Orange-spotted trevally

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Orange-spotted trevally
C. bajad.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Carangiformes
Family: Carangidae
Genus: Carangoides
Species:
C. bajad
Binomial name
Carangoides bajad
(Forsskål, 1775)
Carangoides bajad distribution.png
Approximate range of the orangespotted trevally
Synonyms
  • Scomber ferdau bajad
    Forsskål, 1775
  • Caranx bajad
    (Forsskål, 1775)
  • Caranx immaculatus
    Ehrenberg, 1833
  • Caranx auroguttatus
    Cuvier, 1833
  • Carangoides auroguttatus
    (Cuvier, 1833)
  • Caranx fulvoguttatus var. flava
    Klunzinger, 1871

The orange-spotted trevally, Carangoides bajad (also known as the gold-spotted trevally) is a species of inshore marine fish in the jack family, Carangidae. The species is fairly common in tropical to subtropical waters of the Indo-Pacific, ranging from Madagascar in the west to Japan in the east, typically inhabiting inshore reefs. The species has characteristic orange-yellow spots on its sides, although counts of fin rays and scutes are needed to distinguish it from related species with similar colouring. Orange-spotted trevallies are powerful predators, taking a variety of small fish, nekton, and crustaceans, and reach sexual maturity around 25 cm long. It is a moderately large fish, reaching a maximum known length of 55 cm. The species is occasionally taken by fishermen throughout its range, and is generally considered to be bycatch. The exception to this is in the southern Persian Gulf, where it makes up a large proportion of the fishery.

Contents

Taxonomy and naming

The orange-spotted trevally is classified within the genus Carangoides , a group of fish commonly called jacks and trevallies. Carangoides falls into the jack and horse mackerel family Carangidae, the Carangidae are part of the order Carangiformes. [2]

The species was first scientifically described by the Swedish naturalist Peter Forsskål in 1775 based on a specimen taken from the Red Sea which he designated to be the holotype. [3] The specific epithet is an Arabic name of the fish (although it is now usually applied to a catfish, Bagrus bajad , which Forsskål also named), with the letter "j" transcribing a /j/ sound; Forsskål used this technique to name a number of Red Sea fish species. [4] Forsskål at first gave the new taxon subspecies status as Scomber ferdau bajad, relating it to the mackerels, and especially Scomber ferdau, which would later also be transferred to Carangoides. The taxon was later given a species rank, becoming Scomber bajad, then Caranx bajad, before being transferred to its current position as Carangoides bajad. [5]

The species was also independently renamed three times after Forsskål's description, the first coming from Christian Gottfried Ehrenberg, who named the species Caranx immaculatus, although he did not accurately publish the name, leading Georges Cuvier to rename the fish as Caranx auroguttatus in 1833, which was later transferred to Carangoides. In 1871, Carl Benjamin Klunzinger once again proposed a new subspecies (or variety) name for the fish, Caranx fulvoguttatus var. flava. [5] All names except Carangoides bajad are considered to be junior synonyms under the ICZN rules, and are rendered invalid and not used.

Description

An orange-yellow orange-spotted trevally in Egypt Carangoides bajad.jpg
An orange-yellow orange-spotted trevally in Egypt

The orange-spotted trevally has the typical body profile of its genus, having a moderately deep, compressed oblong shape, with the dorsal profile more convex than the ventral profile. It is a moderately large fish, with adults typically reaching a size of 45–60 cm. [6] The largest fish recorded was 72 cm, but are most common at sizes below 40 cm. [7] [6] The dorsal profile of the head is quite straight from the snout to the nape, with an eye diameter smaller than the snout length. Both jaws contain narrow bands of villiform teeth, with these bands becoming wider anteriorly. The dorsal fin is in two parts, the first containing eight spines and the second of one spine followed by 24 to 26 soft rays. [8] The anal fin is similar to the second dorsal fin, although slightly shorter, and consists of two anteriorly detached spines followed by one spine and 21 to 24 soft rays. Another of the major diagnostic features for the species is the length of the anal fin lobe in comparison to the head length, with C. bajad having a head length longer than the anal fin lobe. [8] The lateral line has a slight, broad anterior curve, with this curved section being longer than the posterior straight section. The straight section of the lateral line has 14 to 26 scales followed by 20 to 30 scutes. [9] The chest of the orange-spotted trevally is completely scaled, occasionally having a narrow naked region on the underside near the ventral fins. It has 24 vertebrae and 25 to 43 gill rakers. [8]

