The Lepidosauria are reptiles with overlapping scales. This subclass includes Squamata and Rhynchocephalia. It is a monophyletic group and therefore contains all descendants of a common ancestor. Squamata includes snakes, lizards, and amphisbaenia. Rhynchocephalia was a widespread and diverse group 220-100 million years ago; however, it is now represented only by the genus Sphenodon, which contains a single species of tuatara, native to New Zealand. Lepidosauria is the sister taxon to Archosauria, which includes Aves and Crocodilia. Lizards and snakes are the most speciose group of lepidosaurs and, combined, contain over 9,000 species. There are many noticeable distinguishing morphological differences between lizards, tuataras, and snakes.
Agamidae is a family of over 300 species of iguanian lizards indigenous to Africa, Asia, Australia, and a few in Southern Europe. Many species are commonly called dragons or dragon lizards.
Iguania is an infraorder of squamate reptiles that includes iguanas, chameleons, agamids, and New World lizards like anoles and phrynosomatids. Consisting of nearly 13,000 named species. T Using morphological features as a guide to evolutionary relationships, the Iguania are believed to form the sister group to the remainder of the Squamata. However, molecular information has placed Iguania well within the Squamata as sister taxa to the Anguimorpha and closely related to snakes.. The order has been under debate and revisions after being classified by Charles Lewis Camp in 1923 due to difficulties finding adequate synapomorphic morphological characteristics. The majority of Iguanias are arboreal but there are several terrestrial groups. They usually have primitive fleshy, non-prehensile tongues, although the tongue is highly modified in chameleons. The group has a fossil record that extends back to the Early Jurassic .. Today they are scattered occurring in Madagascar, the Fiji and Friendly Islands and Western Hemisphere
Lazarussuchus is a genus of basal choristodere, a type of amphibious reptile, known from France, Germany, and the Czech Republic. Fossils have been found in Late Paleocene, Late Oligocene, and Early Miocene deposits, meaning that the genus lasted about 40 million years. Two species have been named: the type species L. inexpectatus ("unexpected") from the late Oligocene of France and Germany and L. dvoraki from the early Miocene of the Czech Republic. It was not a large animal; the skull of L. inexpectatus was only about 4.53 centimeters long (1.78 in). A complete specimen of Lazarussuchus with preserved soft tissue was found from the Late Paleocene of France, but has not been assigned to a species.
Susan E. Evans is British palaeontologist and herpetologist. She is the author or co-author of over 100 peer-reviewed papers and book chapters.
Huehuecuetzpalli mixtecus is an extinct lizard from the Early Cretaceous Tlayúa Formation in Tepexi de Rodríguez, Central Mexico. Although it is not the oldest known lizard, Huehuecuetzpalli may be the most basal member of Squamata, making it an important taxon in understanding the origins of squamates. It may or may not be a basal member of Iguania, a large clade of lizards that traditionally includes the iguanas and their close relatives, chameleons, and agamids: if it is an iguanian, H. mixtecus represents the earliest major offshoot of the squamate evolutionary tree.
Bahndwivici is an extinct genus of lizard known from a nearly complete and articulated skeleton discovered in rocks of the Green River Formation of Wyoming, United States. The skeleton is very similar to that of the modern Chinese crocodile lizard, Shinisaurus.
Afairiguana avius is an extinct iguanid lizard known from a nearly complete and articulated skeleton discovered in rocks of the Early Eocene-aged Green River Formation of Wyoming, United States. As of the initial description, the skeleton represents the oldest complete iguanian from the Western Hemisphere, and is the oldest representative of the extant iguanid family of anoles, Polychrotidae.
Barbaturex is an extinct genus of giant herbivorous iguanian lizards from the Eocene of Myanmar. It is represented by a single species, Barbaturex morrisoni, which is known from several partial dentaries and a fused pair of frontals, two bones that form part of the top of the skull. Based on the size of these bones, Barbaturex morrisoni is estimated to have been about 1 metre (3.3 ft) from snout to vent, and possibly up to 6 feet (1.8 m) including the tail. Barbaturex morrisoni was named after The Doors frontman Jim Morrison, a play on his epithet "The Lizard King". The genus's name is a portmanteau of the Latin words Barbatus and rex, meaning "bearded king", in reference to ridges along the mandible and the lizard's large size.
