Green jack

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Green jack
Green jack.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Carangiformes
Family: Carangidae
Genus: Caranx
Species:
C. caballus
Binomial name
Caranx caballus
Günther, 1868
Caranx caballus distribution map.png
Approximate range of the green jack
Synonyms [1]
  • Trachurus boops
    Girard, 1858
  • Caranx girardi
    Steindachner, 1870
  • Carangoides caballus
    (Günther, 1868)

The green jack (Caranx caballus), also known as the horse jack, is an abundant species of coastal marine fish in the jack family Carangidae. The species is distributed in the eastern Pacific Ocean along the American coastline from Santa Cruz Island off California in the north to Peru in the south, as well as a number of islands including the Galapagos and recently, Hawaii. The green jack is distinguished from other similar carangid species by a number of features including gill raker and lateral line scale counts, and the presence of an adipose eyelid. It is a moderately large species, growing to at least 55 cm in length and 2.81 kg, although unconfirmed reports suggest a much larger maximum size. It lives in a wide range of continental shelf environments including estuaries, bays, reefs and offshore seamounts, living both pelagically and demersally. The green jack is a predatory species, preying on a variety of fish, crustaceans and cephalopods, as well as zooplankton. Most fish are sexually mature by the time they attain 38 cm, with spawning occurring between May and October. The species is of high importance to fisheries throughout its range, caught by pelagic trawls, a variety of netting methods and hook and line. The green jack is also of interest to anglers, taken by bait and lures, although is considered to only be fair in eating quality.

Contents

Taxonomy and naming

The green jack is formally classified within the genus Caranx , one of a number of groups of fish referred to as jacks and trevallies. Caranx is further classified in the family Carangidae, part of the order Carangiformes. [2]

The species was first scientifically described by the French biologist Charles Frédéric Girard in 1858, who named the species Trachurus boops, placing the species in the horse mackerel genus based on the holotype taken off of San Diego, California. [3] With the instability of carangid taxonomy at the time, the species was reassigned to a number of genera, including Caranx, which created a taxonomic homonym with the Caranx boops described by Georges Cuvier in 1833. In 1868, British zoologist Albert Günther unaware of the prior naming independently renamed the species Caranx caballus based on specimens collected from Panama, one of which was designated to be the holotype. [4] The species was further renamed in 1870 by Franz Steindachner, who attempted to remove the Caranx boops homonym by simply renaming the species Caranx girardi. Later reviews of the Carangidae revealed this synonymy, and due to Gunther's' earlier naming, his species name was kept and attributed to him, while Cuvier's' Caranx boops was later moved to Selar boops . [5] The specific name caballus is Latin for 'horse', reflecting the common name of horse mackerel given to this and many similar species at the time. [6] William Smith-Vaniz has suggested the possibility C. caballus is conspecific with a very similar Atlantic species; Caranx crysos , but no studies into this relationship have been undertaken. [7] Its position in the genus Caranx has also been questioned recently by John Randall due to its unusual dentition. The common name 'green jack' is in reference to the species' common coloration, while 'horse jack' is derived from the Latin name. In Spanish-speaking nations, it is often known under broadly applied names including cocinero and caballa. [8]

Description

The green jack is moderately large in size, growing to a maximum confirmed length of 55 cm and 2.81 kg in weight. [8] Unconfirmed reports indicate the species may grow much larger; up to 1 m in size, but it is most commonly seen at lengths below 40 cm. [7] The green jack is morphologically similar to a number of other carangids, having an elongated, moderately compressed fusiform body with dorsal and ventral profiles of approximately equal convexity and a slightly pointed snout. The posterior section of the eye is covered by a moderately well developed adipose eyelid, and the posterior extremity of the jaw is vertically under the center of the eye. [7] The dorsal fin is in two parts, the first consisting of 8 spines and the second of 1 spine followed by 22 to 25 soft rays. The anal fin consists of 2 anteriorly detached spines followed by 1 spine and 16 to 21 soft rays. The lobes of both the second dorsal and anal fin are slightly elongated and almost entirely covered in small scales, but are still much shorter than the head length. The lateral line has a pronounced but short anterior arch, with the curved section intersecting the straight section below the spine of the second dorsal fin. The straight section contains 0 to 7 scales followed by 42 to 56 very strong scutes, and 43 to 47 scales overall. [6] The chest is completely scaled. [9] The upper jaw contains an irregular series of outer canines with an inner band of small, regularly spaced teeth, while the lower jaw contains a single band of small teeth. The species has 40 to 45 gill rakers in total; 10 to 15 on the upper limb and 27 to 30 on the lower limb, with this the only feature that differs between C. caballus and C. crysos. There are 25 vertebrae present. [6]

