"Labyrinthodontia" is an informal grouping of extinct predatory amphibians which were major components of ecosystems in the late Paleozoic and early Mesozoic eras. Traditionally considered a subclass of the class Amphibia, modern classification systems recognize that labyrinthodonts are not a formal natural group (clade) exclusive of other tetrapods. Instead, they consistute an evolutionary grade, ancestral to living tetrapods such as lissamphibians and amniotes. "Labyrinthodont"-grade vertebrates evolved from lobe-finned fishes in the Devonian, though a formal boundary between fish and amphibian is difficult to define at this point in time.
Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and armour-like bony plates that distinguish them from the modern soft-bodied lissamphibians.
Zatracheidae is a family of Late Carboniferous and Early Permian temnospondyls known from North America and Europe. Zatracheidids are distinguished by lateral (sideways) bony protuberances of the quadratojugal bone of the skull, and a large opening in the snout called the internarial fontanelle that is bordered by enlarged premaxillae. The skull is flattened, with small orbits or eye sockets set far back. The opening in the snout may have housed a gland for producing a sticky substance so that prey would adhere to the tongue. If so, this indicates that these animals spent a large part of their time on land.
Proterogyrinus is an extinct genus of early tetrapods from the order Embolomeri. Fossil remains of Proterogyrinus have been found in Scotland, UK, and West Virginia, United States, and date back to the Serpukhovian, which is from about 331 to 323 million years ago. The genus was originally named by renowned vertebrate paleontologist Alfred Sherwood Romer in 1970. A comprehensive redescription was later published by Canadian paleontologist Robert Holmes in 1984. The generic name "Proterogyrinus" is Greek for "earlier wanderer" or "earlier tadpole". This name was chosen by Romer in keeping with a trend of naming long-bodied early tetrapods with the suffix "-gyrinus".
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous (Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous (Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape.
Pholiderpeton is an extinct genus of embolomere amphibian which lived in the Late Carboniferous period (Bashkirian) of England. The genus was first named by Thomas Henry Huxley in 1869 to include the species P. scutigerum, based on the disarticulated front half of a skeleton discovered near Bradford, Yorkshire. Associated fossil wood suggests that this specimen died inside a Lepidodendron tree trunk.
Diplovertebron is an extinct genus of embolomere that lived in the Late Carboniferous period (Moscovian), about 310 million years ago. Diplovertebron was a medium-sized animal, around 50 cm in length. Members of the genus inhabited European Carboniferous swamps in what is now the Czech Republic. They were closely related to larger swamp-dwelling tetrapods like Proterogyrinus and Anthracosaurus. However, Diplovertebron were much smaller than these large, crocodile-like creatures. Known from a single species, Diplovertebron punctatum, this genus has had a complicated history closely tied to Gephyrostegus, another genus of small, reptile-like amphibians.
Anthracosaurus is an extinct genus of embolomere, a possible distant relative of reptiles that lived during the Late Carboniferous in what is now Scotland, England, and Ohio. It was a large, aquatic eel-like predator. It has a robust skull about 40 centimetres (1.3 ft) in length with large teeth in the jaws and on the roof of the mouth. Anthracosaurus probably inhabited swamps, rivers and lakes. Its name is Greek for "coal lizard".
Pteroplax is an extinct genus of embolomerous anthracosaur. Only one species has been described; the skull table noted is the lectotype of this species. Pteroplax dates from the late Carboniferous Period, about 315 million years ago. It is known with certainty only from Newsham in Blyth, Northumberland, England. At that site, it shared a coal-swamp lake habitat with the larger embolomere, "Eogyrinus". Pteroplax probably grew up to about 10 ft (3.0 m) in length and was largely aquatic, feeding upon fish and smaller tetrapods. It likely had a long, eel-like body, with short limbs and a long tail.
Gephyrostegus is a genus of extinct gephyrostegid reptiliomorph amphibian. It was a small animal at 22 cm snout-vent length, of generally lizard-like build and presumably habit. It had large eyes and a large number of small, pointed teeth, indicating it was an active insectivorous hunter. The remains have been found in Nýřany, Czech Republic, dating from around 310 million years ago.