The orange-spotted trevally has a silvery grey- to brassy-coloured body, becoming paler to a silvery white ventrally. Many conspicuous orange to yellow spots occur on the sides of the fish, giving the species its name, and make for an easy way to identify the fish in the field. [8] An entirely yellow variant has been reported, [10] although the fish is thought to be able to rapidly change between its normal colour configuration and this colour. [8] The colour of the fins ranges from hyaline to lemon yellow, and no dark opercular spot is present. [3]

Distribution and habitat

Group of orange-spotted trevallies in the red Sea, Egypt Grupo de jureles lentejuela (Carangoides bajad), parque nacional Ras Muhammad, Egipto, 2022-03-28, DD 20.jpg
Group of orange-spotted trevallies in the red Sea, Egypt

The orange-spotted trevally inhabits the tropical to subtropical waters of the Indian and western Pacific Oceans. It is distributed from Madagascar and the Comoros Islands in the west, [11] northwards to the Red Sea, the Gulf of Aden, and the Persian Gulf, and probably east toward India, although few fish have been recorded in this region. [5] It is commonly found from the Gulf of Thailand to Okinawa, Japan in the east, and southward to Indonesia, the Philippines, and New Britain. [8] In 2005, orange-spotted trevally reportedly had been caught in the Mediterranean Sea, indicating they had become Lessepsian migrants, passing through the Suez Canal to extend their range. [12] This claim has been disputed, though, with the source of the report coming under question due to the presence of other species unknown to the Mediterranean shown in the purported photograph. [13]

The orange-spotted trevally is a coastal species, most common in inshore waters over rocky and coral reefs, where it is found both solitary and in schools at depths of 2 to 50 m. They are often observed patrolling the edges of seaward reefs, and have been known to mingle with Parupeneus cyclostomus . [14]

Biology and fisheries

The orange-spotted trevally is a strong-swimming predator that takes a variety of prey, including fish, crustaceans, and nekton. [15] They reach sexual maturity around 1–3 years of age when they are about 15–30 cm in length. [6] [11] The main spawning period in the Persian Gulf is between June and September. [6] Whether further differences in the spawning period occurs throughout its range is currently unknown. This species also exhibits seasonal oscillation in growth rate, which was fastest during November–April and slowest during May–September. [11]

The orange-spotted trevally is occasionally taken throughout its range by hook and line, gill nets, and other artisanal gear, although in most areas it is bycatch and does not form a large part of these fisheries. [8] One fishery is highly dependent on the species in the southern Persian Gulf. There, the orange-spotted trevally is one of the most common fish found just above the sea floor, and is taken by wire traps and sold fresh at local markets. The combined catch of C. bajad and Gnathanodon speciosus totals around 1100 tonnes per year. [16] The development of the fishing fleet of the United Arab Emirates has caused a number of species to be overexploited, but the orange-spotted trevally is still being taken at sustainable levels. [16]

Related Research Articles

<span class="mw-page-title-main">Golden trevally</span> Species of fish

The golden trevally, also known as the golden kingfish, banded trevally or king trevally, is a species of large marine fish classified in the jack and horse mackerel family Carangidae, and the only member of the monospecific genus Gnathanodon. The golden trevally is widely distributed throughout the tropical and subtropical waters of the Indian and Pacific Oceans, ranging from South Africa in the west to Central America in the east, extending to Japan in the north and Australia in the south. The species predominantly occupies inshore waters where it inhabits both reef and sandy substrates. The golden trevally is easily distinguished from its relatives by its fleshy, rubbery lips and unique colouration, which ranges from bright yellow with black bars as a juvenile to a golden-silvery colour as an adult. It is known to grow to 120 centimetres (47 in) in length and 15 kilograms (33 lb) in weight. The golden trevally schools as a juvenile, often closely following larger objects including sharks and jellyfish. The species uses its protractile jaws to suck out prey from the sand or reef, and consumes a variety of fish, crustaceans and molluscs. Spawning aggregations gather at night at different times of the year throughout its range. The golden trevally is a considerable constituent of several Middle Eastern fisheries and being of minor importance to many others, with a worldwide annual catch of 1,187 metric tons to 3,475 metric tons recorded between 2000 and 2010. The golden trevally is a popular gamefish, taken by bait, lure, fly and also spear throughout its range. Several Asian countries currently farm the fish in caged aquaculture. Due to their brilliant colouration, juveniles are popular in marine aquaria.