In life, the green jack has a greenish blue colour overall, with an olive green to dark blue back and a golden to grey coloured belly, with a distinct back blotch on the outer edge of the operculum. Individuals in schools often have a very evident pearly white marking near this black blotch. Juveniles have 7 dark vertical stripes on their flanks which fade with age. [10] The caudal fin is grey with dark tips, with all other fins being light grey to hyaline in colour. [7]

Distribution and habitat

The green jack is distributed throughout the coastal waters of the tropical and subtropical waters of the eastern Pacific Ocean. Its range extends along the coast of the Americas from Santa Cruz Island, California, south through Mexico and Central America, and down to Peru. The species also inhabits a number of offshore islands including the Galápagos Islands. [10] The species had been captured on occasion from the Hawaiian Islands since 1923, but was misidentified until William Smith-Vaniz re-examined the specimens in 1981 and identified them as Caranx caballus. [6] Due to the heavy fishing pressures throughout Hawaii and the rarity of captures, it was concluded these few individuals were strays carried by ocean currents from the eastern Pacific, as these were the only known specimens. In 1998 however, the species appeared in fishermen's catches in large numbers, and has ever since, indicating a population has been established in the island chain. [6]

The green jack inhabits a variety of continental shelf environments to a depth of at least 100 m, predominantly inshore reef systems, as well as shallow bays, estuaries and lagoons. [7] It leads both a demersal and pelagic lifestyle, forming large schools which can move large distances offshore, allowing them to reach offshore island as well as deep, isolated seamounts. [11]

Biology

Illustration of the green jack Green jack.png
Illustration of the green jack

The green jack is a gregarious species, forming moderately large schools of fish in the marine environment. Smaller shoals are formed by juveniles when entering shallower waters including bays and estuaries. The species is predatory in nature, taking small fish, cephalopods, crabs, shrimps and other crustaceans, either in midwater or on the sea floor. [12] Green jack are also known to consume zooplankton, especially in deeper waters around islands and seamounts where the plankton is aggregated. [11] Throughout its range, the species is one of the most abundant species, making it an important part of the ecology, not only as a predator but as prey. It is known to be taken by larger fish such as marlin and sharks, [13] as well a variety of other animals.[ citation needed ]

Reproduction in the species has been quite well studied, with the commercial value of the species having led to a number of growth and population dynamics studies being undertaken. Spawning appears to take place from May to October, with peaks in June and October.[ citation needed ] Larval growth has been extensive described, with the larvae identifiable by their pigmentation pattern and meristic count of 25 vertebrae. The body shape of early larvae resembles that of Chloroscombrus orquetu , however, differences in pigmentation, particularly on the dorsal body margin, differentiate the species [14] Studies off the coast of Mexico have found the larvae tend to occur in deeper waters than most other carangid larvae inhabit, from 90 to 100 m depth, possibly indicating spawning occurs in deeper waters. [15] Otolith studies indicate that both the females and males grow at the same rate; 16.82 cm after the first year, 27.78 after the second, 34.66 after the third,[ citation needed ] with modelling suggesting the maximum length of 55 cm is reached at around 8 years of age.[ citation needed ] Most individuals reach sexual maturity before they reach 38 cm in length. [12] Juveniles tend to move to protected waters such as estuaries, moving to deeper waters at adulthood.[ citation needed ]

