Tabulata Temporal range: | |
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Tabulate coral (a syringoporid); Boone Limestone (Lower Carboniferous) near Hiwasse, Arkansas. Scale bar is 2.0 cm. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Cnidaria |
Subphylum: | Anthozoa |
Subclass: | † Tabulata |
Suborders | |
Tabulata, commonly known as tabulate corals, are an order of extinct forms of coral. They are almost always colonial, forming colonies of individual hexagonal cells known as corallites defined by a skeleton of calcite, similar in appearance to a honeycomb. Adjacent cells are joined by small pores. Their distinguishing feature is their well-developed horizontal internal partitions (tabulae) within each cell, but reduced or absent vertical internal partitions (septa). They are usually smaller than rugose corals, but vary considerably in shape, from flat to conical to spherical.
Around 300 species have been described. Among the most common tabulate corals in the fossil record are Aulopora , Favosites , Halysites , Heliolites , Pleurodictyum , Sarcinula and Syringopora . Tabulate corals with massive skeletons often contain endobiotic symbionts, such as cornulitids and Chaetosalpinx . [1] [2]
Like rugose corals, they lived entirely during the Paleozoic, being found from the Ordovician to the Permian. With Stromatoporoidea and rugose corals, the tabulate corals are characteristic of the shallow waters of the Silurian and Devonian. Sea levels rose in the Devonian, and tabulate corals became much less common. They finally became extinct in the Permian–Triassic extinction event.
The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Ma) to the start of the Silurian Period 443.8 Mya.
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
The rugosa, also called the tetracorallia or horn coral, are an extinct order of solitary and colonial corals that were abundant in Middle Ordovician to Late Permian seas.
Tentaculites is an extinct genus of conical fossils of uncertain affinity, class Tentaculita, although it is not the only member of the class. It is known from Lower Ordovician to Upper Devonian deposits both as calcitic shells with a brachiopod-like microstructure and carbonaceous 'linings'. The "tentaculites" are also referred to as the styliolinids.
Stromatoporoidea is an extinct clade of sea sponges common in the fossil record from the Middle Ordovician to the Late Devonian. They can be characterized by their densely layered calcite skeletons lacking spicules. Stromatoporoids were among the most abundant and important reef-builders of their time, living close together in flat biostromes or elevated bioherms on soft tropical carbonate platforms.
Cornulitida is an extinct order of encrusting animals from class Tentaculita, which were common around the globe in the Ordovician to Devonian oceans, and survived until the Carboniferous. Organisms that may be the oldest cornulitids have been found in Cambrian sediments of Jordan.
Favosites is an extinct genus of tabulate coral characterized by polygonal closely packed corallites. The walls between corallites are pierced by pores known as mural pores which allowed transfer of nutrients between polyps. Favosites, like many corals, thrived in warm sunlit seas, feeding by filtering microscopic plankton with their stinging tentacles and often forming part of reef complexes. The genus had a worldwide distribution from the Late Ordovician to Late Permian.
Halysites is an extinct genus of tabulate coral. Colonies range from less than one to tens of centimeters in diameter, and they fed upon plankton.
Anticalyptraea is a fossil genus of encrusting tentaculitoid tubeworms from the Silurian to Devonian of Europe and North America . Anticalyptraea commonly encrust various invertebrate fossils such as stromatoporoids, rugose corals, bryozoans, brachiopods and crinoids, but they can also be common on the hardgrounds.
Chaetosalpinx is an ichnogenus of bioclaustrations. Chaetosalpinx includes straight to sinuous cavities that are parallel to the host's axis of growth. The cavity is circular to oval in cross-section and it lacks a wall lining or floor-like tabulae. They are common in tabulate and rugose corals from Late Ordovician to Devonian of Europe and North America. They may have been parasites.
Fossils of many types of water-dwelling animals from the Devonian period are found in deposits in the U.S. state of Michigan. Among the more commonly occurring specimens are bryozoans, corals, crinoids, and brachiopods. Also found, but not so commonly, are armored fish called placoderms, snails, sharks, stromatolites, trilobites and blastoids.
Syringopora is an extinct genus of phaceloid tabulate coral. It has been found in rocks ranging in age from the Ordovician to the Permian, although it was most widespread during the Silurian, Devonian, and Carboniferous periods. Among other places, it has been found in the Columbus Limestone in Ohio, and in the Spring Branch Member of the Lecompton Limestone in Kansas.
Paleontology in Michigan refers to paleontological research occurring within or conducted by people from the U.S. state of Michigan. During the Precambrian, the Upper Peninsula was home to filamentous algae. The remains it left behind are among the oldest known fossils in the world. During the early part of the Paleozoic Michigan was covered by a shallow tropical sea which was home to a rich invertebrate fauna including brachiopods, corals, crinoids, and trilobites. Primitive armored fishes and sharks were also present. Swamps covered the state during the Carboniferous. There are little to no sedimentary deposits in the state for an interval spanning from the Permian to the end of the Neogene. Deposition resumed as glaciers transformed the state's landscape during the Pleistocene. Michigan was home to large mammals like mammoths and mastodons at that time. The Holocene American mastodon, Mammut americanum, is the Michigan state fossil. The Petoskey stone, which is made of fossil coral, is the state stone of Michigan.
