| Siphonophorida Temporal range: | |
|---|---|
 | |
| Illacme plenipes | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Subphylum: | Myriapoda |
| Class: | Diplopoda |
| Subclass: | Chilognatha |
| Infraclass: | Helminthomorpha |
| Subterclass: | Colobognatha |
| Order: | Siphonophorida Newport, 1844 |
| Families | |
| Synonyms | |
Californizoniinae Verhoeff, 1941 | |
Siphonophorida (Greek for "tube bearer") is an order of millipedes containing two families and over 100 species.
Millipedes in the order Siphonophorida are long and worm-like, reaching up to 36 millimetres (1.4 in) in length and up to 190 body segments. Eyes are absent, and in many species the head is elongated into a long beak, with mandibles highly reduced. The beak may serve in a suctorial function. The body has a dense covering of fine setae. Each body segment consists of a dorsal tergite, two lateral pleurites, and ventral sternite, which are unfused. The male reproductive appendages (gonopods) are simple and leg-like, consisting of the ninth and 10th leg pairs. This lack of specialization has led to Siphonophorida being called a "taxonomist's nightmare", [1] and Jeekel (cited in [1] ) jokingly gave the order the "taxonomists' award for least popular group among diplopods". [2]
Siphonophorida occurs from southwestern USA to Brazil and Peru in the western hemisphere, as well as South Africa, India, southeast Asia and Australia. [3] The oldest fossil of the group is from the Late Cretaceous (Cenomanian) aged Burmese amber, assignable to the extant genus Siphonophora. [4]
Two families are traditionally recognized. Differences between the two families include antennae and head structure: the large family Siphonophoridae, with over 100 species, has elongate beak-like mouthparts, and straight antennae with sensory pits on the fifth and sixth antennal segments ("antennomeres)". The family Siphonorhinidae, with only around 10 species, lacks the beak-like mouthparts, and has geniculate (elbowed) antennae lacking sensory pits on the fifth and sixth antennomere. The siphonorhinid genus Nematozonium with two species, is sometimes placed in its own monotypic family, Nematozoniidae. [1]
Family Siphonophoridae Newport, 1844
Family Siphonorhinidae Cook, 1895
Centipedes are predatory arthropods belonging to the class Chilopoda of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (metameric) creatures with one pair of legs per body segment. All centipedes are venomous and can inflict painful bites, injecting their venom through pincer-like appendages known as forcipules. Despite the name, no centipede has exactly 100 pairs of legs; they can have a varying number of legs, ranging from 15 pairs to 191 pairs, always an odd number. They are predominantly carnivorous.
Passalidae is a family of beetles known variously as "bessbugs", "bess beetles", "betsy beetles" or "horned passalus beetles". Nearly all of the 500-odd species are tropical; species found in North America are notable for their size, ranging from 20–43 mm, for having a single "horn" on the head, and for a form of social behavior unusual among beetles.
Symphylans, also known as garden centipedes or pseudocentipedes, are soil-dwelling arthropods of the class Symphyla in the subphylum Myriapoda. Symphylans resemble centipedes, but are very small, non-venomous, and only distantly related to both centipedes and millipedes. They can move rapidly through the pores between soil particles, and are typically found from the surface down to a depth of about 50 centimetres (20 in). They consume decaying vegetation, but can do considerable harm in an agricultural setting by consuming seeds, roots, and root hairs in cultivated soil.
This glossary of entomology describes terms used in the formal study of insect species by entomologists.
Illacme plenipes is a siphonorhinid millipede found in the central region of the U.S. state of California. It has up to 750 legs. One of two known species in the genus Illacme, it was first seen in 1926, but was not rediscovered until 2005, almost 80 years after its discovery, by Paul Marek, then a Ph.D. student at East Carolina University.
Glomerida is an order of pill-millipedes found primarily in the Northern Hemisphere. Also known as northern pill millipedes, they superficially resemble pill-bugs or woodlice, and can enroll into a protective ball. They have twelve body segments, 17 to 19 pairs of legs, and males have enlarged rear legs involved in mating. The order includes about 30 genera and at least 280 species, including Glomeris marginata, the common European pill-millipede. The order contains members in Europe, South-east Asia and the Americas from California to Guatemala. Although historically considered closely related with the similar sphaerotheriidans that also enroll, some DNA evidence suggest they may be more closely related to glomeridesmidans, a poorly known order that does not enroll.
