Ototriton Temporal range: Early Eocene | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Squamata |
Family: | Rhineuridae |
Genus: | † Ototriton Loomis, 1919 |
Type species | |
†Ototriton solidus Loomis, 1919 |
Ototriton is an extinct genus of rhineurid amphisbaenian or worm lizard from the Early Eocene of the western United States, including the type and only species Ototriton solidus. Paleontologist F. B. Loomis named Ototriton in 1919 on the basis of a single skull from the Wind River Formation in Wyoming, misinterpreting it as the skull of a salamander. [1] Unlike salamanders and like other rhineurids, Ototriton has a shovel-shaped snout that it presumably used for burrowing underground. Ototriton is one of the earliest known rhineurids and also one of the largest. [2]
Several other species have been assigned to Ototriton since Loomis named the genus in 1919. In 1928, paleontologist Charles W. Gilmore assigned a vertebra from the Bridger Formation of Wyoming, first classified as Glyptosaurus anceps, to Ototriton based on its large size, but later attributed it to the snake Lestophis crassus. In 1945, Gilmore and G. I. Jepsen named a new species of Ototriton, O. minor, on the basis of another skull from the Wind River Formation, distinguishing it from O. solidus on the basis of its smaller size. O. minor was later classified as its own genus, Jepsibaena. The most recent study to consider the specimen places it in the species Protorhineura hatcherii. [2]
Gastornis is an extinct genus of large flightless birds that lived during the late Paleocene and Eocene epochs of the Cenozoic era. The genus is currently thought to contain three or four distinct species, known from incomplete fossil remains, found in western-central Europe. More complete specimens are known from a fourth, North American species, which had previously been classified in the distinct genus Diatryma. Many scientists now consider Diatryma to be so similar to the other species of Gastornis that it should also be included in that genus. A fifth species, also previously classified in its own genus, is known from China.
Ceratosaurus was a carnivorous theropod dinosaur in the Late Jurassic period. This genus was first described in 1884 by American paleontologist Othniel Charles Marsh based on a nearly complete skeleton discovered in Garden Park, Colorado, in rocks belonging to the Morrison Formation. The type species is Ceratosaurus nasicornis.
Brontosaurus is a genus of gigantic quadruped sauropod dinosaurs. Although the type species, B. excelsus, had long been considered a species of the closely related Apatosaurus, researchers proposed in 2015 that Brontosaurus is a genus separate from Apatosaurus and that it contains three species: B. excelsus, B. yahnahpin, and B. parvus.
Charles Whitney Gilmore was an American paleontologist who gained renown in the early 20th century for his work on vertebrate fossils during his career at the United States National Museum. Gilmore named many dinosaurs in North America and Mongolia, including the Cretaceous sauropod Alamosaurus, Alectrosaurus, Archaeornithomimus, Bactrosaurus, Brachyceratops, Chirostenotes, Mongolosaurus, Parrosaurus, Pinacosaurus, Styracosaurus ovatus and Thescelosaurus.
Cimolestes is a genus of early eutherians with a full complement of teeth adapted for eating insects and other small animals. Paleontologists have disagreed on its relationship to other mammals, in part because quite different animals were assigned to the genus, making Cimolestes a grade taxon of animals with similar features rather than a genus of closely related ones. Fossils have been found in North America, where they first appeared during the Late Cretaceous. According to some paleontologists, they died out at the start of the Paleocene, while others report the genus from the early Eocene.
Edmontosaurus annectens is a species of flat-headed and duck-billed (hadrosaurid) dinosaur from the very end of the Cretaceous Period, in what is now North America. Remains of E. annectens have been preserved in the Frenchman, Hell Creek, and Lance Formations. All of these formations are dated to the late Maastrichtian stage of the Late Cretaceous Period, representing the last three million years before the extinction of the dinosaurs. Also, E. annectens is also from the Laramie Formation, and magnetostratigraphy suggests an age of 69-68 Ma for the Laramie Formation. Edmontosaurus annectens is known from numerous specimens, including at least twenty partial to complete skulls, discovered in the U.S. states of Montana, South Dakota, North Dakota, Wyoming and Colorado and the Canadian province of Saskatchewan. It was a large animal, up to approximately 12 metres (39 ft), potentially up to 15 m (49 ft) in length, with an extremely long and low skull. E. annectens exhibits one of the most striking examples of the "duckbill" snout common to hadrosaurs. It has a long taxonomic history, and specimens have at times been classified in the genera Diclonius, Trachodon, Hadrosaurus, Claosaurus, Thespesius, Anatosaurus and Anatotitan, before being grouped together in Edmontosaurus.
