Squamata is the largest order of reptiles, comprising lizards, snakes and amphisbaenians, which are collectively known as squamates or scaled reptiles. With over 10,000 species, it is also the second-largest order of extant (living) vertebrates, after the perciform fish, and roughly equal in number to the Saurischia. Members of the order are distinguished by their skins, which bear horny scales or shields. They also possess movable quadrate bones, making it possible to move the upper jaw relative to the neurocranium. This is particularly visible in snakes, which are able to open their mouths very wide to accommodate comparatively large prey. Squamata is the most variably sized order of reptiles, ranging from the 16 mm (0.63 in) dwarf gecko to the 5.21 m (17.1 ft) green anaconda and the now-extinct mosasaurs, which reached lengths of over 14 m (46 ft).
Anguidae refers to a large and diverse family of lizards native to the Northern Hemisphere. Common characteristics of this group include a reduced supratemporal arch, striations on the medial faces of tooth crowns, osteoderms, and a lateral fold in the skin of most taxa. The group includes the slowworms, glass lizards, and alligator lizards, among others. The family is divided into three subfamilies, and contains approximately 100 species in ten genera.
Ophidia is a group of squamate reptiles including modern snakes and all reptiles more closely related to snakes than to other living groups of lizards.
Mosasaurs are an extinct group of large marine reptiles containing 38 genera in total. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. Mosasaurs probably evolved from an extinct group of aquatic lizards known as aigialosaurs in the Early Cretaceous. During the last 20 million years of the Cretaceous period, with the extinction of the ichthyosaurs and pliosaurs, mosasaurs became the dominant marine predators. They became extinct as a result of the K-Pg event at the end of the Cretaceous period, about 66 million years ago.
Aeolosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. Aeolosaurus is well known for a titanosaur, as it is represented by the remains of several individuals belonging to at least three species. However, like most titanosaurs, no remains of the skull are known. The holotype of Aeolosaurus rionegrinus consists of a series of seven tail vertebrae, as well as parts of both forelimbs and the right hindlimb. It was discovered in the Angostura Colorada Formation in Argentina, which dates from the Campanian stage of the Late Cretaceous, about 83 to 74 million years ago.
Carcharodontosaurids were a group of carnivorous theropod dinosaurs. In 1931 Ernst Stromer named Carcharodontosauridae as a family, which, in modern paleontology, indicates a clade within Carnosauria. Carcharodontosaurids included some of the largest land predators ever known: Giganotosaurus, Mapusaurus, Carcharodontosaurus, and Tyrannotitan all rivaled or slightly exceeded Tyrannosaurus in length. A 2015 paper by Christophe Hendrickx and colleagues gives a maximum length estimate of 14 meters for the largest carcharodontosaurids, while the smallest carcharodontosaurids were estimated to have been at least 6 meters long.
Amazonsaurus is a genus of diplodocoid sauropod dinosaur from the Early Cretaceous Period of what is now South America. It would have been a large-bodied quadrupedal herbivore with a long neck and whiplash tail. Although more derived diplodocoids were some of the longest animals ever to exist, Amazonsaurus was probably not more than 12 meters (40 ft) long. Gregory S. Paul estimated in 2010 its weight at 5000 kg.
Adamantisaurus is a poorly-known genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. It is only known from six tail vertebrae but, as a sauropod, it can be assumed that this dinosaur was a very large animal with a long neck and tail.
Pycnonemosaurus is a genus of carnivorous theropod dinosaur that belonged to the family Abelisauridae. It was found in the Upper Cretaceous Bauru-type red conglomerate sandstones of the Adamantina Formation, Mato Grosso, Brazil, and it lived about 70 million years ago during the Late Cretaceous.
Yabeinosaurus is an extinct genus of lizard from the Early Cretaceous Jehol Group of northeastern China. The type species Yabeinosaurus tenuis is known from many well-preserved skeletons belonging to both juvenile and adult individuals. For about 60 years Yabeinosaurus was known only from juvenile specimens, leading scientists to believe that it was a small lizard with weakly developed bones. Because of this, it was thought to be closely related to geckos. Larger specimens up to 35 centimetres (14 in) long were first described in 2005, showing that Yabeinosaurus was a relatively large lizard when fully grown. Recent phylogenetic analyses indicate that Yabeinosaurus is not closely related to geckos but rather a very basal ("primitive") lizard close to the split between Iguania and Scleroglossa, one of the earliest divergences in the evolutionary history of lizards. Whether or not it lies outside this split or within Scleroglossa is uncertain.
