The Oligocene is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present. As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich; the name comes from the Ancient Greek ὀλίγος and καινός, and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period.
Barn-owls are one of the two families of owls, the other being the true owls or typical owls, Strigidae. They are medium to large owls with large heads and characteristic heart-shaped faces. They have long, strong legs with powerful talons. They also differ from the Strigidae in structural details relating in particular to the sternum and feet.
The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents. These birds are adapted for swimming, floating on the water surface, and in some cases diving in at least shallow water. The family contains around 174 species in 43 genera.
The rails, or Rallidae, are a large cosmopolitan family of small- to medium-sized, ground-living birds. The family exhibits considerable diversity and includes the crakes, coots, and gallinules. Many species are associated with wetlands, although the family is found in every terrestrial habitat except dry deserts, polar regions, and alpine areas above the snow line. Members of the Rallidae occur on every continent except Antarctica. Numerous island species are known. The most common rail habitats are marshland and dense forest. They are especially fond of dense vegetation.
Sparassodonta is an extinct order of carnivorous metatherian mammals native to South America. They were once considered to be true marsupials, but are now thought to be either a sister taxon to them, or considerably distantly related, part of a separate clade of Gondwanan metatherians. A number of these mammalian predators closely resemble placental predators that evolved separately on other continents, and are cited frequently as examples of convergent evolution. They were first described by Florentino Ameghino, from fossils found in the Santa Cruz beds of Patagonia. Sparassodonts were present throughout South America's long period of "splendid isolation" during the Cenozoic; during this time, they shared the niches for large warm-blooded predators with the flightless terror birds. Previously, it was thought that these mammals died out in the face of competition from "more competitive" placental carnivorans during the Pliocene Great American Interchange, but more recent research has showed that sparassodonts died out long before eutherian carnivores arrived in South America. Sparassodonts have been referred to as borhyaenoids by some authors, but currently the term Borhyaenoidea refers to a restricted subgroup of sparassodonts comprising borhyaenids and their close relatives.
Hyaenodon ("hyena-tooth") is the type genus of the Hyaenodontidae, a family of extinct carnivorous fossil mammals from Eurasia, North America, and Africa, with species existing temporally from the Eocene until the middle Miocene, existing for about 26.1 million years .
Rhineuridae is a family of amphisbaenians that includes one living genus and species, Rhineura floridana, as well as many extinct species belonging to both Rhineura and several extinct genera. The living R. floridana is found only in Florida no further north than the panhandle, but extinct species ranged across North America, some occurring as far west as Oregon. The family has a fossil record stretching back 60 million years to the Paleocene and was most diverse in the continental interior during the Eocene and Oligocene.
Herpetotheriidae is an extinct family of metatherians. Although sometimes placed in the living opossum family, they may form the sister clade to all living marsupials. Species of this family are generally reconstructed as terrestrial. Fossils of herpetotheriids come from North America, Asia, Europe, Africa, and perhaps South America. The oldest representative is Maastrichtidelphys from the latest Cretaceous (Maastrichtian) of the Netherlands and the youngest member is Amphiperatherium from the Middle Miocene of Europe.
Archaeohippus is an extinct three toed member of the family Equidae known from fossils of early Oligocene to middle Miocene age. The genus is noted for several distinct skeletal features. The skull possesses deeply pocketed fossa in a notably long preorbital region. The genus is considered an example of phyletic dwarfism with adults estimated at being on average 20kg in weight. This is in contrast to the most common equid of the period, Miohippus. Characters of the teeth show a mix of both primitive and advanced traits. The advanced traits are very similar to those shown in the genus Parahippus. The noted similarities of Archaeohippus and Parahippus show them to be descended from a common ancestor and are considered sister species.
Brachyhyops is an extinct genus of entelodont artiodactyl mammal that lived during the Eocene Epoch of western North America and southeastern Asia. The first fossil remains of Brachyhyops are recorded from the late Eocene deposits of Beaver Divide in central Wyoming and discovered by paleontology crews from the Carnegie Museum of Natural History during the early 20th century. The type species, Brachyhyops wyomingensis, is based on a single skull and was named by E.H. Colbert in 1937, but was not officially described until 1938. During the latter half of the 20th century, additional specimens from North America have been recorded from Saskatchewan and as far south as Texas, indicating that Brachyhyops had a broad distribution and was well-dispersed throughout western North America.
Boavus is an extinct genus of boa known primarily from Eocene-aged strata of North America. At least three species are known from the Middle Eocene Green River lagerstätte in Wyoming, two species from Eocene strata of Uinta County, and at least one species is known from the middle to late Eocene-aged Sespe Formation of California.
