The Messel pit is a disused quarry near the village of Messel about 35 km (22 mi) southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of well-preserved fossils dating from the middle of the Eocene, it has significant geological and scientific importance. Over 1000 species of plants and animals have been found at the site. After almost becoming a landfill, strong local resistance eventually stopped these plans and the Messel pit was declared a UNESCO World Heritage Site on 9 December 1995. Significant scientific discoveries about the early evolution of mammals and birds are still being made at the Messel pit, and the site has increasingly become a tourist site as well.
Viverravidae is an extinct monophyletic family of mammals from extinct superfamily Viverravoidea within the clade Carnivoramorpha, that lived from the early Palaeocene to the late Eocene in North America, Europe and Asia. They were once thought to be the earliest carnivorans and ancestral to extant ones, but now are placed outside the order Carnivora based on cranial morphology as relatives to extant carnivorans.
Miacis is an extinct genus of placental mammals from clade Carnivoraformes, that lived in North America from early to middle Eocene.
Didymictis is an extinct genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America and Europe from the late Paleocene to middle Eocene.
Palaeopython is an extinct genus of snake from the Eocene of Europe. The genus has been used to refer to large Western and Central European snake vertebrae from the Eocene. P. cadurcensis and the tentatively-referred "P." neglectus originate from France; P. ceciliensis originates from Germany; and P. helveticus originates from Switzerland. A species known from multiple well-preserved specimens found in the Messel Pit of Germany, P. fischeri, was named by Stephan Schaal in 2004, but examination of the genus showed that it represented a distinct lineage; it was renamed Eoconstrictor fischeri in 2020 by Agustín Scanferla and Krister T. Smith. Another species from France, P. filholii, was moved to the genus Phosphoroboa in 2021 by Georgalis, Márton Rabi, and Smith. An additional species, P. sardus, was described in 1901 by Alessandro Portis from the Middle Miocene of Monte Albu. However, a reevaluation of the holotype specimen of this species revealed it to actually belong to an indeterminate acanthomorph fish.
Saniwa is an extinct genus of varanid lizard that lived during the Eocene epoch. It is known from well-preserved fossils found in the Bridger and Green River Formations of Wyoming, United States. The type species S. ensidens was described in 1870 as the first fossil lizard known from North America. A second species, S.orsmaelensis, is recognised from remains found in Europe. It is a close relative of Varanus, the genus that includes monitor lizards.
Afairiguana avius is an extinct iguanid lizard known from a nearly complete and articulated skeleton discovered in rocks of the Early Eocene-aged Green River Formation of Wyoming, United States. As of the initial description, the skeleton represents the oldest complete iguanian from the Western Hemisphere, and is the oldest representative of the extant iguanid family of anoles, Polychrotidae.
The Willwood Formation is a sedimentary sequence deposited during the late Paleocene to early Eocene, or Clarkforkian, Wasatchian and Bridgerian in the NALMA classification.
Barbaturex is an extinct genus of giant herbivorous iguanian lizards from Pondaung Formation, the Eocene of Myanmar. It is represented by a single species, Barbaturex morrisoni, which is known from several partial dentaries and a fused pair of frontals, two bones that form part of the top of the skull. Based on the size of these bones, Barbaturex morrisoni is estimated to have been about 1 metre (3.3 ft) from snout to vent, and possibly up to 6 feet (1.8 m) including the tail. Barbaturex morrisoni was named after The Doors frontman Jim Morrison, a play on his epithet "The Lizard King". The genus's name is a portmanteau of the Latin words Barbatus and rex, meaning "bearded king", in reference to ridges along the mandible and the lizard's large size.
Shinisauria is a clade or evolutionary grouping of anguimorph lizards that includes the living Chinese crocodile lizard Shinisaurus and several of its closest extinct relatives. Shinisauria was named in 2008 by Jack Lee Conrad as a stem-based taxon to include all anguimorphs more closely related to Shinisaurus than to Anguis fragilis, Heloderma suspectum or Varanus varius. Several recent phylogenetic analyses of lizard evolutionary relationships place Shinisauria in a basal position within the clade Platynota, which also includes monitor lizards, helodermatids, and the extinct mosasaurs. Shinisaurians were once thought to be closely related to the genus Xenosaurus, but they are now considered distant relatives within Anguimorpha. The fossil record of shinisaurians extends back to the Early Cretaceous with Dalinghosaurus, which is from the Aptian aged Yixian Formation of China. Two other extinct shinisaurians are currently known: Bahndwivici from the Eocene of Wyoming and Merkurosaurus from the Late Oligocene of Germany and the Early Miocene of the Czech Republic. An indeterminate shinisaurian is known from an isolated tail found in the Eocene aged Messel pit in Germany.
Anniealexandria is an extinct genus of amphisbaenian lizard known by the type species Anniealexandria gansi from the earliest Eocene of Wyoming. Anniealexandria is the only known member of the family Bipedidae in the fossil record, which otherwise only includes the extant genus Bipes from Mexico. It was named in 2009 in honor of Annie Montague Alexander, founder of the University of California Museum of Paleontology. Remains of Anniealexandria are known only from a single fossil locality in the Bighorn Basin called Castle Gardens, but within the locality its fossils are common in the Willwood Formation, usually consisting of isolated jaw bones and vertebrae. Anniealexandria seems to have been a common component of a paleofauna that included fifteen other lizard species and existed in western North America during a period of global warming in the latest Paleocene and earliest Eocene.
