| Palaeosaniwa | |
|---|---|
 | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Order: | Squamata |
| Suborder: | Anguimorpha |
| Genus: | † Palaeosaniwa |
| Species: | †P. canadensis |
| Binomial name | |
| †Palaeosaniwa canadensis Gilmore, 1928 | |
| Synonyms [1] | |
| |
Palaeosaniwa canadensis is an extinct species of carnivorous lizard from the late Cretaceous of North America. The name, given by Charles Whitney Gilmore in 1928, means "ancient Saniwa from Canada".
Palaeosaniwa is among the largest terrestrial lizards known from the Mesozoic era, with an estimated body mass of 6 kg (13 lb) and a snout–vent length about 82 cm (32 in). [3] Its total body length would have been around 1–2 m (3 ft 3 in – 6 ft 7 in), smaller than the adult Komodo dragon and other Mesozoic lizards of East Asia ( Asprosaurus and Chianghsia ). [4] [5] [6] Some specimens attributed to cf. P. canadensis is estimated to have skull length up to 15 centimetres (5.9 in) which is comparable to a young specimen of komodo dragon around 1.6 m (5.2 ft) long. [7] It is similar to modern varanid lizards (particularly the Komodo dragon) in having bladelike teeth with minute serrations. These teeth would have been effective for seizing and cutting large prey items, and suggest that Palaeosaniwa fed on other vertebrates. Adult Palaeosaniwa would have been large enough to prey on any of the avialans or mammals known from the time, small non-avian dinosaurs, and the eggs and juveniles of large dinosaurs.
Palaeosaniwa was originally described from the late Campanian of Alberta. More recently it has been reported from the late Campanian of Montana, and the late Maastrichtian of Montana and Wyoming. It is known primarily from isolated teeth and vertebrae, but two partial skeletons have also been discovered. The type species, P. canadensis, is from Alberta. Although the Maastrichtian Palaeosaniwa has traditionally been referred to this species, it succeeds it by roughly ten million years. Given the distance in time between these animals, they are likely to represent distinct species, but the available fossils are too incomplete to be certain.
Palaeosaniwa is a member of the Platynota, a group that includes the monitor lizards (Varanidae) and Gila monsters (Helodermatidae). Originally, it was thought to be a member of the Varanidae, but has also been interpreted as a relative of the Helodermatidae. [8] The most recent analysis places Palaeosaniwa outside of either Varanidae or Helodermatidae, as a stem member of the Varanoidea. Its precise affinities remain poorly understood, but it may be related to other Late Cretaceous, North American carnivorous lizards such as Parasaniwa , Paraderma , Labrodioctes , and Cemeterius .
The Varanidae are a family of lizards in the superfamily Varanoidea and order Anguimorpha. The family, a group of carnivorous and frugivorous lizards, includes the living genus Varanus and a number of extinct genera more closely related to Varanus than to the earless monitor lizard (Lanthanotus). Varanus includes the Komodo dragon, crocodile monitor, savannah monitor, the goannas of Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the earless monitor lizard and Chinese crocodile lizard. The oldest members of the family are known from the Late Cretaceous of Mongolia.
Mosasaurs are an extinct group of large aquatic reptiles within the family Mosasauridae that lived during the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes.
Saurornitholestes is a genus of carnivorous dromaeosaurid theropod dinosaur from the late Cretaceous of Canada (Alberta) and the United States.
Chirostenotes is a genus of oviraptorosaurian dinosaur from the late Cretaceous of Alberta, Canada. The type species is Chirostenotes pergracilis.
The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller.
Globidens is an extinct genus of mosasaurid oceanic lizard classified as part of the Globidensini tribe in the Mosasaurinae subfamily. Globidens belongs to the family Mosasauridae, which consists of several genera of predatory marine lizards of various sizes that were prevalent during the Late Cretaceous. Specimens of Globidens have been discovered in Angola, Brazil, Colombia, Morocco, Syria and the United States. Among mosasaurs, Globidens is probably most well known for the highly rounded, globe-like teeth that give it its name.