Spirostreptida is an order of long, cylindrical millipedes. There are approximately 1000 described species, making Spirostreptida the third largest order of millipedes after Polydesmida and Chordeumatida.
Sisyridae, commonly known as spongeflies or spongillaflies, are a family of winged insects in the order Neuroptera. There are approximately 60 living species described, and several extinct species identified from the fossil record.
Julida is an order of millipedes. Members are mostly small and cylindrical, typically ranging from 10–120 millimetres (0.39–4.72 in) in length. Eyes may be present or absent, and in mature males of many species, the first pair of legs is modified into hook-like structures. Additionally, both pairs of legs on the 7th body segment of males are modified into gonopods.
Zigrasimecia is an extinct genus of ants which existed in the Cretaceous period approximately 98 million years ago. The first specimens were collected from Burmese amber in Kachin State, 100 kilometres (62 mi) west of Myitkyina town in Myanmar. In 2013, palaeoentomologists Phillip Barden and David Grimaldi published a paper describing and naming Zigrasimecia tonsora. They described a dealate female with unusual features, notably the highly specialized mandibles. Other features include large ocelli, short scapes, 12 antennomeres, small eyes, and a clypeal margin that has a row of peg-like denticles. The genus Zigrasimecia was originally incertae sedis within Formicidae until a second species, Zigrasimecia ferox, was described in 2014, leading to its placement in the subfamily Sphecomyrminae. Later, it was considered to belong to the distinct subfamily Zigrasimeciinae.
Platydesmida is an order of millipedes containing two families and over 60 species. Some species practice paternal care, in which males guard the eggs.
Callipodida is an order of millipedes containing around 130 species, many characterized by crests or ridges.
Colobognatha is a clade of helminthomorph millipedes containing four orders: Platydesmida, Polyzoniida, Siphonocryptida, and Siphonophorida.
Siphoniulus is a poorly known genus of millipede containing only two living species: S. alba from Indonesia, and S. neotropicus from Mexico and Guatemala. An additional two fossil species are known from Cretaceous amber. Siphoniulus species are the only members of the family Siphoniulidae and order Siphoniulida, making Siphoniulida the smallest millipede order. Few specimens are known, and their classification is contentious, although most recent studies place them as basal members of the Helminthomorpha.
Siphonorhinidae is a family of millipede in the order Siphonophorida. There are at least 4 genera and about 12 described species in Siphonorhinidae.
Parastemmiulus is an extinct genus of millipede in the family Stemmiulidae known from a fossil found in Mexico. There is one described species in the genus, Parastemmiulus elektron. The species is one of three millipedes described from Mexican amber, and the oldest Stemmiulidae fossil species as of 2013.
Anbarrhacus is an extinct genus of millipede in the family Platyrhacidae known from a fossil found in North America. There is one described species in the genus, Anbarrhacus adamantis, which is one of three millipedes described from Mexican amber.
Camelomecia is an extinct genus of stem-group ants not placed into any Formicidae subfamily. Fossils of the single known species, Camelomecia janovitzi, are known from the Middle Cretaceous of Asia. The genus is one of several ants described from Middle Cretaceous ambers of Myanmar.
Illacme is a genus of millipedes in the family Siphonorhinidae. It includes two species. Illacme plenipes was first described in 1928 from San Benito County, California, and rediscovered in 2005. In 2016, Illacme tobini was described based on a single male specimen collected in 2006 from Lange Cave, in Sequoia National Park, 240 km (150 mi) east of the known habitat for I. plenipes.
Siphonophora albiceps is a species of millipede belonging to the family Siphonophoridae. The species is distinct for the contrast of its white front with its dark middle, its small body, and large middle joints in its first pair of legs. It was first described by Harold Loomis in 1970 and its type locality is in the U.S. Virgin Islands.
Media related to Siphonophorida at Wikimedia Commons
Data related to Siphonophorida at WikispeciesOrders of the class Diplopoda (millipedes) | |||||||||||||||||||||||||||||||
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