<span class="mw-page-title-main">Shrimp scad</span> Species of fish

The shrimp scad, is a species of widespread tropical marine fish of the jack family, Carangidae. The shrimp scad is widely distributed in the tropical and subtropical western Indian Ocean and areas of the eastern Pacific Ocean, ranging from South Africa in the west to Hawaii in the east, including Japan and Australia to the north and south. The species is commonly found on inshore reefs and sandy substrates. It has the common body profile of a scad, and may be difficult to differentiate from others in the genus Alepes. It is one of the larger scads, growing to 40 cm, but often is encountered at much smaller sizes. The shrimp scad often forms large schools, and is carnivorous, consuming a variety of crustaceans and small fish. It is of moderate importance to fisheries throughout its range.

<span class="mw-page-title-main">Malabar trevally</span> Species of fish

The Malabar trevally, also known as the Malabar jack, Malabar kingfish or nakedshield kingfish, is a species of large inshore marine fish of the jack family, Carangidae. It is distributed throughout the Indian and west Pacific Oceans from South Africa in the west to Japan and Australia in the east, inhabiting reefs and sandy bays on the continental shelf. The Malabar trevally is similar to many of the other species in the genus Carangoides, with the number of gill rakers and the grey-brown colour of the tongue being the diagnostic features. The Malabar trevally is a predator, taking a variety of small fish, cephalopods and crustaceans. The species is of minor economic importance throughout its range, caught by a variety of net and handline methods.

<i>Carangoides</i> Genus of fishes

Carangoides is a genus of tropical to subtropical marine fishes in the jack family, Carangidae. They are small- to large-sized, deep-bodied fish characterised by a certain gill raker and jaw morphology, often appearing very similar to jacks in the genus Caranx. They inhabit the subtropical and tropical regions of the Indian, Pacific, and Atlantic Oceans, often occupying coastal areas, including reefs, bays, and estuaries, rarely venturing far offshore. They are all predatory fishes, taking a variety of smaller fishes, crustaceans and cephalopods as prey. The genus was first erected in 1851 by Pieter Bleeker for an unknown taxon and currently contains 20 species. Many make up significant proportions of various fisheries, although a number of ciguatera cases have been attributed to them.

<span class="mw-page-title-main">Longfin trevally</span> Species of fish

The longfin trevally, also known as the longfin kingfish, longfin cavalla or armed trevally, is a species of inshore marine fish in the jack family, Carangidae. The species is common in tropical to subtropical waters of the Indo-Pacific, ranging from South Africa in the west to Japan in the east, typically inhabiting inshore reefs and bays. The species is easily distinguished by its elongate dorsal and anal fin lobes and filamentous dorsal rays, as well as its scaleless breast. Longfin trevally are pelagic predators, taking a variety of small fish, cephalopods and crustaceans, and reach sexual maturity at around 21 cm. The species has a maximum known length of 57 cm and weight of 3.5 kg. The longfin trevally has a very complex taxonomic history which is closely intertwined with another currently valid species, Carangoides ciliarius, which may yet prove to be synonymous. Longfin trevally are of minor importance to fisheries throughout their range and are considered good table fish, and are occasionally taken by anglers.

