, and the Noctuoidea. Uniquely, they have a pair of pocket-like organs on the seventh abdominal spiracle of the adult moth which are only possibly sound receptive organs. They are quite large and brightly coloured moths that occur in southern Europe and feed on species of Euphorbia. Sometimes they are attracted to light. The family was first described by Pierre Chrétien in 1916.
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