Leidyosuchus is an extinct genus of alligatoroid from the Late Cretaceous of Alberta. It was named in 1907 by Lawrence Lambe, and the type species is L. canadensis. It is known from a number of specimens from the middle Campanian age Dinosaur Park Formation. It was a medium-sized alligatorid, with a maximum skull length greater than 40 centimeters (16 in).
Brachyhyops is an extinct genus of entelodont artiodactyl mammal that lived during the Eocene Epoch of western North America and southeastern Asia. The first fossil remains of Brachyhyops are recorded from the late Eocene deposits of Beaver Divide in central Wyoming and discovered by paleontology crews from the Carnegie Museum of Natural History during the early 20th century. The type species, Brachyhyops wyomingensis, is based on a single skull and was named by E.H. Colbert in 1937, but was not officially described until 1938. During the latter half of the 20th century, additional specimens from North America have been recorded from Saskatchewan and as far south as Texas, indicating that Brachyhyops had a broad distribution and was well-dispersed throughout western North America.
Asiatosuchus is an extinct genus of crocodyloid crocodilians that lived in Eurasia during the Paleogene. Many Paleogene crocodilians from Europe and Asia have been attributed to Asiatosuchus since the genus was named in 1940. These species have a generalized crocodilian morphology typified by flat, triangular skulls. The feature that traditionally united these species under the genus Asiatosuchus is a broad connection or symphysis between the two halves of the lower jaw. Recent studies of the evolutionary relationships of early crocodilians along with closer examinations of the morphology of fossil specimens suggest that only the first named species of Asiatosuchus, A. grangeri from the Eocene of Mongolia, belongs in the genus. Most species are now regarded as nomina dubia or "dubious names", meaning that their type specimens lack the unique anatomical features necessary to justify their classification as distinct species. Other species such as "A." germanicus and "A." depressifrons are still considered valid species, but they do not form an evolutionary grouping with A. grangeri that would warrant them being placed together in the genus Asiatosuchus.
Eostrix is a genus of extinct primitive owls in the family Protostrigidae, along with Oligostrix and Minerva. These owls date from the early Eocene of the United States, Europe, and Mongolia. They have been described based on fossil remains. The genus was created by Pierce Brodkorb in 1971 to place a fossil species known until that time as Protostrix mimica.
Orthogenysuchus is an extinct genus of caimanine alligatorids. Fossils have been found from the Wasatch Beds of the Willwood Formation of Wyoming, deposited during the early Eocene. The type species is O. olseni. The holotype, known as AMNH 5178, is the only known specimen belonging to the genus and consists of a skull lacking the lower jaws. The braincase is filled in by the matrix and most of the suture lines between bones are indiscernible, making comparisons with other eusuchian material difficult.
"Crocodylus" affinis is an extinct species of crocodyloid from the Eocene of Wyoming. Fossils were first described from the Bridger Formation by American paleontologist Othniel Charles Marsh in 1871. Marsh described the species, along with every other species of crocodyloid in the Bridger Formation, under the genus Crocodylus. The known specimen of "Crocodylus" affinis is a skull found at Grizzly Buttes, Wyoming, measuring 13 inches in length on the upper surface. Recent phylogenetic studies of crocodyloids show that "C." affinis is not a species of Crocodylus, but a genus has not yet been erected to include the species. Other Bridger species such as Crocodylus clavis and Brachyuranochampsa zangerli have been synonymized with "C." affinis.
Saniwa is an extinct genus of varanid lizard that lived about 48 million years ago during the Eocene epoch. It is known from well-preserved fossils found in the Bridger and Green River Formations of Wyoming, and evidence indicates Saniwa also lived in Europe. The type species S. ensidens was described in 1870 as the first fossil lizard known from North America. Several other species have since been added, but their validity is uncertain. It is a close relative of Varanus, the genus that includes monitor lizards.
Afairiguana avius is an extinct iguanid lizard known from a nearly complete and articulated skeleton discovered in rocks of the Early Eocene-aged Green River Formation of Wyoming, United States. As of the initial description, the skeleton represents the oldest complete iguanian from the Western Hemisphere, and is the oldest representative of the extant iguanid family of anoles, Polychrotidae.