<span class="mw-page-title-main">Longnose trevally</span> Species of fish

The longnose trevally, also known as the tea-leaf trevally, club-nosed trevally, grunting trevally or dusky trevally, is a species of inshore marine fish in the jack family, Carangidae. The species is distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans from South Africa to New Zealand and Japan, inhabiting coastal waters, especially reefs, to a depth of 90 m. The longnose trevally is distinguished from similar species by a combination of a scaleless breast and the number of gill rakers and fin rays. It is a moderately large fish, growing to a maximum known length of 72 cm and 4.35 kg. The longnose trevally is a predatory fish, consuming small fish, crustaceans and molluscs. The species is of minor commercial importance throughout its range, and is considered to be a good table fish.

<span class="mw-page-title-main">Coastal trevally</span> Species of fish

The coastal trevally, also known as the onion trevally, Japanese trevally or bluefin kingfish, is a species of inshore marine fish in the jack family Carangidae. The species is distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans, from South Africa in the west to Japan and New Caledonia in the east, reaching as far south as Australia. The species is found on deep coastal reefs, both in schools and as solitary individuals, where they prey on small midwater organisms including crustaceans, small fish and cephalopods. The species is taken as bycatch in a number of fisheries throughout its range by a number of fishing methods and is of little commercial value, but is considered to be a good table fish. A mistype in the original volume in which Eduard Rüppell named the species led to the combination Carangoides caeruleopinnatus, which has incorrectly spread through the literature.

<span class="mw-page-title-main">Shadow trevally</span> Species of fish

The shadow trevally, also known as the shadow kingfish, twothread trevally or Aldabra trevally, is a species of inshore marine fish in the jack family Carangidae. The species is patchily distributed throughout the tropical and subtropical waters of the Indian and west Pacific Oceans, from South Africa in the west to Japan and Samoa in the east, reaching as far south as Indonesia and New Caledonia. It is most easily distinguished from similar species by as series of dark rectangular blotches under the second dorsal fin, giving a 'shadowed' appearance, from which its common name is derived. The shadow trevally is a reasonably large fish, growing to 85 cm in length and at least 2.6 kg in weight. It inhabits shallow coastal waters, including reefs, bays, and estuaries, where it takes small fish and benthic crustaceans as prey. Nothing is known of the species' ecology and reproductive biology. It is of little importance to fisheries, and is occasionally taken by bottom trawls and other artisanal fishing gear.

<i>Caranx</i> Genus of fishes

Caranx is a genus of tropical to subtropical marine fishes in the jack family Carangidae, commonly known as jacks, trevallies and kingfishes. They are moderate- to large-sized, deep-bodied fishes which are distinguished from other carangid genera by specific gill raker, fin ray and dentition characteristics. The genus is represented in the Pacific, Indian and Atlantic Oceans, inhabiting both inshore and offshore regions, ranging from estuaries and bays to deep reefs and offshore islands. All species are powerful predators, taking a variety of fish, crustaceans and cephalopods, while they in turn are prey to larger pelagic fishes and sharks. A number of fish in the genus have a reputation as powerful gamefish and are highly sought by anglers. They often make up high amounts of the catch in various fisheries, but are generally considered poor to fair table fishes.

<span class="mw-page-title-main">Whitefin trevally</span> Species of fish

The whitefin trevally, also known as the horse trevally, is a species of deep water offshore fish in the jack family Carangidae. The species inhabits the tropical to temperate waters of the Indo-Pacific and central Pacific, ranging from South Africa in the west to Hawaii in the east. The whitefin trevally is a moderate-sized fish, growing to 37 cm, and is distinguished by a number of morphological traits, including fin size, gill raker count, and colour. It inhabits the continental shelf and slope at depths to 200 m over sand and mud substrates, where it preys on fish, crustaceans, and cephalopods. Studies in Japan indicate a length at sexual maturity of 17.4 cm on average, with spawning occurring between May and October, with each individual spawning multiple times. Whitefin trevallies are of high importance to fisheries in Japan, where they are taken by trawlers, although the catch numbers have halved since the 1980s. It is of minor importance elsewhere throughout its range, but is considered a good table fish.

