Song sparrow | |
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in Brooklyn, New York, USA | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Passeriformes |
Family: | Passerellidae |
Genus: | Melospiza |
Species: | M. melodia |
Binomial name | |
Melospiza melodia (Wilson, 1810) | |
Part of the range of M. melodia Breeding range Year-round range Wintering range | |
Synonyms | |
Melospiza cinerea |
The song sparrow (Melospiza melodia) is a medium-sized New World sparrow. Among the native sparrows in North America, it is easily one of the most abundant, variable and adaptable species.
Adult song sparrows have brown upperparts with dark streaks on the back and are white underneath with dark streaking and a dark brown spot in the middle of the breast. They have a brown cap and a long brown rounded tail. Their face is gray with a brown streak through each eye. They are highly variable in size across numerous subspecies (for subspecies details, see below). The body length ranges from 11 to 18 cm (4.3 to 7.1 in) and wingspan can range from 18 to 25.4 cm (7.1 to 10.0 in). [2] [3] Body mass ranges from 11.9 to 53 g (0.42 to 1.87 oz). [4] The average of all races is 32 g (1.1 oz) but the widespread nominate subspecies (M. m. melodia) weighs only about 22 g (0.78 oz) on average. The maximum lifespan in the wild is 11.3 years. [5] The eggs of the song sparrow are brown with greenish-white spots. Females lay three to five eggs per clutch, with an average incubation time of 13–15 days before hatching.
In the field, they are most easily confused with the Lincoln's sparrow and the Savannah sparrow. The former can be recognized by its shorter, grayer tail and the differently-patterned head, the brown cheeks forming a clear-cut angular patch. The Savannah sparrow has a forked tail and yellowish flecks on the face when seen up close.
Though a habitat generalist, the song sparrow favors brushland and marshes, including salt marshes across most of Canada and the United States. They also thrive in human dominated areas such as in suburbs, agricultural fields, and along roadsides. Permanent residents of the southern half of their range, northern populations of the song sparrow migrate to the southern United States or Mexico during winter and intermingle with the native, non-migratory population. The song sparrow is a very rare vagrant to western Europe, with a few recorded in Great Britain and Norway.
These birds forage on the ground, in shrubs or in very shallow water. They eat mainly insects and seeds. Birds in salt marshes may also eat small crustaceans. They nest either in a sheltered location on the ground or in trees or shrubs. Song sparrows with areas of shrub cover in their territory, away from the intertidal coastline, have greater over-winter survival, as well as higher reproductive success. [6]
The song sparrow has been the subject of several studies detailing the physiological reactions of bird species to conditions such as daylight length and differing climatic conditions. Most birds gain mass in their reproductive organs in response to some signal, either internal or external as the breeding season approaches. The exact source of this signal varies from species to species – for some, it is an endogenous process separate from environmental cues, while other species require extensive external signals of changing daylight length and temperature before beginning to increase the mass of their reproductive organs. Male specimens of M. melodia gain significant testicular mass in response both to changes in the daily photoperiod and as a result of endogenous chemical signals. [7] Females also undergo significant ovarian growth in response to both photo-period and endogenous signals. Hormone levels in both males and females fluctuate throughout the breeding season, having very high levels in March and late April and then declining until May. [8] These studies suggest that there are multiple factors at work that influence when and how the song sparrow breeds other than just increasing day length.
Due to the myriad subspecies of the song sparrow and the extremely varied climate of southern California, where many of these subspecies make their homes, physiological studies were undertaken to determine how climatic conditions and local environment influenced the bill size of M. melodia subspecies. The bill of a bird is highly important for thermoregulation as the bare surface area makes a perfect place to radiate excess heat or absorb solar energy to maintain homeostasis. [9] Knowing this, comparisons of bill length between individual song sparrows collected in different habitats were made with regard to the primary habitat type or microclimate that they were collected in. Larger beaked subspecies were strongly correlated with hotter microclimates - a correlation that follows from the conditions of Allen's Rule. [10]
The sparrow species derives its name from its colorful repertoire of songs. Enthusiasts report that one of the songs heard often in suburban locations closely resembles the opening four notes of Ludwig van Beethoven's Symphony No. 5. The male uses a fairly complex song to declare ownership of its territory and attract females.
Singing itself consists of a combination of repeated notes, quickly passing isolated notes, and trills. The songs are very crisp, clear, and precise, making them easily distinguishable by human ears. A particular song is determined by not only pitch and rhythm but also the timbre of the trills. Although one bird will know many songs—as many as 20 different tunes with as many as 1000 improvised variations on the basic theme, [11] —unlike thrushes, the song sparrow usually repeats the same song many times before switching to a different song.
