Anolbanolis | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Squamata |
Suborder: | Iguania |
Clade: | Pleurodonta |
Genus: | † Anolbanolis Smith, 2009 |
Species | |
†A. banalisSmith, 2009 (type) |
Anolbanolis is an extinct genus of iguanian lizards that lived in the Bighorn Basin of what is now Wyoming during the earliest Eocene. The type species A. banalis was named by paleontologist Krister Smith in 2009 from a collection of isolated skull fragments found in a layer of the Willwood Formation that dates to a brief period of global warming called the Paleocene-Eocene thermal maximum (PETM) about 56 million years ago. The genus name Anolbanolis means "poor anole" from the Greek anolbos ("poor, wretched") and the name Anolis, in reference to the scrappy nature of known fossil material and its close resemblance to lizards in the genus Anolis . Smith suggested that Anolbanolis may be a close relative of Anolis or Polychrus , which are common today in Central and South America but not found as far north as Wyoming, fitting with the idea that the Bighorn Basin was warmer and wetter in the Eocene than it is currently. According to Smith, the species name banalis is a reference to it being a "banal" lizard in the Willwood, being "quite abundant in the type locality," and not unusual because "the discovery of Eocene boreal fossil members of living subtropical and tropical clades is becoming commonplace." [1] In 2011 Smith named a second species of Anolbanolis, A. geminus, which lived in the Bighorn Basin shortly after the PETM and was about 50 percent larger than A. banalis. Smith concluded on the basis of features in A. geminus that Anolbanolis is more closely related to Anolis than it is to Polychrus. [2]
Microcosmodon is a mammal genus from the Paleocene of North America. It was a member of the extinct order Multituberculata, and lies within the suborder Cimolodonta and family Microcosmodontidae. The genus Microcosmodon was named by G.L. Jepsen in 1930.
Polychrus is the only extant genus of polychrotid lizards in the world. Commonly called bush anoles, they are found in Central and South America, as well as nearby Trinidad and Tobago.
The Bighorn Basin is a plateau region and intermontane basin, approximately 100 miles (160 km) wide, in north-central Wyoming in the United States. It is bounded by the Absaroka Range on the west, the Pryor Mountains on the north, the Bighorn Mountains on the east, and the Owl Creek Mountains and Bridger Mountains on the south. It is drained to the north by tributaries of the Bighorn River, which enters the basin from the south, through a gap between the Owl Creek and Bridger Mountains, as the Wind River, and becomes the Bighorn as it enters the basin. The region is semi-arid, receiving only 6–10 in (15–25 cm) of rain annually.
Gastornis is an extinct genus of large flightless birds that lived during the late Paleocene and Eocene epochs of the Cenozoic era. The genus is currently thought to contain three or four distinct species, known from incomplete fossil remains, found in western-central Europe. More complete specimens are known from a fourth, North American species, which had previously been classified in the distinct genus Diatryma. Many scientists now consider Diatryma to be so similar to the other species of Gastornis that it should also be included in that genus. A fifth species, also previously classified in its own genus, is known from China.
Sifrhippus is an extinct genus of equid containing the species S. sandrae and S. grangeri. Sifrhippus is the oldest known equid, living during the early Eocene. Its fossils were discovered in the Bighorn Basin of Wyoming. A 2012 study found Arenahippus to be synonymous with Sifrhippus.
Lithornis is a genus of extinct paleognathous birds. Lithornis were able to fly well, but are closely related to today's tinamous and ratites.
Paracathartes is a genus of extinct bird from the Wasachtian horizon of lower Eocene Wyoming, USA. One species, Paracathartes howardae has been described.
Orthogenysuchus is an extinct genus of caimanine alligatorids. Fossils have been found from the Wasatch Beds of the Willwood Formation of Wyoming, deposited during the early Eocene. The type species is O. olseni. The holotype, known as AMNH 5178, is the only known specimen belonging to the genus and consists of a skull lacking the lower jaws. The braincase is filled in by the matrix and most of the suture lines between bones are indiscernible, making comparisons with other eusuchian material difficult.
Bahndwivici is an extinct genus of lizard known from a nearly complete and articulated skeleton discovered in rocks of the Green River Formation of Wyoming, United States. The skeleton is very similar to that of the modern Chinese crocodile lizard, Shinisaurus.
Afairiguana avius is an extinct iguanid lizard known from a nearly complete and articulated skeleton discovered in rocks of the Early Eocene-aged Green River Formation of Wyoming, United States. As of the initial description, the skeleton represents the oldest complete iguanian from the Western Hemisphere, and is the oldest representative of the extant iguanid family of anoles, Polychrotidae.
Homogalax is an extinct genus of tapir-like odd-toed ungulate. It was described on the basis of several fossil finds from the northwest of the USA, whereby the majority of the remains come from the state of Wyoming. The finds date to the Lower Eocene between 56 and 48 million years ago. In general, Homogalax was very small, only reaching the weight of today's peccaries, with a maximum of 15 kg. Phylogenetic analysis suggests the genus to be a basal member of the clade that includes today's rhinoceros and tapirs. In contrast to these, Homogalax was adapted to fast locomotion.
The Willwood Formation is a sedimentary sequence deposited during the late Paleocene to early Eocene, or Clarkforkian, Wasatchian and Bridgerian in the NALMA classification.