The Macropodidae are an extant family of marsupial with the distinction of the ability to move bipedally on the hind legs, sometimes by jumping, as well as quadrupedally. They are herbivores, but some fossil genera like Ekaltadeta are hypothesised to have been carnivores. The taxonomic affiliations within the family and with other groups of marsupials is still in flux.
Tomistominae is a subfamily of crocodylians that includes one living species, the false gharial. Many more extinct species are known, extending the range of the subfamily back to the Eocene epoch. In contrast to the false gharial, which is a freshwater species that lives only in southeast Asia, extinct tomistomines had a global distribution and lived in estuaries and along coastlines.
The Hoko River Formation is a Late Eocene marine sedimentary geologic formation. The formation is exposed in outcrops along the Strait of Juan de Fuca on the Olympic Peninsula in Washington state, USA. It is known for containing numerous fossils of crabs. It overlies the older Lyre Formation and underlies the younger Makah Formation.
Dyticonastis is an extinct genus of amphisbaenians, or worm lizards, that includes a single species, Dyticonastis rensbergeri, that lived during the late Oligocene and early Miocene in what is now Oregon. Fossils of the species come from the John Day Formation. It belongs to Rhineuridae, a family that includes many other extinct North American amphisbaenians but only one living species, Rhineura floridana, from Florida. Dyticonastis rensbergeri occurs the farthest west of all rhineurid species. Like all rhineurids, Dyticonastis has a shovel-like snout adapted for burrowing underground, but it differs from most other members of the group in having a relatively shallow angle to its snout wedge and in having a widened snout tip. The only other rhineurids that share these features are species of the genus Spathorhynchus, which lived from the Middle Eocene to the Early Oligocene in what is now Wyoming. A 2007, phylogenetic analysis of amphisbaenians found that Dyticonastis and Spathorhynchus are each other's closest relatives, suggesting that both taxa may have evolved through vicariant speciation; the growth of the Rocky Mountains during the earliest stages of the Laramide orogeny in the early Paleogene would have separated North American rhineurids into eastern and western populations, with the western population producing Dyticonastis and Spathorhynchus.
Ototriton is an extinct genus of rhineurid amphisbaenian or worm lizard from the Early Eocene of the western United States, including the type and only species Ototriton solidus. Paleontologist F. B. Loomis named Ototriton in 1919 on the basis of a single skull from the Wind River Formation in Wyoming, misinterpreting it as the skull of a salamander. Unlike salamanders and like other rhineurids, Ototriton has a shovel-shaped snout that it presumably used for burrowing underground. Ototriton is one of the earliest known rhineurids and also one of the largest.
Macrorhineura is an extinct genus of rhineurid amphisbaenian or worm lizard, including the type and only species Macrorhineura skinneri, named in 1970 on the basis of the front half of a skull from the Early Miocene Sharps Formation in Wounded Knee, South Dakota. Although the skull is incomplete, features such as a pointed, shovel-shaped snout indicate that it belongs to the family Rhineuridae. Within Rhineuridae, Macrorhineura is most closely related to Ototriton and Hyporhina, two genera from the Eocene and Oligocene of Colorado and Wyoming, based on the shared feature of equally sized dentary teeth in the lower jaw. Together they form a clade or evolutionary grouping of mid-continental rhineurids, which became isolated from a more western clade of rhineurids that includes Dyticonastis and Spathorhynchus. Rhineurids were relatively common across much of North America during the Paleogene, but their range contracted in the Neogene as the climate became colder, leaving only one living species in Florida, Rhineura floridana. The presence of Macrorhineura in the Miocene shows that mid-continental rhineurids persisted into the Neogene, although by this time their distribution range was already shrinking.
Hyporhina is an extinct genus of amphisbaenians or worm lizards that lived from the Late Eocene to the Middle Oligocene in what is now the western United States.
Archaerhineura was a genus of amphisbaenian lizards in the family Rhineuridae that is now extinct. The only species is Archaerhineura mephitis, named in 2015 on the basis of a single fragment of the lower jaw from the Polecat Bench Formation in Park County, Wyoming, which dates to the late Paleocene. Archaerhineura is one of the oldest amphisbaenians and was part of an evolutionary radiation of Rhineuridae in the Paleocene several million years after the Cretaceous–Paleogene extinction event. This rhineurid radiation coincided with the radiation of another group of amphisbaenians, Amphisbaeniformes, which includes the still-extant families Blanidae and Amphisbaenidae. The presence of Archaerhineura and other Paleocene rhineurids in the western United States indicates that amphisbaenians, which would later have a nearly global distribution, originated in North America.