Anolbanolis is an extinct genus of iguanian lizards that lived in the Bighorn Basin of what is now Wyoming during the earliest Eocene. The type species A. banalis was named by paleontologist Krister Smith in 2009 from a collection of isolated skull fragments found in a layer of the Willwood Formation that dates to a brief period of global warming called the Paleocene-Eocene thermal maximum (PETM) about 56 million years ago. The genus name Anolbanolis means "poor anole" from the Greek anolbos and the name Anolis, in reference to the scrappy nature of known fossil material and its close resemblance to lizards in the genus Anolis. Smith suggested that Anolbanolis may be a close relative of Anolis or Polychrus, which are common today in Central and South America but not found as far north as Wyoming, fitting with the idea that the Bighorn Basin was warmer and wetter in the Eocene than it is currently. According to Smith, the species name banalis is a reference to it being a "banal" lizard in the Willwood, being "quite abundant in the type locality," and not unusual because "the discovery of Eocene boreal fossil members of living subtropical and tropical clades is becoming commonplace." In 2011 Smith named a second species of Anolbanolis, A. geminus, which lived in the Bighorn Basin shortly after the PETM and was about 50 percent larger than A. banalis. Smith concluded on the basis of features in A. geminus that Anolbanolis is more closely related to Anolis than it is to Polychrus.
Babibasiliscus is an extinct genus of casquehead lizard that lived in what is now Wyoming during the early Eocene, approximately 48 million years ago. The genus is known from a single species, Babibasiliscus alxi, which was named by paleontologist Jack Conrad in 2015 on the basis of a fossilized skull from the Bridger Formation in the Green River Basin. The name Babibasiliscus comes from the Shoshoni word babi, meaning "older male cousin", and Basiliscus, a modern-day genus of casquehead lizards. The specimen is undeformed and nearly complete except for the tip of the snout and the top of the skull, making it unclear whether the distinctive bony crest of living corytophanids was present in prehistoric relatives like Babibasiliscus. The skull is about 42 millimetres (2 in) in length and the entire body is estimated to have been about 0.6 metres (2 ft) long. Bones on the right side of lower jaw of the specimen are thickened and fused together, suggesting that the jaw had broken and healed when the animal was alive.
Geiseltaliellus is an extinct genus of iguanian lizards that lived in what is now western Europe during the Eocene. It belongs to the family Corytophanidae, which includes modern casquehead lizards. Many fossils are known from Germany, France, and Belgium, with the most well preserved coming from the Messel pit lagerstätte in Messel, Germany. German paleontologist Oskar Kuhn named the genus in 1944 after the Geiseltal valley where the first specimens were found, designating the type species Geiseltaliellus longicaudus. Three new species — G. louisi, G. lamandini, and G. grisolli — were named in the 1990s and 2000s on the basis of more fragmentary remains from France and Belgium, although G. louisi has since been synonymized with G. longicaudus. In 2009 the Messel pit specimens were recognized as belonging to a species distinct from that of the G. longicaudus specimens in Geiseltalt and were collectively reclassified under a new name, G. maarius.
Magnuviator is a genus of extinct iguanomorph lizard from the Late Cretaceous of Montana, US. It contains one species, M. ovimonsensis, described in 2017 by DeMar et al. from two specimens that were discovered in the Egg Mountain nesting site. Magnuviator is closest related to the Asian Saichangurvel and Temujinia, which form the group Temujiniidae. Unlike other members of the Iguanomorpha, however, Magnuviator bears a distinct articulating notch on its tibia for the ankle bones, which has traditionally been considered a characteristic of non-iguanomorph lizards. The morphology of its teeth suggests that its diet would have mainly consisted of wasps, like the modern phyrnosomatid iguanians Callisaurus and Urosaurus, although it also shows some adaptations to herbivory.
Palaeovaranidae, formerly known as Necrosauridae, is an extinct clade of anguimorph lizards known from the Paleogene of Europe. They have sometimes been recovered as members of Varanoidea. It contains three genera.
Eolacertidae is an extinct family of lacertoid lizards known from the Paleocene and Eocene of Europe. They are the closest known relatives of Lacertidae.
Paranecrosaurus is an extinct genus of lizard from the Eocene Messel Pits of Germany. It contains a single species, Paranecrosaurus feisti, originally described as a species of Saniwa. It was carnivorous, as indicated by the presence of the lizard Cryptolacerta as stomatch contents. It is placed in the family Palaeovaranidae.
This is an overview of the paleofauna of the Eocene Messel Formation as explored by the Messel Pit excavations in Germany. A former quarry and now UNESCO World Heritage Site, the Messel Formation preserves what once were a series of anoxic lakes surrounded by a sub-tropical rainforest during the Middle Eocene, approximately 47 Ma.
This list of fossil reptiles described in 2016 is a list of new taxa of fossil reptiles that were described during the year 2016, as well as other significant discoveries and events related to reptile paleontology that occurred in 2016.