<span class="mw-page-title-main">Blue trevally</span> Species of fish

The blue trevally, also known as the banded trevally, barred trevally, Ferdau's trevally or Forskaal's jackfish, is a common, widespread species of pelagic marine fish classified in the jack family, Carangidae. The blue trevally is distributed throughout the tropical and subtropical waters of the Indo-Pacific and central Pacific regions, ranging from South Africa in the west to Hawaii in the east. It is a moderately large fish, growing to a recorded maximum length of 70 cm, with the number of rays in the second dorsal fin and the colouring serving as diagnostic features of the species. The species inhabits waters to depths of 60 m, generally inhabiting reefs, beaches, lagoons, and areas with sandy substrates. It is a predatory fish, taking other fish, prawns, crabs, and molluscs, and very little is known of the species' reproductive biology. The blue trevally is of varying importance to fisheries throughout its range, with some regions having high catches of the fish. It is considered to be a gamefish, and is sought after for its excellent eating qualities.

<span class="mw-page-title-main">Yellowspotted trevally</span> Species of fish

The yellowspotted trevally, also known as the yellowspotted kingfish, goldspotted trevally, tarrum, or yellowspot, is a widespread species of large inshore marine fish in the jack family Carangidae. The yellowspotted trevally inhabits the tropical and subtropical waters of the western Indo-Pacific region, from South Africa in the west to Japan and Australia in the east. The species is known to grow to a maximum length of at least 1.2 m, and is distinguished by gill raker and fin morphology, as well as the distinctive golden spots which give the fish its name. The yellowspotted trevally generally prefers inshore rocky and coral reefs, but is occasionally found over deep offshore sand banks to a depth of 100 m. It is a predatory fish, taking fish, cephalopods, and crustaceans, and shows diet partitioning with other trevallies in studies conducted in Australian waters. Reproduction is poorly studied, although observational evidence suggests spawning occurs in aggregations, probably during summer in South Africa. It is generally of minor importance to commercial fisheries throughout its range, but is considered an excellent sportfish by anglers and spearfishermen, and a good table fish.

<span class="mw-page-title-main">Bumpnose trevally</span> Species of fish

The bumpnose trevally, also known as the bumpnose kingfish or onion kingfish, is a species of relatively small inshore marine fish classified in the jack family Carangidae. The bumpnose trevally is fairly common in the tropical and subtropical waters of the Indo-west Pacific region, ranging from South Africa in the west to Japan and Samoa in the east. It is a small species by carangid standards, reaching a maximum known length of 32 cm, and can be distinguished from the similarly shaped Carangoides armatus by a distinct 'bump' on the snout, which gives the fish its common name. The species inhabits coastal waters, often living along bays and beaches, where it takes shrimp, small crabs, and juvenile fish as prey. The bumpnose trevally is of minor importance to fisheries throughout its range, taken by hook and line, trawls, and seine nets. It is also of minor importance to anglers, taken by baits from beaches and piers, and is considered a modest table fish.

<span class="mw-page-title-main">Duskyshoulder trevally</span> Species of fish

The duskyshoulder trevally or epaulet trevally, is a species of small inshore marine fish in the jack family, Carangidae. It is distributed through the eastern Indian and western Pacific Oceans, ranging from eastern India to northern Australia and Taiwan. It is relatively small by carangid standards, reaching only 27 cm maximum length, and can be distinguished by the large, black blotches on its shoulders. The duskyshoulder trevally is an inshore fish living in waters less than 50 m deep, over sandy substrates in bays and on the continental shelf. It is a predatory fish, taking demersal fishes, crustaceans, and cephalopods, with nothing known of its reproductive habits. It is of little value to fisheries, often taken as bycatch in prawn trawling operations.

<span class="mw-page-title-main">Coachwhip trevally</span> Species of fish

The coachwhip trevally, also known as the oblong trevally or oblique-banded trevally, is a species of inshore marine fish classified in the jack family Carangidae. The coachwhip trevally is distributed through the Indo-west Pacific region, ranging from South Africa in the west to Fiji and Japan in the east. It is a moderately large fish, growing to a known maximum length of 46 cm and can be distinguished from similar species by an array of detailed morphological features including dentition, fin ray counts and scale patterns. The coachwhip trevally inhabits coastal waters throughout its range, known to prefer estuarine waters in a number of localities. Nothing is known of its diet or reproductive biology, and is of little importance to fisheries, occasionally taken as bycatch in trawl and hook and line fisheries.