Relationship to humans

The green jack is one of the most abundant species of fish within its range, and as such is quite important to fisheries throughout its range. It is taken by pelagic trawls, inshore gill nets, purse nets, and seines, as well as hook and line gear. [7] Catch statistics are not kept for the fish throughout most of its range; however, the Mexican state of Colima is one exception. Records from 1980 onward show a catch between 9 and 250 tons, with an average of 93 tons, with the catch best between September and December. Like most carangid species in the area, it is considered a lower class product, fetching between US$ 0.50 and 0.80 per kilogram, making it accessible to the low-income population.[ citation needed ] The fish are sold both for consumption and as bait, either fresh, smoked or salted. Some hope the Hawaiian population becomes commercially viable in the future, as they are often landed in other carangid catches at the present. [6]

The green jack is also of modest interest to anglers in the region, with the species a good light-tackle fish. Lures are most often used on the species, including jigs, spoons, and squids, although baits of live fish and strip baits work equally as well. [16] The IGFA all tackle world record for the species stands at 2.81 kg (6 lb 3oz) caught off of Cabo San Lucas, Mexico in 2000. [17]

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The crevalle jack, also known as the common jack, black-tailed trevally, couvalli jack, black cavalli, jack crevale, or yellow cavalli is a common species of large marine fish classified within the jack family, Carangidae. The crevalle jack is distributed across the tropical and temperate waters of the Atlantic Ocean, ranging from Nova Scotia, Canada to Uruguay in the western Atlantic and Portugal to Angola in the eastern Atlantic, including the Mediterranean Sea. It is distinguishable from similar species by its deep body, fin colouration and a host of more detailed anatomical features, including fin ray and lateral line scale counts. It is one of the largest fish in the genus Caranx, growing to a maximum known length of 124 cm and a weight of 32 kg, although is rare at lengths greater than 60 cm. The crevalle jack inhabits both inshore and offshore waters to depths of around 350 m, predominantly over reefs, bays, lagoons and occasionally estuaries. Young fish dispersed north by currents in the eastern Atlantic are known to migrate back to more tropical waters before the onset of winter; however, if the fish fail to migrate, mass mortalities occur as the temperature falls below the species' tolerance.

<span class="mw-page-title-main">Pacific jack mackerel</span> Species of fish

The Pacific jack mackerel, also known as the Californian jack mackerel or simply jack mackerel, is an abundant species of pelagic marine fish in the jack family, Carangidae. It is distributed along the western coast of North America, ranging from Alaska in the north to the Gulf of California in the south, inhabiting both offshore and inshore environments. The Pacific jack mackerel is a moderately large fish, growing to a maximum recorded length of 81 cm, although commonly seen below 55 cm. It is very similar in appearance to other members of its genus, Trachurus, especially T. murphyi, which was once thought to be a subspecies of T. symmetricus, and inhabits waters further south. Pacific jack mackerel travel in large schools, ranging up to 600 miles offshore and to depths of 400 m, generally moving through the upper part of the water column.

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<span class="mw-page-title-main">Bigeye trevally</span> Species of fish

The bigeye trevally, also known as the bigeye jack, great trevally, six-banded trevally and dusky jack, is a species of widespread large marine fish classified in the jack family Carangidae. The bigeye trevally is distributed throughout the tropical waters of the Indian and Pacific Oceans, ranging from South Africa in the west to California and Ecuador in the east, including Australia to the south and Japan in the north. The bigeye trevally is best distinguished by its colouration, having a dark second dorsal fin with a white tip on the lobe, and also possessing a small dark spot on the operculum. Other more detailed anatomical features also set the species apart from other members of Caranx. The species is known to grow to a length of 120 cm and 18 kg.

<span class="mw-page-title-main">Giant trevally</span> Species of fish

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<span class="mw-page-title-main">Bar jack</span> Species of fish

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<span class="mw-page-title-main">Indian threadfish</span> Species of fish

The Indian threadfish, also known as the Indian threadfin, diamond trevally, mirror fish or plumed trevally, is a large species of coastal marine fish of the jack family, Carangidae. The species is widespread in the waters of the tropical Indo-West Pacific Ocean, ranging from east Africa to India, Asia, Indonesia and Australia. Adult fish tend to inhabit coastal waters over reefs down to 100 m in depth, while juveniles inhabit a variety of environments including estuaries and seagrass beds. The Indian threadfish is similar to the other two species in the genus Alectis, with a slight concavity in the profile of the head the most obvious distinguishing feature. It is a large species, growing to 165 cm and 25 kg in weight. The species is carnivorous, consuming fishes, cephalopods and crustaceans. The Indian threadfish is of minor commercial importance, and has been the subject of aquaculture in Singapore.

