Jennifer Alice Clack, was an English palaeontologist and evolutionary biologist. She specialised in the early evolution of tetrapods, specifically studying the "fish to tetrapod" transition: the origin, evolutionary development and radiation of early tetrapods and their relatives among the lobe-finned fishes. She is best known for her book Gaining Ground: the Origin and Early Evolution of Tetrapods, published in 2002 and written with the layperson in mind.
Romer's gap is an example of an apparent gap in the tetrapod fossil record used in the study of evolutionary biology. Such gaps represent periods from which excavators have not yet found relevant fossils. Romer's gap is named after paleontologist Alfred Romer, who first recognised it in 1956. Recent discoveries in Scotland are beginning to close this gap in palaeontological knowledge.
Reptiliomorpha is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians (lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes Homo sapiens, but not Ascaphus truei. Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing Homo sapiens, but not Pipa pipa, Caecilia tentaculata, and Siren lacertina.
The Visean, Viséan or Visian is an age in the ICS geologic timescale or a stage in the stratigraphic column. It is the second stage of the Mississippian, the lower subsystem of the Carboniferous. The Visean lasted from 346.7 to 330.9 Ma. It follows the Tournaisian age/stage and is followed by the Serpukhovian age/stage.
Westlothiana is a genus of reptile-like tetrapod that lived about 338 million years ago during the latest part of the Viséan age of the Carboniferous. Members of the genus bore a superficial resemblance to modern-day lizards. The genus is known from a single species, Westlothiana lizziae. The type specimen was discovered in the East Kirkton Limestone at the East Kirkton Quarry, West Lothian, Scotland in 1984. This specimen was nicknamed "Lizzie the lizard" by fossil hunter Stan Wood, and this name was quickly adopted by other paleontologists and the press. When the specimen was formally named in 1990, it was given the specific name "lizziae" in homage to this nickname. However, despite its similar body shape, Westlothiana is not considered a true lizard. Westlothiana's anatomy contained a mixture of both "labyrinthodont" and reptilian features, and was originally regarded as the oldest known reptile or amniote. However, updated studies have shown that this identification is not entirely accurate. Instead of being one of the first amniotes, Westlothiana was rather a close relative of Amniota. As a result, most paleontologists since the original description place the genus within the group Reptiliomorpha, among other amniote relatives such as diadectomorphs and seymouriamorphs. Later analyses usually place the genus as the earliest diverging member of Lepospondyli, a collection of unusual tetrapods which may be close to amniotes or lissamphibians, or potentially both at the same time.
Proterogyrinus is an extinct genus of early tetrapods from the order Embolomeri. Fossil remains of Proterogyrinus have been found in Scotland, UK, and West Virginia, United States, and date back to the Serpukhovian, which is from about 331 to 323 million years ago. The genus was originally named by renowned vertebrate paleontologist Alfred Sherwood Romer in 1970. A comprehensive redescription was later published by Canadian paleontologist Robert Holmes in 1984. The generic name "Proterogyrinus" is Greek for "earlier wanderer" or "earlier tadpole". This name was chosen by Romer in keeping with a trend of naming long-bodied early tetrapods with the suffix "-gyrinus".
Eucritta is an extinct genus of stem-tetrapod from the Viséan epoch in the Carboniferous period of Scotland. The name of the type and only species, E. melanolimnetes is a homage to the 1954 horror film Creature from the Black Lagoon.
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous (Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous (Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape.
Pholiderpeton is an extinct genus of embolomere amphibian which lived in the Late Carboniferous period (Bashkirian) of England. The genus was first named by Thomas Henry Huxley in 1869 to include the species P. scutigerum, based on the disarticulated front half of a skeleton discovered near Bradford, Yorkshire. Associated fossil wood suggests that this specimen died inside a Lepidodendron tree trunk.
Diplovertebron is an extinct genus of embolomere that lived in the Late Carboniferous period (Moscovian), about 310 million years ago. Diplovertebron was a medium-sized animal, around 50 cm in length. Members of the genus inhabited European Carboniferous swamps in what is now the Czech Republic. They were closely related to larger swamp-dwelling tetrapods like Proterogyrinus and Anthracosaurus. However, Diplovertebron were much smaller than these large, crocodile-like creatures. Known from a single species, Diplovertebron punctatum, this genus has had a complicated history closely tied to Gephyrostegus, another genus of small, reptile-like amphibians.