Shinisauria is a clade or evolutionary grouping of anguimorph lizards that includes the living Chinese crocodile lizard Shinisaurus and several of its closest extinct relatives. Shinisauria was named in 2008 as a stem-based taxon to include all anguimorphs more closely related to Shinisaurus than to any other lizard. Several recent phylogenetic analyses of lizard evolutionary relationships place Shinisauria in a basal position within the clade Platynota, which also includes monitor lizards, helodermatids, and the extinct mosasaurs. Shinisaurians were once thought to be closely related to the genus Xenosaurus, but they are now considered distant relatives within Anguimorpha. The fossil record of shinisaurians extends back to the Early Cretaceous with Dalinghosaurus, which is from the Yixian Formation of China. Two other extinct shinisaurians are currently known: Bahndwivici from the Eocene of Wyoming and Merkurosaurus from the Early Miocene of the Czech Republic.
Spathorhynchus is an extinct genus of amphisbaenians or worm lizards that existed from the Middle Eocene to the Early Oligocene in what is now Wyoming. It includes two species, the type species S. fossorium, named in 1973 from the Middle Eocene Bridger and Wind River Formations, and the species S. natronicus, named in 1977 from the Lower Oligocene White River Formation. Spathorhynchus belongs to the family Rhineuridae, which includes many other extinct species that ranged across North America at various times in the Cenozoic but only has one surviving member, Rhineura floridana, from Florida. Spathorhynchus differs from all other rhineurids except Dyticonastis from the Late Oligocene-Early Miocene of Oregon in having a slightly widened, spatula-shaped snout tip with a low angle of about 30 degrees. The two taxa may be closely related, having evolved in isolation in western North America after the formation of the Rocky Mountains separated them from rhineurids further east.
Suzanniwana is an extinct genus of iguanian lizards that lived in western North America during the earliest Eocene, approximately 56 million years ago. Two species are known from the Bighorn Basin of Wyoming: the type species S. patriciana named in 2009, and the species S. revenanta named in 2013. Suzanniwana lived during the Paleocene–Eocene Thermal Maximum, a brief period of global warming that resulted in warmer and drier conditions in the Bighorn Basin. It likely stemmed from a lineage that had migrated into the basin from regions farther to the south, following a latitudinal band of constant climatic conditions that moved northward as the planet warmed. Suzanniwana shares many skeletal features with modern casquehead lizards of the family Corytophanidae and may be a stem-corytophanid. It also closely resembles Geiseltaliellus, an iguanian from the middle Eocene Messel pit in Germany.
Anolbanolis is an extinct genus of iguanian lizards that lived in the Bighorn Basin of what is now Wyoming during the earliest Eocene. The type species A. banalis was named by paleontologist Krister Smith in 2009 from a collection of isolated skull fragments found in a layer of the Willwood Formation that dates to a brief period of global warming called the Paleocene-Eocene thermal maximum (PETM) about 56 million years ago. The genus name Anolbanolis means "poor anole" from the Greek anolbos and the name Anolis, in reference to the scrappy nature of known fossil material and its close resemblance to lizards in the genus Anolis. Smith suggested that Anolbanolis may be a close relative of Anolis or Polychrus, which are common today in Central and South America but not found as far north as Wyoming, fitting with the idea that the Bighorn Basin was warmer and wetter in the Eocene than it is currently. According to Smith, the species name banalis is a reference to it being a "banal" lizard in the Willwood, being "quite abundant in the type locality," and not unusual because "the discovery of Eocene boreal fossil members of living subtropical and tropical clades is becoming commonplace." In 2011 Smith named a second species of Anolbanolis, A. geminus, which lived in the Bighorn Basin shortly after the PETM and was about 50 percent larger than A. banalis. Smith concluded on the basis of features in A. geminus that Anolbanolis is more closely related to Anolis than it is to Polychrus.