<span class="mw-page-title-main">Shrimp scad</span> Species of fish

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<i>Caranx lugubris</i> Species of fish

Caranx lugubris, the black jack, black trevally, black kingfish, coal fish or black ulua, is a species of large ocean fish in the jack family Carangidae. The species has a circumtropical distribution, found in oceanic, offshore waters of the tropical zones of the Pacific, Atlantic and Indian Oceans. The species is particularly prevalent around offshore islands such as the Caribbean islands in the Atlantic, Hawaii and French Polynesia in the Pacific and the Seychelles and Maldives in the Indian Ocean. Black jack are rare in shallow waters, preferring deep reefs, ledges and seamounts in clear waters. The species is easily distinguished by its black to grey fins and jet black scutes, with the head having a steep profile near the snout. The largest recorded length is 1 m and weight of 17.9 kg. The black jack lives either individually or in small schools, and is known to school with other species. It is a predatory fish, taking a variety of fish, crustaceans and molluscs as prey. Sexual maturity is reached at 34.6 cm in females and 38.2 cm in males, with spawning taking place between February and September in the Caribbean. The early life history of the species is very poorly understood. Black jack are of high importance to many island fisheries, but are rarely encountered in most continental fisheries. The species has a reputation as a gamefish, and is variably considered a terrible or excellent food fish, although several cases of ciguatera poisoning have been attributed to the species. The species was initially named Caranx ascensionis by Georges Cuvier, however several issues with the use of this name have seen Felipe Poey's name Caranx lugubris become the valid scientific name.

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<i>Caranx</i> Genus of fishes

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<span class="mw-page-title-main">Blue trevally</span> Species of fish

The blue trevally, also known as the banded trevally, barred trevally, Ferdau's trevally or Forskaal's jackfish, is a common, widespread species of pelagic marine fish classified in the jack family, Carangidae. The blue trevally is distributed throughout the tropical and subtropical waters of the Indo-Pacific and central Pacific regions, ranging from South Africa in the west to Hawaii in the east. It is a moderately large fish, growing to a recorded maximum length of 70 cm, with the number of rays in the second dorsal fin and the colouring serving as diagnostic features of the species. The species inhabits waters to depths of 60 m, generally inhabiting reefs, beaches, lagoons, and areas with sandy substrates. It is a predatory fish, taking other fish, prawns, crabs, and molluscs, and very little is known of the species' reproductive biology. The blue trevally is of varying importance to fisheries throughout its range, with some regions having high catches of the fish. It is considered to be a gamefish, and is sought after for its excellent eating qualities.

<span class="mw-page-title-main">Pacific crevalle jack</span> Species of fish

The Pacific crevalle jack is a species of large marine fish classified in the jack family Carangidae. The species is distributed through the tropical waters of the eastern Pacific Ocean from California in the north to Peru in the south, including several offshore islands. The species is best identified by its deep body and mostly unscaled breast, as well as other more detailed anatomical features. The species is known to grow to a maximum length of 101 cm and a weight of 19.7 kg. Disagreement on the status of the species has been significant in the scientific literature, with many claiming it to be conspecific with or subspecific to the Atlantic Caranx hippos. The most recent review of the crevalle jacks strongly concluded it to be a separate species based on the development of hyperostosis and fin colouring. It is predominantly a schooling inshore species, inhabiting sandy and rocky substrates, although larger individuals are occasionally found living pelagically to depths of 350 m. The Pacific crevalle jack is a fast-swimming predator, taking a variety of fish, crustaceans, and other small invertebrates. Spawning is thought to occur year-round, although peaks occur during November and May. The Pacific crevalle jack is an important species to commercial fisheries, with data available from Colima in Mexico indicating it accounts for up to 15% of the entire yearly catch. Pacific crevalle jack are highly rated gamefish, taken by both bait and a variety of lures, but are considered relatively poor quality food.