Megalocephalus is an extinct genus of baphetid amphibian from the late Carboniferous of the British Isles and the United States (Ohio). It contains two species, M. pachycephalus and M. lineolatus.
Neopteroplax is an extinct genus of eogyrinid embolomere closely related to European genera such as Eogyrinus and Pteroplax. Members of this genus were among the largest embolomeres in North America. Neopteroplax is primarily known from a large skull found in Ohio, although fragmentary embolomere fossils from Texas and New Mexico have also been tentatively referred to the genus. Despite its similarities to specific European embolomeres, it can be distinguished from them due to a small number of skull and jaw features, most notably a lower surangular at the upper rear portion of the lower jaw.
Dictyocephalus is an extinct genus of prehistoric temnospondyls; the only species is Dictyocephalus elegans. This taxon was one of the first metoposaurids to be discovered in North America, being discovered by Ebeneezer Emmons and briefly described by Joseph Leidy in 1856 in the Newark supergroup exposures of Chatham County, North Carolina. At the time, Leidy was uncertain of much of the anatomy of D. elegans, which is represented only by a small partial skull and made only brief descriptions and measurements of a few elements, with an estimated size based on the long-snouted trematosaur Trematosaurus. Emmons provided the first figures of the specimen the following year. Romer (1947) briefly mentioned that the specimen was indistinguishable from "Buettneria" (Koskinonodon).
Lydekkerina is an extinct genus of stereospondyl temnospondyl. It is the type genus of the family Lydekkerinidae. Fossils have been collected from Early Triassic deposits in South Africa and Australia. The type species is L. huxleyi, first described in 1889. While most other stereospondyls were semiaquatic, Lydekkerina was exclusively terrestrial.
Micropholis is an extinct genus of dissorophoid temnospondyl. Fossils have been found from the Lystrosaurus Assemblage Zone of the Karoo Basin in South Africa and are dated to the Induan. Fossils have also been found from the lower Fremouw of Antarctica.Micropholis is the only post-Permian dissorophoid and the only dissorophoid in what is presently the southern hemisphere and what would have been termed Gondwana during the amalgamation of Pangea.
Anaschisma is an extinct genus of large temnospondyls. These animals were part of the family called Metoposauridae, which filled the crocodile-like predatory niches in the late Triassic. It had large skull about 62 centimetres (24 in) long, and possibly reached 3 metres (9.8 ft) long. It was an ambush hunter, snapping up anything small enough to fit in its huge jaws. It was very common during the Late Triassic in what is now the American Southwest.
Parioxys is an extinct genus of temnospondyl amphibian from the Early Permian of Texas.
Spondylerpeton is an extinct genus of embolomere closely related to "Cricotus" (Archeria) in the family Archeriidae. This genus is known from fragmentary remains, namely a short series of tail vertebrae preserved in an ironstone nodule. These remains were found in the Mazon Creek beds of Illinois, an area famed for its preservation of Carboniferous plants and animals. Spondylerpeton individuals were probably about three to four feet in length, by far the largest animals known to have inhabited the Mazon Creek area during this era.
Papposaurus is an extinct genus of proterogyrinid embolomere which lived in the Mississippian of Scotland. It is known from a single species, Papposaurus traquiairi, which is based on an isolated femur discovered in ironstone near Loanhead. Though originally compared closely to reptiles, subsequent study has revealed closer similarity to basal embolomeres such as Proterogyrinus and Archeria. With such limited remains, Papposaurus may not be a valid genus. The femur was redescribed in 1986 by T. R. Smithson, who considered Papposaurus traquairi a nomen vanum possibly synonymous with Proterogyrinus scheelei.
Eobaphetes is an extinct genus of embolomere which likely lived in the Pennsylvanian of Kansas. The genus is based on several skull and jaw fragments of a single individual. They were originally described under the species Erpetosuchus kansasensis, but this was later changed to Eobaphetes kansasensis when it was determined that Erpetosuchus was preoccupied by a Triassic reptile.
