European spadefoot toads Temporal range: | |
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Pelobates fuscus fuscus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Amphibia |
Order: | Anura |
Suborder: | Mesobatrachia |
Superfamily: | Pelobatoidea |
Family: | Pelobatidae Bonaparte, 1850 |
Genera | |
See text | |
The distribution of extant pelobatids (in black). |
The European spadefoot toads are a family of frogs, the Pelobatidae, with only one extant genus Pelobates, containing six species. They are native to Europe, the Mediterranean, northwestern Africa, and western Asia.
The European spadefoot toad grows up to 10 cm (3.9 in) in length and is often inconspicuously coloured. They have squat bodies with smooth skin and eyes with vertical pupils. They are predominantly fossorial (burrowing) frogs, which dig into sandy soils. Pelobatidae frogs burrow backwards and they spend much of their time in the ground. They prefer open areas with loose soil as opposed to dense compact soil to facilitate the burrowing and have hardened protrusions on their feet to aid in digging, which is the source of the common name. They emerge from the ground during periods of rain and breed in pools, which are usually temporary. [2]
All of the species from this family have free-living, aquatic tadpoles. The eggs are laid in temporary ponds that may quickly evaporate, so the tadpole stage is unusually brief, with rapid development to the adult form in as little as two weeks. To further speed their growth, some of the tadpoles are cannibalistic, eating their brood-mates to increase their supply of protein. [2]
The seven species of American spadefoot toads (genera Scaphiopus and Spea ) were previously also included in the family Pelobatidae, but are now generally regarded as the separate family Scaphiopodidae.
Family Pelobatidae
The earliest fossil genus of pelobatids, Elkobatrachus , was described in 2006. [4]
In the Jurassic Morrison Formation, pelobatids are represented by the ilium of an unnamed but indeterminate species. [1] This ilium is larger than that of Enneabatrachus, a contemporary discoglossid species. [1] A specimen has been recovered from Quarry 9 of Como Bluff in Wyoming. [1] Pelobatids are present in stratigraphic zones 5 and 6 of the formation. [5]
The Oligocene site of Enspel in Germany preserves evidence of pelobatid tadpoles feeding on pollen. [6]
A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura. The oldest fossil "proto-frog" Triadobatrachus is known from the Early Triassic of Madagascar, but molecular clock dating suggests their split from other amphibians may extend further back to the Permian, 265 million years ago. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest. Frogs account for around 88% of extant amphibian species. They are also one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from taxonomy or evolutionary history.
Bombinatoridae is a family of toads found in Eurasia. Species of the family have flattened bodies and some are highly toxic.
The Alytidae are a family of primitive frogs. Their common name is painted frogs or midwife toads. Most are endemic to Europe, but three species occur in northwest Africa, and a species formerly thought to be extinct is found in Israel.
The Mesobatrachia is a paraphyletic group of relatively primitive frogs. At the end of 2016, it contained 3 superfamilies, 6 families, 16 genera, and 244 species. Recognized as a group in 1993, the name is contrasted with the primitive Archaeobatrachia and the more diverse and advanced Neobatrachia.
Caeciliidae is the family of common caecilians. They are found in Central and South America. Like other caecilians, they superficially resemble worms or snakes.
The Mexican burrowing toad is the single living representative of the family Rhinophrynidae. It is a unique species in its taxonomy and morphology, with special adaptations to assist them in digging burrows where they spend most of their time. These adaptations include a small pointed snout and face, keratinized structures and a lack of webbing on front limbs, and specialized tongue morphology to assist in feeding on ants and termites underground. The body is nearly equal in width and length. It is a dark brown to black color with a red-orange stripe on its back along with splotches of color on its body. The generic name Rhinophrynus means 'nose-toad', from rhino- (ῥῑνο-), the combining form of the Ancient Greek rhis and phrunē.
The Rhinophrynidae are a family of frogs containing one extant genus, the monotypic Rhinophrynus, and a number of fossil genera. The family is sometimes known as the Mexican burrowing toads or simply burrowing toads.
The Scaphiopodidae are a family of American spadefoot toads, which are native to North America. The family is small, comprising only eleven different species.
A true toad is any member of the family Bufonidae, in the order Anura. This is the only family of anurans in which all members are known as toads, although some may be called frogs. The bufonids now comprise more than 35 genera, Bufo being the best known.
Scaphiopus is a genus of North American amphibian commonly referred to as the North American spadefoots, southern spadefoots, or eastern spadefoot toads. They differ greatly from true toads by having eyes with vertical pupils, no parotoid gland, and relatively smooth skin. Their most distinctive feature is a spade-like projection on their hind feet, from which their common name is derived. This projection enables spadefoot toads to dig in loose soils with ease. Its scientific name means ‘spade-foot’ as well, from the Ancient Greek skaphís and pous.
The plains spadefoot toad is a species of American spadefoot toad which ranges from southwestern Canada, throughout the Great Plains of the western United States, and into northern Mexico. Like other species of spadefoot toads, they get their name from a spade-like projection on their hind legs which allows them to dig into sandy soils. Their name, in part, comes from their keratinized metatarsals, which are wide instead of "sickle shaped". The species name translates as buzzing leaf shaped. This refers to the species' distinguishing features; its buzzing mating call, and its leaf-shaped digging metatarsals. It was first described by Cope in 1863.
Pelobates varaldii, the Moroccan spadefoot toad, Moroccan spadefoot, or Varaldi's spadefoot toad, is a species of frog in the family Pelobatidae. As currently known, it is endemic to the coastal north-western Morocco, although there is an unconfirmed record from the Spanish territory of Melilla that could possibly represent this species. The specific name varaldii honours Marcel Varaldi who collected amphibians and reptiles in Morocco.
The Pelobatoidea are a superfamily of frogs. They typically combine a toad-like body shape with a frog-like, pointed face. Phylogenetically they stand between primitive frogs on the one side and higher frogs on the other and are therefore – among other things by characteristics of bone construction – in the suborder Mesobatrachia.
Eopelobates is an extinct genus of frogs in the family Pelobatidae. Closely related to the living European spadefoot toad, it is known from the Eocene of western North America, and the Eocene–Pliocene of Europe. It is suggested that the distribution over both Europe and North America is due to dispersal during the Paleocene-Eocene Thermal Maximum.
Pelobates cultripes is a toad species in the family Pelobatidae. It is known under many different common names, including the western spadefoot, Iberian spadefoot toad, Spanish spadefoot toad, and Wagler's spadefoot toad. It is found in most of the Iberian Peninsula with isolated populations southern and western France.
Pelobates syriacus, the eastern spadefoot or Syrian spadefoot, is a species of toad in the family Pelobatidae, native to an area extending from Eastern Europe to Western Asia.
Robert Frederick Inger was an American herpetologist. During his lifetime, he wrote numerous books and publications about herpetology. He was also the curator for amphibians and reptiles at the Field Museum in Chicago, Illinois.