<span class="mw-page-title-main">Longfin crevalle jack</span> Species of fish

The longfin crevalle jack is a recently described species of large marine fish classified in the jack family, Carangidae. It inhabits the subtropical waters of the east Atlantic Ocean, ranging along the African coast from Mauritania south at least to Moçamedes in southern Angola, with the species historically present in the Mediterranean Sea. It is an inshore species, known to occasionally penetrate estuaries, possibly to spawn. The species is very similar to the crevalle jack, Caranx hippos, and is separated by its extended dorsal and anal fin lobes as well as more detailed anatomical features including dorsal and anal fin ray counts. The fish is known to reach 127 cm in length and 20.9 kg in weight. The longfin crevalle jack is a predatory fish, taking small fish as its main prey. Due to longstanding confusion between C. fischeri and C. hippos, the importance of each species to African fisheries is poorly understood, with a known combined catch in this region of between 2,233 and 10,054 tonnes per year in 1995–2004. Like its close relatives, the species is considered a powerful gamefish and highly sought after, although their coarse flesh makes for relatively poor quality food.

<span class="mw-page-title-main">Blacktip trevally</span> Species of fish

The blacktip trevally, also known as the blacktip kingfish or yellowtail kingfish, is a species of large marine fish classified in the jack family Carangidae. The blacktip trevally is distributed throughout the tropical to subtropical Indian and West Pacific Oceans, ranging from South Africa in the west to Fiji, Japan and northern Australia in the east. It inhabits coastal waters throughout its range, preferring moderately deep clear waters over rocky and coral reefs. The blacktip trevally is easily distinguished by its yellow fins and a dark upper caudal fin lobe which gives the species its common name, as well as a host of other anatomical features. The species is known to reach a maximum size of 1 m. It is a benthopelagic predator, commonly forming small shoals where it takes a variety of fishes, cephalopods and crustaceans as prey. Little is known of reproduction in the species, and spawning is assumed to take place in more tropical regions of its range, with juveniles known to inhabit bays and large estuaries. Blacktip trevally are often caught using hook and line and various nets in commercial fisheries although don't make up a large part of the market. They are also popular with anglers due to their fighting ability and decent table qualities.

<span class="mw-page-title-main">Brassy trevally</span> Species of fish

The brassy trevally, Caranx papuensis is a species of large marine fish classified in the jack family, Carangidae.

<span class="mw-page-title-main">False scad</span> Species of fish

The false scad, also known as the spotfin scad, ten-finned horse mackerel and yellow horse mackerel, is a species of medium-sized marine fish classified in the jack family Carangidae. The false scad is distributed throughout the tropical and temperate waters of the eastern Atlantic Ocean from Namibia in the south to Spain and throughout most of the Mediterranean in the north. The species has an atypical body form compared to other species of Caranx, and can be distinguished from these on its elongate 'scad-like' body shape alone. Distinguishing the species from members of Decapterus and Trachurus is more difficult, requiring detailed anatomical analysis. The false scad is known to grow to a length of 60 cm and a weight of 1 kg. The species lives both pelagically and demersally in continental shelf waters, ranging from depths of 15 to 200 m. It is a predatory fish, taking small fishes, crustaceans including euphausiids and shrimps, and cephalopods as its main prey, with significant dietary shifts as it ages. The false scad reaches sexual maturity at 2 years of age, with spawning occurring between April and July in shallow inshore waters, where the juveniles remain. The species is of major importance to fisheries throughout its range, but particularly from Morocco to Senegal, with annual catches ranging from 500 to 19000 t. The species is caught using trawls, seine nets and gill nets and sold fresh, frozen or salted.