Palaeosaniwa canadensis is an extinct species of carnivorous lizard from the late Cretaceous of North America. The name, given by Charles Whitney Gilmore in 1928, means "ancient Saniwa from Canada".
Kuwajimalla kagaensis is an extinct species of plant-eating lizard from the Early Cretaceous Kuwajima Formation in Japan. K. kagaensis is the type species.
Pristiguana is an extinct genus of primitive iguanids from the Maastrichtian Marília Formation of Brazil. The type species is P. brasiliensis.
Brasiliguana is a genus of iguanian lizard which lived during the late Cretaceous period in what is now Brazil. It is known from the holotype MN 7230-V, an isolated left maxilla with partially preserved teeth, which was found in the Upper Cretaceous Adamantina Formation, part of the Bauru Group of São Paulo State, southeast Brazil. Brasiliguana was named by William R. Nava and Agustín G. Martinelli in 2011 and the type species is Brasiliguana prudentis. The generic name refers to its provenance from the Late Cretaceous of Brazil and iguana, from the South American and Caribbean aboriginal language meaning "lizard". The specific name, prudentis, refers to Presidente Prudente Municipality, where the holotype was found.
Ovoo gurvel is an extinct monitor lizard from the Late Cretaceous of Mongolia. It is one of the smallest and earliest monitor lizards. It was described in 2008. Ovoo possesses a pair of two small bones in its skull that are not seen in any other lizard.
Polyglyphanodontia is an extinct clade of lizards from the Cretaceous that includes around a dozen genera. Polyglyphanodontians were the dominant group of lizards in North America and Asia during the Late Cretaceous. Most polyglyphanodontians are Late Cretaceous in age, though the oldest one, Kuwajimalla kagaensis, is known from the Early Cretaceous Kuwajima Formation (Japan). Early Cretaceous South American taxon Tijubina, and possibly also Olindalacerta, might also fall within Polyglyphanodontia or be closely allied to the group, but if so, they would be two of only three Gondwanan examples of an otherwise Laurasian clade. They produced a remarkable range of forms. Chamopsiids, including Chamops, were characterized by large, blunt, crushing teeth, and were most likely omnivores. Macrocephalosaurus, from the Gobi Desert, was a specialized herbivore; it grew to roughly a meter long and had multicusped, leaf-shaped teeth like those of modern iguanas. Polyglyphanodon, from the Maastrichtian of Utah, was another herbivore, but its teeth formed a series of transverse blades, similar to those of Trilophosaurus. Peneteius had remarkable, multicusped teeth, similar to those of mammals. The polyglyphanodontids first appear in the latter part of the Early Cretaceous in North America, and became extinct during the Cretaceous-Paleogene extinction event. Polyglyphanodontians closely resembled the teiid lizards, and purported teiid lizards from the Late Cretaceous appear to be polyglyphanodontians.
The Lacertoidea is a group of lizards that includes the Lacertidae, Teiidae, Gymnophthalmidae, and the burrowing Amphisbaenia.
Pariguana is an extinct genus of iguanid lizard from the Late Cretaceous of western North America. It is known from a single type species, Pariguana lancensis, named in 2012 on the basis of a partial lower jaw from the Lance Formation in eastern Wyoming. This jaw bone comes from a layer dated approximately 650,000 years before the Cretaceous–Paleogene extinction event. Pariguana is the oldest definitive iguanid from North America, and may represent the first stage of the iguanian evolutionary radiation from Asia into North America.
Gueragama is an extinct genus of iguanian lizard from the Late Cretaceous of Brazil. It belongs to a group of iguanians called Acrodonta, whose living members include chameleons and agamids and are currently restricted to the Old World. Gueragama is the only acrodont known from South America, providing evidence that the group once ranged across much of Gondwana and only became restricted to the Old World after the supercontinent broke apart. The type species, Gueragama sulamericana, was named in 2015 on the basis of an isolated lower jaw from the Turonian- to Campanian-age Goio-Erê Formation in the Bauru Basin, which was deposited in a desert environment. Unlike modern acrodontans, whose teeth implant on the margins of the jaws, Gueragama has teeth that implant along the inner surface of the lower jaw, a feature common in most non-acrodontan lizards and characteristic of the second major group of iguanians, Pleurodonta. The non-acrodont dentition of Gueragama is evidence of its basal position within Acrodonta, and is shared with the taxon of Late Cretaceous acrodontan relatives, the family Priscagamidae.
Sachicasaurus is an extinct genus of brachauchenine pliosaurid known from the Barremian of the Paja Formation, Altiplano Cundiboyacense in the Colombian Eastern Ranges of the Andes. The type species is S. vitae.