Chamaeleontiformes is a hypothesized clade of iguanian lizards defined as all taxa sharing a more recent common ancestor with Chamaeleo chamaeleon than with Hoplocercus spinosus, Polychrus marmoratus, or Iguana iguana. It was named by paleontologist Jack Conrad in 2008 to describe a clade recovered in his phylogenetic analysis that included the extinct genus Isodontosaurus, the extinct family Priscagamidae, and the living clade Acrodonta, which includes agamids and chameleons. It is a stem-based taxon and one of two major clades within Iguania, the other being Pleurodonta. Below is a cladogram from Daza et al. (2012) showing this phylogeny:
Suratagama is an extinct genus of agamid lizard known from the type species Suratagama neeraae from the early Eocene of India. It was named in 2013 on the basis of three isolated jaw bones from the Cambay Shale in Gujarat.
Priscagamidae is an extinct family of iguanian lizards known from the Late Cretaceous of Mongolia and China and the Eocene of India, spanning a range from 75 to 54 million years ago. It includes the genera Heterodontagama, Mimeosaurus, Phrynosomimus, Priscagama, and possibly Pleurodontagama. The first fossils of priscagamids were found in the Djadochta and Khermeen Tsav formations of Mongolia. More recently they have been found in the Cambay Formation in India, leading to the naming of Heterodontagama in 2013. Priscagamidae was originally described as a subfamily of Agamidae called Priscagaminae in 1984, but it was reclassified as a distinct family in 1989. Most phylogenetic analyses still find a close relationship between Priscagamidae and Agamidae, although a 2015 study found it to be basal to all other iguanian clades, warranting its removal from Iguania and placement in a larger clade called Iguanomorpha.
Heterodontagama is an extinct genus of iguanian lizard from the Early Eocene of India. It belongs to the extinct family Priscagamidae, which is otherwise only known from the Late Cretaceous of Mongolia. The type species Heterodontagama borsukae was named in 2013 from several isolated upper and lower jaws found in an exposure of the Cambay Shale in an open-pit coal mine in Gujarat.
Gueragama is an extinct genus of iguanian lizard from the Late Cretaceous of Brazil. It belongs to a group of iguanians called Acrodonta, whose living members include chameleons and agamids and are currently restricted to the Old World. Gueragama is the only acrodont known from South America, providing evidence that the group once ranged across much of Gondwana and only became restricted to the Old World after the supercontinent broke apart. The type species, Gueragama sulamericana, was named in 2015 on the basis of an isolated lower jaw from the Turonian- to Campanian-age Goio-Erê Formation in the Bauru Basin, which was deposited in a desert environment. Unlike modern acrodontans, whose teeth implant on the margins of the jaws, Gueragama has teeth that implant along the inner surface of the lower jaw, a feature common in most non-acrodontan lizards and characteristic of the second major group of iguanians, Pleurodonta. The non-acrodont dentition of Gueragama is evidence of its basal position within Acrodonta, and is shared with the taxon of Late Cretaceous acrodontan relatives, the family Priscagamidae.
Bharatagama is an extinct genus of iguanian lizard from the Early Jurassic of India. It is one of the oldest known lizards and the oldest known iguanian. The type and only species is Bharatagama rebbanensis, named in 2002. Over one hundred fossils of Bharatagama have been found in the Kota Formation, which outcrops in the Pranhita–Godavari Basin and dates back to about 190 million years ago (Ma). Despite its abundance, Bharatagama is known only from isolated jaw bones mixed together in microvertebrate assemblages with equally fragmentary remains of fish, sphenodontians, dinosaurs, crocodylomorphs, and mammals. These fossils represent all stages of development, from hatchlings to adults. The total length of the skull in adult specimens is estimated to have been about 15 millimetres (0.59 in).
Geiseltaliellus is an extinct genus of iguanian lizards that lived in what is now western Europe during the Eocene. It belongs to the family Corytophanidae, which includes modern casquehead lizards. Many fossils are known from Germany, France, and Belgium, with the most well preserved coming from the Messel pit lagerstätte in Messel, Germany. German paleontologist Oskar Kuhn named the genus in 1944 after the Geiseltal valley where the first specimens were found, designating the type species Geiseltaliellus longicaudus. Three new species — G. louisi, G. lamandini, and G. grisolli — were named in the 1990s and 2000s on the basis of more fragmentary remains from France and Belgium, although G. louisi has since been synonymized with G. longicaudus. In 2009 the Messel pit specimens were recognized as belonging to a species distinct from that of the G. longicaudus specimens in Geiseltalt and were collectively reclassified under a new name, G. maarius.