<span class="mw-page-title-main">Senegal jack</span> Species of fish

The Senegal jack, also known as the African jack, is a species of large marine fish classified in the jack family Carangidae. The species is distributed through the tropical waters of the eastern Atlantic Ocean, ranging along the west African coast from Angola in the south to Mauritania in the north. It can be distinguished from co-occurring relatives by its longer dorsal fin lobe, as well as a host of other anatomical features. The Senegal jack grows to a known maximum length of 1 m. It is a coastal species, known to live semi-pelagically, inhabiting both the sea floor and surface waters to depths of around 200 m. The Senegal jack is a predatory species, taking fish, crabs and shrimps as its main prey items. The species reaches sexual maturity at 21 cm in females and 24 cm in males, with spawning occurring in two periods; February to April and September to November. The species is of minor importance to fisheries, and is not discriminated from other jacks in catch statistics. It is taken by trawls, seines and hook and line, and sold fresh or preserved.

<span class="mw-page-title-main">Tille trevally</span> Species of fish

The tille trevally, also known as the tille kingfish, is a species of large marine fish classified in the jack family, Carangidae. The tille trevally is distributed through the tropical and subtropical waters of the Indo-Pacific region, ranging from South Africa in the west to Fiji, Japan and Australia in the east. The species is best distinguished by its rounded, strongly convex anterior profile, with other detailed anatomical features also useful. The tille trevally reaches a maximum length of 80 cm and a weight of 7.2 kg. It is predominantly an inshore species, inhabiting coastal reef and lagoon environments, although has been recorded on deep offshore seamounts. It is a predatory fish, taking various species of fish and crustaceans as prey, with little known of its reproductive cycle. It is of minor importance to fisheries throughout its range, taken by hook and line, gill nets and purse seines. The tille trevally is also considered a good game fish, and an excellent table fish. The species acquired its scientific and common names from a local name used by Pondicherry fishermen, koton tille, which Georges Cuvier then used when he named the species in 1833.

References

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  10. 1 2 Goodson, Gar; Phillip J. Weisgerber (1988). Fishes of the Pacific Coast: Alaska to Peru, Including the Gulf of California and the Galapagos Islands. U.S.: Stanford University Press. p. 25. ISBN   978-0-8047-1385-6.
  11. 1 2 Klimley, A.P.; S.B. Butler (1988). "Immigration and emigration of a pelagic fish assemblage to seamounts in the Gulf of California related to water mass movements using satellite imagery". Marine Ecology Progress Series. 49 (1): 11–20. Bibcode:1988MEPS...49...11K. doi: 10.3354/meps049011 .
  12. 1 2 David, Usan (2006). "Study of a Caranx caballus fishery in Las Perlas Archipelago, Panama" (PDF). Master of Science Thesis. Heriot-Watt University, Edinburgh. Retrieved 2008-09-30.{{cite journal}}: Cite journal requires |journal= (help)
  13. Abitia-Cardenas, L.A.; F. Galvan-Magaña; J. Rodriguez-Romero (1997). "Food habits and energy values of prey of striped marlin, Tetrapterus audax, off the east coast of Mexico" (PDF). Fishery Bulletin. 95 (2): 360–368. Retrieved 2008-09-30.
  14. Sumida, Barbara Y.; H.G. Moser; E.H. Ahlstrom (1985). "Descriptions of Larvae of California Yellowtail, Seriola lalandi and three other Carangids from the Eastern Tropical Pacific: Chloroscombrus orqueta, Caranx caballus, and Caranx sexfasciatus" (PDF). CalCOFI Report. XXVI: 139–159. Archived from the original (PDF) on 2008-12-01. Retrieved 2008-09-30.
  15. Rodriguez Navarro, M.D.C; Hernandez Vazquez, S.; Funes Rodriguez, R.; Flores Vargas, R. (2001). "Distribution and abundance of fish larvaes of Haemulidae, Sciaenidae and Carangidae on the continental shelf from Jalisco and Colima, Mexico". Boletin del Centro de Investigaciones Biologicas Universidad del Zulia. 35 (1): 1–24.
  16. Fichter, George S.; Phil Francis; Tom Dolan; Ken Martin; Harry McKnaught (2001). Fishing: A Guide to Fresh and Salt-Water. Macmillan. p. 20. ISBN   978-1-58238-141-1.
  17. "Jack, Green". igfa.org. IGFA. Retrieved 14 June 2019.