<span class="mw-page-title-main">Barcheek trevally</span> Species of fish

The barcheek trevally, also known as the barcheek kingfish, shortridge trevally or oblique-banded trevally, is a species of moderately large marine fish of the jack family Carangidae. The barcheek trevally is distributed throughout the tropical waters of the Indo-west Pacific region, ranging from South Africa in the west to Japan, Australia and a number of small central Pacific islands in the east. The species inhabits inshore and offshore waters, found along the slopes of lagoons and out to deeper reefs on the continental shelf, where it preys on small fish and benthic crustaceans. It is a moderately large fish, growing to a maximum recorded length of 50 cm, and can be distinguished from similar species by its somewhat protruding lower jaw and the dark banding on its operculum. It is of minor importance to fisheries throughout its range, taken by trawling, hook and line methods and various inshore fish netting methods.

<span class="mw-page-title-main">Brownback trevally</span> Species of fish

The brownback trevally, also known as the brown-backed trevally, is a species of small inshore marine fish classified in the jack family, Carangidae. The brownback trevally is distributed in two populations through the tropical waters of the Indo-west Pacific region, ranging from the Persian Gulf east to India, South East Asia and the Indonesian islands. The species is distinguished from similar species by its completely scaled breast and black-tipped second dorsal fin, and is known to reach a maximum length of 25 cm. The brownback trevally inhabits inshore waters including bays and estuaries, where it preys on demersal crustaceans and small fish. Other aspects of its biology are poorly known, and it is of minor importance to fisheries, occasionally caught by hook and line or trawls. William Smith-Vaniz has recently suggested the two distinct populations may actually represent two distinct species.

<span class="mw-page-title-main">Imposter trevally</span> Species of fish

The imposter trevally, also known as the imposter jack or white-tongued trevally, is a species of small coastal marine fish in the jack family Carangidae. The imposter trevally is distributed through the tropical waters of Indian and west Pacific oceans, from the Gulf of Oman in the west to Japan and Australia in the east. The species is quite similar to the Malabar trevally, but can be distinguished by its gill raker count, and is identifiable from other species by having a white to pale grey tongue. It is a relatively small species, growing to a maximum recorded length of 30 cm. The imposter trevally inhabits coastal waters of the continental shelf in depths of up to 140 m, often associating with closely related carangid species. It preys on a range of small fish, crustaceans and cephalopods, but little is known of its reproductive biology. The imposter trevally is of minor importance to fisheries throughout its range, taken by hook and line, bottom trawls and several types of artisanal gear.

<span class="mw-page-title-main">Blacktip trevally</span> Species of fish

The blacktip trevally, also known as the blacktip kingfish or yellowtail kingfish, is a species of large marine fish classified in the jack family Carangidae. The blacktip trevally is distributed throughout the tropical to subtropical Indian and West Pacific Oceans, ranging from South Africa in the west to Fiji, Japan and northern Australia in the east. It inhabits coastal waters throughout its range, preferring moderately deep clear waters over rocky and coral reefs. The blacktip trevally is easily distinguished by its yellow fins and a dark upper caudal fin lobe which gives the species its common name, as well as a host of other anatomical features. The species is known to reach a maximum size of 1 m. It is a benthopelagic predator, commonly forming small shoals where it takes a variety of fishes, cephalopods and crustaceans as prey. Little is known of reproduction in the species, and spawning is assumed to take place in more tropical regions of its range, with juveniles known to inhabit bays and large estuaries. Blacktip trevally are often caught using hook and line and various nets in commercial fisheries although don't make up a large part of the market. They are also popular with anglers due to their fighting ability and decent table qualities.

<span class="mw-page-title-main">Brassy trevally</span> Species of fish

The brassy trevally, Caranx papuensis is a species of large marine fish classified in the jack family, Carangidae.