Song sparrows typically learn their songs from a handful of other birds that have neighboring territories. They are most likely to learn songs that are shared between these neighbors. Ultimately, they will choose a territory close to or replacing the birds that they have learned from. This allows the song sparrows to address their neighbors with songs shared with those neighbors. It has been demonstrated that song sparrows are able to distinguish neighbors from strangers on the basis of song, and also that females are able to distinguish (and prefer) their mate's songs from those of other neighboring birds, and they prefer songs of neighboring birds to those of strangers. [12]
A 2022 study by Duke University also found that male song sparrows memorize a 30-minute long playlist of their songs and use that information to curate both their current playlist and the following one. The findings suggest that male song sparrows deliberately shuffle and repeat their songs possibly to keep a female's attention. [13]
Common predators of the song sparrow include cats, hawks, and owls; however snakes, dogs, and the American kestrel are treated ambiguously, suggesting that they are less of a threat[ citation needed ]. The song sparrow recognizes enemies by both instinctual and learned patterns (including cultural learning), and adjusts its future behavior based on both its own experiences in encounters, and from watching other birds interact with the enemies. Comparisons of experiments on hand-raised birds to observation of birds in the wild suggest that the fear of owls and hawks is instinctual, but fear of cats is learned. [14]
Song sparrows' nests are parasitized by the brown-headed cowbird. The cowbirds' eggs closely resemble song sparrows' eggs, although the cowbirds' eggs are slightly larger. Song sparrows recognize cowbirds as a threat and attack the cowbirds when they are near the nest. There is some evidence that this behavior is learned rather than instinctual. [14] A more recent study found that the behavior of attacking female cowbirds near nests may actually attract cowbird parasitism because the female cowbirds use such behavior to identify female song sparrows that are more likely to successfully raise a cowbird chick. [15] One study found that while cowbird parasitism did result in more nest failure, overall there were negligible effects on song sparrow populations when cowbirds were introduced to an island. The study pointed to a number of explanatory factors including song sparrows raising multiple broods, and song sparrows' abilities to raise cowbird chicks with their own. [16]
The song sparrow is one of the most polytypic bird species in North America, rivaling other variable species on a global scale, such as the horned lark, the yellow wagtail, the golden whistler, and the island thrush. There have been a total of 52 subspecies named altogether, of which 25 are still recognized following a systematic morphological review. [17] A high quality reference genome has recently been created for this species, which will allow for comprehensive genomic analyses of the differences between the several morphologically diverged subspecies. [18]
Small, brownish, long-winged forms with strong black streaks.
Large, dark, diffuse dark streaks. A study of mtDNA allozyme variation of most forms in this group concluded that they are of comparatively recent origin and that island populations are apparently derived independently from each other. [19]
Small, well-marked and short-winged brownish forms. All resident, except occasional birds from upland populations.
Small, pale, streaks rufous; all resident.
Black-spotted, white throats; all resident.
Seen as a whole, the song sparrow is widespread and common enough to be classified as Species of Least Concern by the IUCN. The taxa mailliardi, maxillaris, samuelis (all Category 3), pusillula (Category 2), and graminea (Category 1) are listed as Species of Special Concern in California. [22]
The horned lark or shore lark is a species of lark in the family Alaudidae found across the northern hemisphere. It is known as "horned lark" in North America and "shore lark" in Europe.
The dark-eyed junco is a species of junco, a group of small, grayish New World sparrows. The species is common across much of temperate North America and in summer it ranges far into the Arctic. It is a variable species, much like the related fox sparrow, and its systematics are still not completely resolved.
The chipping sparrow is a species of New World sparrow, a passerine bird in the family Passerellidae. It is widespread, fairly tame, and common across most of its North American range.
The brown-headed cowbird is a small, obligate brood parasitic icterid native to temperate and subtropical North America. It is a permanent resident in the southern parts of its range; northern birds migrate to the southern United States and Mexico in winter, returning to their summer habitat around March or April.
The Savannah sparrow is a small New World sparrow that is the only member of the genus Passerculus. It is a widespread and abundant species that occupies open grassland habitats in North America.
The swamp sparrow is a medium-sized New World sparrow related to the song sparrow.
Lincoln's sparrow is a small sparrow native to North America. It is a less common passerine bird that often stays hidden under thick ground cover, but can be distinguished by its sweet, wrenlike song. Lincoln's sparrow is one of three species in the genus Melospiza which also includes the song sparrow and the swamp sparrow. It lives in well-covered brushy habitats, often near water. This bird is poorly documented because of its secretive nature and breeding habits solely in boreal regions.
The grasshopper sparrow is a small New World sparrow. It belongs to the genus Ammodramus, which contains three species that inhabit grasslands and prairies. Grasshopper sparrows are sometimes found in crop fields and they will readily colonize reclaimed grassland. In the core of their range, grasshopper sparrows are dependent upon large areas of grassland where they avoid trees and shrubs. They seek out heterogenous patches of prairie that contain clumps of dead grass or other vegetation where they conceal their nest, and also contain barer ground where they forage for insects, spiders, and seeds. Grasshopper sparrows are unusual among New World sparrows in that they sing two distinct song types, the prevalence of which varies with the nesting cycle. The primary male song, a high trill preceded by a stereotyped series of short chips, is reminiscent of the sounds of grasshoppers and is the origin of this species' name. Like some other birds of the central North American grasslands, this species also moves around a lot, not only via annual migrations, but individuals frequently disperse between breeding attempts or breeding seasons. Grasshopper sparrows are in steep decline across their range, even in the core of the breeding distribution in the tallgrass prairies of the central Great Plains. The Florida grasshopper sparrow is highly endangered.