Paleontology in Ohio refers to paleontological research occurring within or conducted by people from the U.S. state of Ohio. Ohio is well known for having a great quantity and diversity of fossils preserved in its rocks. The state's fossil record begins early in the Paleozoic era, during the Cambrian period. Ohio was generally covered by seawater from that time on through the rest of the early Paleozoic. Local invertebrates included brachiopods, cephalopods, coral, graptolites, and trilobites. Vertebrates included bony fishes and sharks. The first land plants in the state grew during the Devonian. During the Carboniferous, Ohio became a more terrestrial environment with an increased diversity of plants that formed expansive swampy deltas. Amphibians and reptiles began to inhabit the state at this time, and remained present into the ensuing Permian. A gap in the local rock record spans from this point until the start of the Pleistocene. During the Ice Age, Ohio was home to giant beavers, humans, mammoths, and mastodons. Paleo-Indians collected fossils that were later incorporated into their mounds. Ohio has been the birthplace of many world famous paleontologists, like Charles Schuchert. Many significant fossils curated by museums in Europe and the United States were found in Ohio. Major local fossil discoveries include the 1965 discovery of more than 50,000 Devonian fish fossils in Cuyahoga County. The Ordovician trilobite Isotelus maximus is the Ohio state invertebrate fossil.
Paleontology in Kentucky refers to paleontological research occurring within or conducted by people from the U.S. state of Kentucky.
Paleontology in Vermont comprises paleontological research occurring within or conducted by people from the U.S. state of Vermont. Fossils are generally uncommon in Vermont. Nevertheless, however, significant finds have been made in the state. Very few fossils are known in Vermont east of the Green Mountains due to the type of rock underlying that area. During the early part of the Paleozoic era, Vermont was covered by a warm, shallow sea that would end up being home to creatures like brachiopods, corals, crinoids, ostracoderms, and trilobites. There are no rocks in the state from the Carboniferous, Permian, Triassic, or Jurassic periods. The few Cretaceous rocks present contain no fossils. The Paleogene and Neogene periods are also absent from the local rock record. During the Ice Age, glaciers scoured the state. At times the state was inundated by seawater, allowing marine mammals to venture in. After the seawater drained away the state was home to mastodons. Local fossils had already attracted scientific attention by the mid-19th century when mastodon remains were found in Rutland County. In 1950 a major Paleozoic invertebrate find occurred. The Pleistocene Beluga whale Delphinapterus leucas is the Vermont state fossil.
Paleontology in Wisconsin refers to paleontological research occurring within or conducted by people from the U.S. state of Wisconsin. The state has fossils from the Precambrian, much of the Paleozoic, some a parts of the Mesozoic and the later part of the Cenozoic. Most of the Paleozoic rocks are marine in origin. Because of the thick blanket of Pleistocene glacial sediment that covers the rock strata in most of the state, Wisconsin’s fossil record is relatively sparse. In spite of this, certain Wisconsin paleontological occurrences provide exceptional insights concerning the history and diversity of life on Earth.
Paleontology in Texas refers to paleontological research occurring within or conducted by people from the U.S. state of Texas. Author Marian Murray has said that "Texas is as big for fossils as it is for everything else." Some of the most important fossil finds in United States history have come from Texas. Fossils can be found throughout most of the state. The fossil record of Texas spans almost the entire geologic column from Precambrian to Pleistocene. Shark teeth are probably the state's most common fossil. During the early Paleozoic era Texas was covered by a sea that would later be home to creatures like brachiopods, cephalopods, graptolites, and trilobites. Little is known about the state's Devonian and early Carboniferous life. Evidence indicates that during the late Carboniferous the state was home to marine life, land plants and early reptiles. During the Permian, the seas largely shrank away, but nevertheless coral reefs formed in the state. The rest of Texas was a coastal plain inhabited by early relatives of mammals like Dimetrodon and Edaphosaurus. During the Triassic, a great river system formed in the state that was inhabited by crocodile-like phytosaurs. Little is known about Jurassic Texas, but there are fossil aquatic invertebrates of this age like ammonites in the state. During the Early Cretaceous local large sauropods and theropods left a great abundance of footprints. Later in the Cretaceous, the state was covered by the Western Interior Seaway and home to creatures like mosasaurs, plesiosaurs, and few icthyosaurs. Early Cenozoic Texas still contained areas covered in seawater where invertebrates and sharks lived. On land the state would come to be home to creatures like glyptodonts, mammoths, mastodons, saber-toothed cats, giant ground sloths, titanotheres, uintatheres, and dire wolves. Archaeological evidence suggests that local Native Americans knew about local fossils. Formally trained scientists were already investigating the state's fossils by the late 1800s. In 1938, a major dinosaur footprint find occurred near Glen Rose. Pleurocoelus was the Texas state dinosaur from 1997 to 2009, when it was replaced by Paluxysaurus jonesi after the Texan fossils once referred to the former species were reclassified to a new genus.
The geological history of North America comprises the history of geological occurrences and emergence of life in North America during the interval of time spanning from the formation of the earth through to the emergence of humanity and the start of prehistory. At the start of the Paleozoic Era, what is now "North" America was actually in the Southern Hemisphere. Marine life flourished in the country's many seas, although terrestrial life had not yet evolved. During the latter part of the Paleozoic, seas were largely replaced by swamps home to amphibians and early reptiles. When the continents had assembled into Pangaea, drier conditions prevailed. The evolutionary precursors to mammals dominated the country until a mass extinction event ended their reign.
Olev Vinn is Estonian paleobiologist and paleontologist.