References

  1. Smith-Vaniz, W.F. & Williams, I. (2017) [errata version of 2016 assessment]. "Carangoides bajad". IUCN Red List of Threatened Species . 2016: e.T46080857A115391273. doi: 10.2305/IUCN.UK.2016-3.RLTS.T46080857A46664509.en . Retrieved 19 February 2022.
  2. J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016). Fishes of the World (5th ed.). Wiley. pp. 380–387. ISBN   978-1-118-34233-6.
  3. 1 2 Ludwig, Albert Carl; Gotthilf Günther (1860). Catalogue of the Fishes in the British Museum. British Museum (Natural History). Dept. of Zoology. p. 238.
  4. Baheyeldin, Khalid (October 17, 2004). "Forsskål's use of Arabic names for fish species". The Baheyeldin dynasty.
  5. 1 2 3 Froese, Rainer; Pauly, Daniel (eds.) (2008). "Carangoides bajad" in FishBase . January 2008 version.
  6. 1 2 3 4 Lin, Yu‐Jia; Rabaoui, Lotfi; Maneja, Rommel H.; Qurban, Mohammad A.; Abdulkader, Khaled; Al‐Nazry, Hussain; Roa‐Ureta, Ruben H. (2019). "Life history traits and temporal trends of abundance of the orange‐spotted trevally ( Carangoides bajad ) from Saudi waters of the Gulf". Journal of Fish Biology. 95 (5): 1184–1194. doi:10.1111/jfb.14116. ISSN   0022-1112. PMID   31389013.
  7. Randall, John E. (1995). Coastal Fishes of Oman. Honolulu: University of Hawaiʻi Press. p. 183. ISBN   0-8248-1808-3.
  8. 1 2 3 4 5 6 7 Carpenter, Kent E.; Volker H. Niem, eds. (1999). FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Volume 4: Bony fishes part 2 (Mugilidae to Carangidae) (PDF). Rome: Food and Agriculture Organization of the United Nations. p. 2694. ISBN   92-5-104587-9.
  9. Lin, Pai-Lei; Shao, Kwang-Tsao (May 15, 1999). "A Review of the Carangid Fishes (Family Carangidae) From Taiwan with Descriptions of Four New Records". Zoological Studies. 38 (1): 33–68.
  10. Myers, R.F.; Lieske, E. (2004). Coral Reef Guide Red Sea. London: Collins. ISBN   978-0-00-715986-4.
  11. 1 2 3 Grandcourt, E.M.; T.Z. Al Abdessalaam; F. Francis; A. Al Shamsi (2004). "Population biology and assessment of representatives of the family Carangidae Carangoides bajad and Gnathanodon speciosus (Forsskal, 1775), in the Southern Arabian Gulf". Fisheries Research. 69 (3): 331–341. doi:10.1016/j.fishres.2004.06.008.
  12. Öztürk, Bayrum, ed. (2005). Marine Life of Turkey in the Aegean and Mediterranean Seas. Istanbul: Turkish Marine Research Foundation (TÜDAV). ISBN   975-8825-07-0.
  13. Golani, Daniel (2006). "The Indian scad (Decapterus russelli), (Osteichthyes: Carangidae), a new Indo-Pacific fish invader of the eastern Mediterranean". Scientia Marina. 70 (4): 603–605. doi: 10.3989/scimar.2006.70n4603 . ISSN   0214-8358.
  14. Siliotti, A. (2002). Fishes of the Red Sea. Verona: Geodia. p. 287. ISBN   88-87177-42-2.
  15. Blaber, S.J.M.; Milton, D.A.; Rawlinson, N.J.F.; Tiroba, G.; Nichols, P.V. (1990). "Diets of lagoon fishes of the Solomon Islands: predators of tuna baitfish and trophic effects of baitfishing on the subsistence fishery". Fisheries Research. 8 (3): 263–286. doi:10.1016/0165-7836(90)90027-S.
  16. 1 2 Grandcourt, E.; T.Z. Al Abdessalaam; F. Francis; A.T. Al Shamsi; K. Al Ali; S. Al Ali (2004). Biological reference points, resource status and management options for the key demersal species of Abu Dhabi Emirate (PDF). Environmental Research and Wildlife Development. pp. 1–28.[ permanent dead link ]