Halisaurus is an extinct genus of mosasaur named by Othniel Charles Marsh in 1869. The holotype, consisting of an angular and a basicranium fragment discovered near Hornerstown, New Jersey, already revealed a relatively unique combination of features and prompted a new genus to be described. Its name is a portmanteau of the Ancient Greek ἅλς and σαῦρος. It was renamed by Marsh to Baptosaurus in 1870, since he believed the name to already be preoccupied by the fish Halosaurus. According to modern rules, a difference of a letter is enough and the substitute name is unneeded, making "Baptosaurus" a junior synonym.
Varanoidea is a superfamily of lizards, including the well-known family Varanidae. Also included in the Varanoidea are the Lanthanotidae, and the extinct Palaeovaranidae.
Centrosaurinae is a subfamily of ceratopsid, a group of large quadrupedal ornithischian dinosaur. Centrosaurine fossil remains are known primarily from the northern region of Laramidia but isolated taxa have been found in China and Utah as well.
Pluridens is an extinct genus of marine lizard belonging to the Mosasauridae. Pluridens is placed in the subfamily Halisaurinae with the genera Phosphorosaurus, Eonatator and Halisaurus. Compared to related halisaurines, Pluridens had longer jaws with more teeth, and smaller eyes. It also grew large size, measuring 5–6 m (16–20 ft) long and perhaps over 9 m (30 ft) in some individuals. The jaws in some specimens are robust, and sometimes show injuries suggestive of combat. The jaws may have been used for fighting over mates or territories.
The Anguimorpha is a suborder of squamates. The group was named by Fürbringer in 1900 to include all autarchoglossans closer to Varanus and Anguis than Scincus. These lizards, along with iguanians and snakes, constitute the proposed "venom clade" Toxicofera of all venomous reptiles.
Taniwhasaurus is an extinct genus of mosasaurs that lived during the Campanian stage of the Late Cretaceous. It is a member of the subfamily Tylosaurinae, a lineage of mosasaurs characterized by a long toothless conical rostrum. Two valid species are attached to the genus, T. oweni and T. antarcticus, known respectively from the fossil record of present-day New Zealand and Antarctica. Two other species have been nominally classified within the genus, T. 'capensis' and T. 'mikasaensis', recorded in present-day South Africa and Japan, but their attribution remains problematic due to the fragmentary state of their fossils. The generic name literally means "taniwha lizard", referring to a supernatural aquatic creature from Māori mythology.
Socognathus is a genus of prehistoric chamopsiid polyglyphanodontian lizards containing species that lived from the Middle Campanian stage to the late Maastrichtian. Several specimens of the type species, Socognathus unicuspis, have been found in Alberta, Canada. A second species, Socognathus brachyodon is known from the late Maastrichtian Lance Formation; its fossils have been found in Wyoming, United States.
Asprosaurus is an extinct genus of anguimorph lizard from the Late Cretaceous of South Korea. Named in 2015 from the Seonso Conglomerate Formation, the type species Asprosaurus bibongriensis is the first Mesozoic lizard to have been discovered on the Korean peninsula. Because Asprosaurus is known only from fragmentary material, its relationships with other lizards are uncertain. However, features of the lower jaw suggest that it may be a member of a clade called Monstersauria, which includes the living Gila monster.
Kaikaifilu is an extinct genus of large mosasaurs that lived during the Maastrichtian stage of the Late Cretaceous, in what is now northern Antarctica. The only species known, K. hervei, was described in 2017 from an incomplete specimen discovered in the López de Bertodano Formation, in Seymour Island, Antarctic Peninsula. The taxon is named in reference to Coi Coi-Vilu, a reptilian ocean deity of the Mapuche cosmology. Early observations of the holotype classify it as a member of the subfamily Tylosaurinae. However, later observations note that several characteristics show that this attribution is problematic.
Leptorhynchos is an extinct genus of caenagnathid theropod from the Late Cretaceous of what is now the US state of Texas, although it has been suggested to also exist in Alberta and South Dakota. The type species is L. gaddisi, and it is currently the only widely accepted valid species. The generic name of Leptorhynchos comes from the Greek "leptos" meaning "small" and "rhynchos" meaning "beak". The specific epithet is in honor of the Gaddis family, who owned the land on which the holotype was discovered.
Xenodens is a potentially dubious extinct genus of marine lizard belonging to the mosasaur family. It contains a single species, X. calminechari, which is known from Late Maastrichtian phosphate deposits in the Ouled Abdoun Basin, Morocco.