The California quail, also known as the California valley quail or Valley quail, is a small ground-dwelling bird in the New World quail family. These birds have a curving crest or plume, made of six feathers, that droops forward: black in males and brown in females; the flanks are brown with white streaks. Males have a dark brown cap and a black face with a brown back, a grey-blue chest and a light brown belly. Females and immature birds are mainly grey-brown with a light-colored belly. Their closest relative is Gambel's quail, which has a more southerly distribution and a longer crest at 2.5 in (6.4 cm), a brighter head and lacks the scaly appearance of the California quail. The two species separated about 1–2 million years ago, during the Late Pliocene or Early Pleistocene. It was selected as the state bird of California in 1931.
The shiny cowbird is a passerine bird in the New World family Icteridae. It breeds in most of South America except for dense forests and areas of high altitude such as mountains. Since 1900 the shiny cowbird's range has shifted northward, and it was recorded in the Caribbean islands as well as the United States, where it is found breeding in southern Florida. It is a bird associated with open habitats, including disturbed land from agriculture and deforestation.
The Bewick's wren is a wren native to North America. It is the only species placed in the genus Thryomanes. At about 14 cm (5.5 in) long, it is grey-brown above, white below, with a long white eyebrow. While similar in appearance to the Carolina wren, it has a long tail that is tipped in white. The song is loud and melodious, much like the song of other wrens. It lives in thickets, brush piles and hedgerows, open woodlands and scrubby areas, often near streams. It eats insects and spiders, which it gleans from vegetation or finds on the ground.
Amak Island is an uninhabited island in Aleutians East Borough, Alaska, United States. The island lies north of the western tip of the Alaska Peninsula, and northwest of the mainland city of Cold Bay. The island's land area is 5.828 square miles (15.09 km2) and its maximum elevation is 1,601 feet (488 m). The island's volcano, Mount Amak, last erupted in 1796.
The black-chinned sparrow is a small bird in the genus Spizella, in the New World sparrow family Passerellidae. It is found in the southwestern United States and throughout much of Mexico north of the Isthmus of Tehuantepec; most populations in the US migrate south after breeding while those in Mexico are resident. It is a slim, long-tailed bird, primarily gray with a reddish-brown back streaked with black, brown wings and tail, a pink beak, and brownish legs and feet. In the breeding season, the male shows black on his throat, chin, and the front of his face. Females, youngsters and nonbreeding males show little or no black in these areas. An unobtrusive bird, it spends much of its time foraging slowly along the ground, either alone or in small groups, sometimes mixing with other Spizella species. It is an omnivore, feeding primarily on seeds during the winter and insects during the summer. It builds a cup-shaped nest of grasses, rootlets, or plant fibers, into which the female lays 2–5 pale blue eggs. The female does most or all of the egg incubation, but both parents feed the hatched nestlings.
The rufous-crowned sparrow is a small American sparrow. This passerine is primarily found across the Southwestern United States and much of the interior of Mexico, south to the transverse mountain range, and to the Pacific coast to the southwest of the transverse range. Its distribution is patchy, with populations often being isolated from each other. Twelve subspecies are generally recognized, though up to eighteen have been suggested. This bird has a brown back with darker streaks and gray underparts. The crown is rufous, and the face and supercilium are gray with a brown or rufous streak extending from each eye and a thick black malar streak.
Cassin's sparrow is a medium-sized sparrow.
The chestnut-backed chickadee is a small passerine bird in the tit family, Paridae, native to western North America.
The sooty fox sparrow contains the darkest-colored taxa in the genus Passerella. It is currently classified as a "subspecies group" within the fox sparrow pending wider-spread acceptance of species status. It has long been suspected to be a separate evolutionary lineage due to morphological distinctness, and this is confirmed by analysis of mtDNA sequence and haplotype data. This group appears to be most closely related to the thick-billed and/or slate-colored fox sparrows.
The seaside sparrow is a species of American sparrow.
New World sparrows are a group of mainly New World passerine birds, forming the family Passerellidae. They are seed-eating birds with conical bills, brown or gray in color, and many species have distinctive head patterns.
The yellow-browed sparrow is a species of bird in the family Passerellidae. First described by Johann Baptist von Spix in 1825, this American sparrow is found across much of the Amazon basin in South America. Its natural habitats are subtropical or tropical moist shrubland, pastureland, and heavily degraded former forest.
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