The Polychrotidae family of iguanian lizards contains the living genus Polychrus and the extinct genus Afairiguana. The family Polychrotidae was once thought to encompass all anoles, including those in the genus Anolis. Studies of the evolutionary relationships of anoles based on molecular information has shown that Polychrus is not closely related to Anolis, but instead closer to Hoplocercidae. It is therefore not part of Dactyloidae and instead is treated as the family, Polychrotidae.
The Wasatch Formation (Tw) is an extensive highly fossiliferous geologic formation stretching across several basins in Idaho, Montana Wyoming, Utah and western Colorado. It preserves fossils dating back to the Early Eocene period. The formation defines the Wasatchian or Lostcabinian, a period of time used within the NALMA classification, but the formation ranges in age from the Clarkforkian to Bridgerian.
Anniealexandria is an extinct genus of amphisbaenian lizard known by the type species Anniealexandria gansi from the earliest Eocene of Wyoming. Anniealexandria is the only known member of the family Bipedidae in the fossil record, which otherwise only includes the extant genus Bipes from Mexico. It was named in 2009 in honor of Annie Montague Alexander, founder of the University of California Museum of Paleontology. Remains of Anniealexandria are known only from a single fossil locality in the Bighorn Basin called Castle Gardens, but within the locality its fossils are common in the Willwood Formation, usually consisting of isolated jaw bones and vertebrae. Anniealexandria seems to have been a common component of a paleofauna that included fifteen other lizard species and existed in western North America during a period of global warming in the latest Paleocene and earliest Eocene.
Suzanniwana is an extinct genus of iguanian lizards that lived in western North America during the earliest Eocene, approximately 56 million years ago. Two species are known from the Bighorn Basin of Wyoming: the type species S. patriciana named in 2009, and the species S. revenanta named in 2013. Suzanniwana lived during the Paleocene–Eocene Thermal Maximum, a brief period of global warming that resulted in warmer and drier conditions in the Bighorn Basin. It likely stemmed from a lineage that had migrated into the basin from regions farther to the south, following a latitudinal band of constant climatic conditions that moved northward as the planet warmed. Suzanniwana shares many skeletal features with modern casquehead lizards of the family Corytophanidae and may be a stem-corytophanid. It also closely resembles Geiseltaliellus, an iguanian from the middle Eocene Messel pit in Germany.
Babibasiliscus is an extinct genus of casquehead lizard that lived in what is now Wyoming during the early Eocene, approximately 48 million years ago. The genus is known from a single species, Babibasiliscus alxi, which was named by paleontologist Jack Conrad in 2015 on the basis of a fossilized skull from the Bridger Formation in the Green River Basin. The name Babibasiliscus comes from the Shoshoni word babi, meaning "older male cousin", and Basiliscus, a modern-day genus of casquehead lizards. The specimen is undeformed and nearly complete except for the tip of the snout and the top of the skull, making it unclear whether the distinctive bony crest of living corytophanids was present in prehistoric relatives like Babibasiliscus. The skull is about 42 millimetres (2 in) in length and the entire body is estimated to have been about 0.6 metres (2 ft) long. Bones on the right side of lower jaw of the specimen are thickened and fused together, suggesting that the jaw had broken and healed when the animal was alive.
Magnuviator is a genus of extinct iguanomorph lizard from the Late Cretaceous of Montana, US. It contains one species, M. ovimonsensis, described in 2017 by DeMar et al. from two specimens that were discovered in the Egg Mountain nesting site. Magnuviator is closest related to the Asian Saichangurvel and Temujinia, which form the group Temujiniidae. Unlike other members of the Iguanomorpha, however, Magnuviator bears a distinct articulating notch on its tibia for the ankle bones, which has traditionally been considered a characteristic of non-iguanomorph lizards. The morphology of its teeth suggests that its diet would have mainly consisted of wasps, like the modern phyrnosomatid iguanians Callisaurus and Urosaurus, although it also shows some adaptations to herbivory.
Wyonycteris is a genus of small mammals that existed in the late Paleocene and early Eocene epochs. The type species is Wyonycteris chalix, which lived in Wyoming during the Clarkforkian North American Land Mammal Age of the Paleocene and was originally proposed to be an early form of insectivorous bat. Later re-examination of the material has put this alliance in doubt, and the genus has instead been proposed as belonging to the subfamily Placentidentinae, within the family Nyctitheriidae. Similar fossil material of the same time period found in Europe was later discovered and described as new species, Wyonycteris richardi.
Azygonyx was a small tillodont mammal, likely the size of a cat to raccoon, that lived in North America during the Paleocene and Eocene in the early part of the Cenozoic Era. The only fossils that have been recovered are from the Willwood and Fort Union Formations in the Bighorn Basin of Wyoming, United States, and date to the Clarkforkian to Wasatchian, about 56 to 50 million years ago. Fifty-six collections that have been recovered thus far include the remains of Azygonyx. Azygonyx survived the Paleocene Eocene Thermal Maximum along with other mammals like Phenacodus and Ectocion, both of which were ground-dwelling mammals. Azygonyx probably was a generalist terrestrial mammal that may have roamed around the ground, but was also capable of climbing trees.
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