Squamata is the largest order of reptiles, comprising lizards and snakes. With over 12,162 species, it is also the second-largest order of extant (living) vertebrates, after the perciform fish. Squamates are distinguished by their skins, which bear horny scales or shields, and must periodically engage in molting. They also possess movable quadrate bones, making possible movement of the upper jaw relative to the neurocranium. This is particularly visible in snakes, which are able to open their mouths very widely to accommodate comparatively large prey. Squamates are the most variably sized living reptiles, ranging from the 16 mm (0.63 in) dwarf gecko to the 6.5 m (21 ft) reticulated python. The now-extinct mosasaurs reached lengths over 14 m (46 ft).
The Iguanidae is a family of lizards composed of the iguanas, chuckwallas, and their prehistoric relatives, including the widespread green iguana.
The Late Cretaceous is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after creta, the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period.
The Maastrichtian is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from 72.1 to 66 million years ago. The Maastrichtian was preceded by the Campanian and succeeded by the Danian. It is named after the city of Maastricht, the capital and largest city of the Limburg province in the Netherlands.
Ichthyornithes is an extinct group of toothed avialan dinosaurs very closely related to the common ancestor of all modern birds. They are known from fossil remains found throughout the late Cretaceous period of North America, though only two genera, Ichthyornis and Janavis, are represented by complete enough fossils to have been named. Ichthyornitheans became extinct at the Cretaceous–Paleogene boundary, along with enantiornitheans, all other non-avian dinosaurs, and many other animal and plant groups.
Palaeosaniwa canadensis is an extinct species of carnivorous lizard from the late Cretaceous of North America. The name, given by Charles Whitney Gilmore in 1928, means "ancient Saniwa from Canada".
The Paleocene, or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek παλαιός palaiós meaning "old" and the Eocene Epoch, translating to "the old part of the Eocene".
Pristiguana is an extinct genus of possible iguanian lizard from the Maastrichtian Marília Formation of Brazil. The type species is P. brasiliensis.
Bottosaurus is an extinct genus of alligatorid from the Late Cretaceous-Early Paleocene of New Jersey, Texas, and possibly North Carolina and South Carolina. Two species are currently accepted, with a third requiring re-evaluation.
Saurocephalus is an extinct genus of ray-finned fishes within the family Saurodontidae. The genus was first described in 1824 and contains six or seven species, including the type species S. lanciformis. Saurocephalus first appeared during the early Valanginian and continued on to the Maastrichtian, where it nearly went extinct. However, the recent discovery of S. lanciformis remains from the earliest Paleocene indicates that it just barely survived into the Cenozoic. This would make it the last surviving ichthyodectiform.
Thoracosaurus is an extinct genus of long-snouted eusuchian which existed during the Late Cretaceous and Early Paleocene in North America and Europe.
Socognathus is a genus of prehistoric chamopsiid polyglyphanodontian lizards containing species that lived from the Middle Campanian stage to the late Maastrichtian. Several specimens of the type species, Socognathus unicuspis, have been found in Alberta, Canada. A second species, Socognathus brachyodon is known from the late Maastrichtian Lance Formation; its fossils have been found in Wyoming, United States.
Obamadon is an extinct genus of polyglyphanodontian lizards from the Late Cretaceous of North America. Fossils have been found in the Hell Creek Formation of Montana and the Lance Formation of Wyoming. Researchers describe it as being distinguished by its "tall, slender teeth with large central cusps separated from small accessory cusps by lingual grooves." The type species was named Obamadon gracilis after United States president Barack Obama, "in reference to the tall, straight teeth, and the manner in which Mr. Obama has acted as a role model of good oral hygiene for the world." According to Nicholas R. Longrich of Yale University, the species "was probably a foot long, [and] with these tall, slender teeth it used to eat insects and plant matter."
Polyglyphanodontia, also known as the Borioteiioidea, is an extinct clade of Cretaceous lizards. Polyglyphanodontians were the dominant group of lizards in North America and Asia during the Late Cretaceous.
Acheroraptor is an extinct genus of dromaeosaurid theropod dinosaur known from the latest Maastrichtian Hell Creek Formation of Montana, United States. It contains a single species, Acheroraptor temertyorum. A. temertyorum is one of the two geologically youngest known species of dromaeosaurids, the other being Dakotaraptor steini, which is also known from Hell Creek. A basal cousin of Velociraptor, Acheroraptor is known from upper and lower jaw material.