Kritosaurini is a clade of saurolophine hadrosaurid dinosaurs from the Late Cretaceous.
Spathorhynchus is an extinct genus of amphisbaenians or worm lizards that existed from the Middle Eocene to the Early Oligocene in what is now Wyoming. It includes two species, the type species S. fossorium, named in 1973 from the Middle Eocene Bridger and Wind River Formations, and the species S. natronicus, named in 1977 from the Lower Oligocene White River Formation. Spathorhynchus belongs to the family Rhineuridae, which includes many other extinct species that ranged across North America at various times in the Cenozoic but only has one surviving member, Rhineura floridana, from Florida. Spathorhynchus differs from all other rhineurids except Dyticonastis from the Late Oligocene-Early Miocene of Oregon in having a slightly widened, spatula-shaped snout tip with a low angle of about 30 degrees. The two taxa may be closely related, having evolved in isolation in western North America after the formation of the Rocky Mountains separated them from rhineurids further east.
Macrorhineura is an extinct genus of rhineurid amphisbaenian or worm lizard, including the type and only species Macrorhineura skinneri, named in 1970 on the basis of the front half of a skull from the Early Miocene Sharps Formation in Wounded Knee, South Dakota. Although the skull is incomplete, features such as a pointed, shovel-shaped snout indicate that it belongs to the family Rhineuridae. Within Rhineuridae, Macrorhineura is most closely related to Ototriton and Hyporhina, two genera from the Eocene and Oligocene of Colorado and Wyoming, based on the shared feature of equally sized dentary teeth in the lower jaw. Together they form a clade or evolutionary grouping of mid-continental rhineurids, which became isolated from a more western clade of rhineurids that includes Dyticonastis and Spathorhynchus. Rhineurids were relatively common across much of North America during the Paleogene, but their range contracted in the Neogene as the climate became colder, leaving only one living species in Florida, Rhineura floridana. The presence of Macrorhineura in the Miocene shows that mid-continental rhineurids persisted into the Neogene, although by this time their distribution range was already shrinking.
Hyporhina is an extinct genus of amphisbaenians or worm lizards that lived from the Late Eocene to the Middle Oligocene in what is now the western United States.
Anolbanolis is an extinct genus of iguanian lizards that lived in the Bighorn Basin of what is now Wyoming during the earliest Eocene. The type species A. banalis was named by paleontologist Krister Smith in 2009 from a collection of isolated skull fragments found in a layer of the Willwood Formation that dates to a brief period of global warming called the Paleocene-Eocene thermal maximum (PETM) about 56 million years ago. The genus name Anolbanolis means "poor anole" from the Greek anolbos and the name Anolis, in reference to the scrappy nature of known fossil material and its close resemblance to lizards in the genus Anolis. Smith suggested that Anolbanolis may be a close relative of Anolis or Polychrus, which are common today in Central and South America but not found as far north as Wyoming, fitting with the idea that the Bighorn Basin was warmer and wetter in the Eocene than it is currently. According to Smith, the species name banalis is a reference to it being a "banal" lizard in the Willwood, being "quite abundant in the type locality," and not unusual because "the discovery of Eocene boreal fossil members of living subtropical and tropical clades is becoming commonplace." In 2011 Smith named a second species of Anolbanolis, A. geminus, which lived in the Bighorn Basin shortly after the PETM and was about 50 percent larger than A. banalis. Smith concluded on the basis of features in A. geminus that Anolbanolis is more closely related to Anolis than it is to Polychrus.
Babibasiliscus is an extinct genus of casquehead lizard that lived in what is now Wyoming during the early Eocene, approximately 48 million years ago. The genus is known from a single species, Babibasiliscus alxi, which was named by paleontologist Jack Conrad in 2015 on the basis of a fossilized skull from the Bridger Formation in the Green River Basin. The name Babibasiliscus comes from the Shoshoni word babi, meaning "older male cousin", and Basiliscus, a modern-day genus of casquehead lizards. The specimen is undeformed and nearly complete except for the tip of the snout and the top of the skull, making it unclear whether the distinctive bony crest of living corytophanids was present in prehistoric relatives like Babibasiliscus. The skull is about 42 millimetres (2 in) in length and the entire body is estimated to have been about 0.6 metres (2 ft) long. Bones on the right side of lower jaw of the specimen are thickened and fused together, suggesting that the jaw had broken and healed when the animal was alive.
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