Anthracosaurus is an extinct genus of embolomere, a possible distant relative of reptiles that lived during the Late Carboniferous in what is now Scotland, England, and Ohio. It was a large, aquatic eel-like predator. It has a robust skull about 40 centimetres (1.3 ft) in length with large teeth in the jaws and on the roof of the mouth. Anthracosaurus probably inhabited swamps, rivers and lakes. Its name is Greek for "coal lizard".
Spathicephalus is an extinct genus of stem tetrapods that lived during the middle of the Carboniferous Period. The genus includes two species: the type species S. mirus from Scotland, which is known from two mostly complete skulls and other cranial material, and the species S. pereger from Nova Scotia, which is known from a single fragment of the skull table. Based on the S. mirus material, the appearance of Spathicephalus is unlike that of any other early tetrapod, with a flattened, square-shaped skull and jaws lined with hundreds of very small chisel-like teeth. However, Spathicephalus shares several anatomical features with a family of stem tetrapods called Baphetidae, leading most paleontologists who have studied the genus to place it within a larger group called Baphetoidea, often as part of its own monotypic family Spathicephalidae. Spathicephalus is thought to have fed on aquatic invertebrates through a combination of suction feeding and filter feeding.
The Allegheny Group, often termed the Allegheny Formation, is a Pennsylvanian-age geological unit in the Appalachian Plateau. It is a major coal-bearing unit in the eastern United States, extending through western and central Pennsylvania, western Maryland and West Virginia, and southeastern Ohio. Fossils of fishes such as Bandringa are known from the Kittaning Formation, which is part of the Allegheny Group.
Neopteroplax is an extinct genus of eogyrinid embolomere closely related to European genera such as Eogyrinus and Pteroplax. Members of this genus were among the largest embolomeres in North America. Neopteroplax is primarily known from a large skull found in Ohio, although fragmentary embolomere fossils from Texas and New Mexico have also been tentatively referred to the genus. Despite its similarities to specific European embolomeres, it can be distinguished from them due to a small number of skull and jaw features, most notably a lower surangular at the upper rear portion of the lower jaw.
East Kirkton Quarry, or simply East Kirkton, is a former limestone quarry in West Lothian, Scotland, now a renowned fossil site. The quarry is known for terrestrial and freshwater fossils about 335 million years old, from the late Viséan stage of the Mississippian subperiod. The quarry is a 200 meter-long (~650 ft) depression located in the town of Bathgate. Geographically, it sits at the Bathgate Hills near the center of the Midland Valley, a fossil-rich region of southeast Scotland. The site is dominated by volcanic tuff, limestone, and silica deposits of large freshwater lakes associated with hot springs and local basaltic (high-iron) volcanism. Three geological intervals are exposed: the East Kirkton Limestone (oldest), Little Cliff Shale (middle), and Geikie Tuff (youngest).
Eldeceeon is an extinct genus of reptiliomorph from the Mississippian of Scotland. It is known from two fossil specimens found within the Viséan-age East Kirkton Quarry in West Lothian. The type and only species, E. rolfei, was named in 1994. Eldeceeon is thought to be closely related to embolomeres, but it has several distinguishing features including long limbs and a short trunk. Initially known from two crushed partial skeletons, additional specimens have been reported by Ruta & Clack (2006). Eldeceeon was redescribed by Ruta, Clack, & Smithson (2020). The redescription supported affinities with Silvanerpeton, reconstructed a skull with larger eyes and a shorter snout, and emphasized potential correlations for an enlarge puboischiofemoralis internus 2 muscle.
The Bluefield Formation is a geologic formation in West Virginia. It preserves fossils dating back to the Mississippian subperiod of the Carboniferous period. Sediments of this age formed along a large marine basin lying in the region of what is now the Appalachian Plateau. The Bluefield Formation is the lowest section of the primarily siliciclastic Mauch Chunk Group, underlying the Stony Gap Sandstone Member of the Hinton Formation and overlying the limestone-rich Greenbrier Group.
The Ballagan Formation is a geologic formation in Scotland and England. It preserves fossils dating back to the early part of the Carboniferous period. Its name comes from the "Ballagan Beds" of Ballagan Glen, near Strathblane, which has a good example of this geological formation.
Stan Wood was a self-taught fossil hunter known for his significant palaeontological discoveries at scale and is arguably most celebrated for refocusing attention on early Carboniferous palaeobiology as the means to closing Romer's gap.
Eobaphetes is an extinct genus of embolomere which likely lived in the Pennsylvanian of Kansas. The genus is based on several skull and jaw fragments of a single individual. They were originally described under the species Erpetosuchus kansasensis, but this was later changed to Eobaphetes kansasensis when it was determined that Erpetosuchus was preoccupied by a Triassic reptile.