The Cretaceous–Paleogene (K–Pg) extinction event, also known as the K–T extinction, was the mass extinction of three-quarters of the plant and animal species on Earth approximately 66 million years ago. The event caused the extinction of all non-avian dinosaurs. Most other tetrapods weighing more than 25 kg (55 lb) also became extinct, with the exception of some ectothermic species such as sea turtles and crocodilians. It marked the end of the Cretaceous period, and with it the Mesozoic era, while heralding the beginning of the current era, the Cenozoic. In the geologic record, the K–Pg event is marked by a thin layer of sediment called the K–Pg boundary, Fatkito boundary or K–T boundary, which can be found throughout the world in marine and terrestrial rocks. The boundary clay shows unusually high levels of the metal iridium, which is more common in asteroids than in the Earth's crust.
Archaerhineura was a genus of amphisbaenian lizards in the family Rhineuridae that is now extinct. The only species is Archaerhineura mephitis, named in 2015 on the basis of a single fragment of the lower jaw from the Polecat Bench Formation in Park County, Wyoming, which dates to the late Paleocene. Archaerhineura is one of the oldest amphisbaenians and was part of an evolutionary radiation of Rhineuridae in the Paleocene several million years after the Cretaceous–Paleogene extinction event. This rhineurid radiation coincided with the radiation of another group of amphisbaenians, Amphisbaeniformes, which includes the still-extant families Blanidae and Amphisbaenidae. The presence of Archaerhineura and other Paleocene rhineurids in the western United States indicates that amphisbaenians, which would later have a nearly global distribution, originated in North America.
Chthonophis is an extinct genus of amphisbaenian lizard with only one known species, Chthonophis subterraneus, from the earliest Paleocene of northeastern Montana. Chthonophis was named in 2015 on the basis of a partial lower jaw from an outcrop of the Fort Union Formation in the Bug Creek Anthills. The surfaces of the bone are well-rounded, suggesting that the remains had been partially digested by another animal before the jaw had been buried and fossilized. Chthonophis is the oldest known amphisbaenian, yet phylogenetic analysis shows that it was not the most basal. Longrich et al. (2015) classified Chthonophis in its own family, Chthonophidae, finding it to be more derived than Rhineuridae but more basal than other clades such as Blanidae and Amphisbaenidae. The existence of a derived amphisbaenian soon after the Cretaceous-Paleogene extinction event suggests that Amphisbaenia has its origins in the Cretaceous, although no Cretaceous amphisbaenians are currently known. Below is a cladogram from Longrich et al. (2015) showing the phylogenetic relationships of Chthonophis:
Odaxosaurus is an extinct genus of anguid lizards that existed in western North America from the Late Cretaceous to the Paleocene. Fossils of the type species Odaxosaurus piger and the species O. priscus are widespread throughout Late Cretaceous formations in the western United States and Canada. First described in 1928 from the Lance Formation in Wyoming, O. piger has since been found in the Hell Creek Formation in Wyoming and Montana, the Frenchman and Scollard formations in Alberta, and the Aguja Formation in Texas. It was one of the few species of lizards to survive the Cretaceous–Paleogene extinction event, which is estimated to have killed off 83% of all lizard species. The second species, O. priscus, was named in 1996 from the Dinosaur Park Formation in Alberta and has since been found in the Kaiparowits Formation in southern Utah. Remains of an anguid from the Kirtland Formation in New Mexico may also belong to Odaxosaurus.
Magnuviator is a genus of extinct iguanomorph lizard from the Late Cretaceous of Montana, US. It contains one species, M. ovimonsensis, described in 2017 by DeMar et al. from two specimens that were discovered in the Egg Mountain nesting site. Magnuviator is closest related to the Asian Saichangurvel and Temujinia, which form the group Temujiniidae. Unlike other members of the Iguanomorpha, however, Magnuviator bears a distinct articulating notch on its tibia for the ankle bones, which has traditionally been considered a characteristic of non-iguanomorph lizards. The morphology of its teeth suggests that its diet would have mainly consisted of wasps, like the modern phyrnosomatid iguanians Callisaurus and Urosaurus, although it also shows some adaptations to herbivory.
