Hyporhina

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Hyporhina
Temporal range: Late EoceneMiddle Oligocene, 40–30  Ma
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Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Family: Rhineuridae
Genus:Hyporhina
Baur, 1893
Species
  • H. antiqua Baur, 1893 (type)
  • H. galbreathiTaylor, 1951

Hyporhina is an extinct genus of amphisbaenians or worm lizards that lived from the Late Eocene to the Middle Oligocene (approximately 40 to 30 million years ago) in what is now the western United States.

A genus is a taxonomic rank used in the biological classification of living and fossil organisms, as well as viruses, in biology. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus.

Amphisbaenia suborder of reptiles

Amphisbaenia is a group of usually legless squamates, comprising over 180 extant species. Amphisbaenians are characterized by their long bodies, the reduction or loss of the limbs, and rudimentary eyes. As many species have a pink body and scales arranged in rings, they have a superficial resemblance to earthworms. While the genus Bipes retains forelimbs, all other genera are limbless. Although superficially similar to the snakes and Dibamidae, recent phylogenetic studies suggest that they are most closely related to the Lacertidae. Amphisbaenians are widely distributed, occurring in North America, Europe, Africa, South America, and the Caribbean. Most species are less than 6 inches (150 mm) long.

The Oligocene is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present. As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich; the name comes from the Ancient Greek ὀλίγος and καινός, and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period.

Contents

Species

It currently includes two species, both from the White River Formation in eastern Wyoming and northeastern Colorado:

The White River Formation is a geologic formation of the Paleogene Period, in the northern Great Plains and central Rocky Mountains, within the United States.

Type species term used in zoological nomenclature (also non-officially in botanical nomenclature)

In zoological nomenclature, a type species is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen(s). A similar concept is used for suprageneric groups called a type genus.

In scientific nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name, although the term is used somewhat differently in the zoological code of nomenclature. For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies. This name is no longer in use: it is now a synonym of the current scientific name, Picea abies.

Taxonomy

Paleontologist Georg Baur named the genus Hyporhina in 1893, making it one of the first prehistoric amphisbaenians to be described. Baur placed it in its own family, Hyporhinidae, because it possessed eye sockets that are enclosed at the back by postorbital bars, a feature that living amphisbaenians lack.

Georg Baur German paleontologist and herpetologist (1859–1898)

Georg Baur (1859–1898) was a vertebrate paleontologist and Neo-Lamarckian who studied reptiles of the Galapagos Islands, particularly the Galápagos tortoises, in the 1890s. He is perhaps best known for his subsidence theory of the origin of the Galapagos Islands, where he postulated the islands were the remains of a former landmass, connected to South America via Cocos Island.

The postorbital bar is a bony arched structure that connects the frontal bone of the skull to the zygomatic arch, which runs laterally around the eye socket. It is a trait that only occurs in mammalian taxa, such as most strepsirrhine primates and the hyrax, while haplorhine primates have evolved fully enclosed sockets. One theory for this evolutionary difference is the relative importance of vision to both orders. As haplorrhines tend to be diurnal, and rely heavily on visual input, many strepsirrhines are nocturnal and have a decreased reliance on visual input.

However, more recent studies have placed it within Rhineuridae, a family that includes the living Rhineura floridana from Florida, and many more extinct species from the western United States that were named after 1893 and have postorbital bars. Hyporhina differs from other rhineurids in having a steep angle to its snout. [1]

Rhineuridae family of reptiles

Rhineuridae is a family of amphisbaenians that includes one living genus and species, Rhineura floridana, as well as many extinct species belonging to both Rhineura and several extinct genera. The living R. floridana is found only in Florida no further north than the panhandle, but extinct species ranged across North America, some occurring as far west as Oregon. The family has a fossil record stretching back 60 million years to the Paleocene and was most diverse in the continental interior during the Eocene and Oligocene.

<i>Rhineura floridana</i> species of reptile

Rhineura floridana, known as the North American worm lizard, Florida worm lizard, or thunderworm, is the only member of the monotypic genus Rhineura. This species is found primarily in Florida but has been recorded in Lanier County, Georgia. No subspecies are currently recognized.

See also

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Archaerhineura was a genus of amphisbaenian lizards in the family Rhineuridae that is now extinct. The only species is Archaerhineura mephitis, named in 2015 on the basis of a single fragment of the lower jaw from the Polecat Bench Formation in Park County, Wyoming, which dates to the late Paleocene. Archaerhineura is one of the oldest amphisbaenians and was part of an evolutionary radiation of Rhineuridae in the Paleocene several million years after the Cretaceous–Paleogene extinction event. This rhineurid radiation coincided with the radiation of another group of amphisbaenians, Amphisbaeniformes, which includes the still-extant families Blanidae and Amphisbaenidae. The presence of Archaerhineura and other Paleocene rhineurids in the western United States indicates that amphisbaenians, which would later have a nearly global distribution, originated in North America.

Anniealexandria is an extinct genus of amphisbaenian lizard known by the type species Anniealexandria gansi from the earliest Eocene of Wyoming. Anniealexandria is the only known member of the family Bipedidae in the fossil record, which otherwise only includes the extant genus Bipes from Mexico. It was named in 2009 in honor of Annie Montague Alexander, founder of the University of California Museum of Paleontology. Remains of Anniealexandria are known only from a single fossil locality in the Bighorn Basin called Castle Gardens, but within the locality its fossils are common in the Willwood Formation, usually consisting of isolated jaw bones and vertebrae. Anniealexandria seems to have been a common component of a paleofauna that included fifteen other lizard species and existed in western North America during a period of global warming in the latest Paleocene and earliest Eocene.

Babibasiliscus is an extinct genus of casquehead lizard that lived in what is now Wyoming during the early Eocene, approximately 48 million years ago. The genus is known from a single species, Babibasiliscus alxi, which was named by paleontologist Jack Conrad in 2015 on the basis of a fossilized skull from the Bridger Formation in the Green River Basin. The name Babibasiliscus comes from the Shoshoni word babi, meaning "older male cousin", and Basiliscus, a modern-day genus of casquehead lizards. The specimen is undeformed and nearly complete except for the tip of the snout and the top of the skull, making it unclear whether the distinctive bony crest of living corytophanids was present in prehistoric relatives like Babibasiliscus. The skull is about 42 millimetres (2 in) in length and the entire body is estimated to have been about 0.6 metres (2 ft) long. Bones on the right side of lower jaw of the specimen are thickened and fused together, suggesting that the jaw had broken and healed when the animal was alive.

References

  1. Hembree, D.I. (2007). "Phylogenetic revision of Rhineuridae (Reptilia: Squamata: Amphisbaenia) from the Eocene to Miocene of North America". The University of Kansas Paleontological Contributions. 15: 1–20.