Black oak | |
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Planted tree | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Rosids |
Order: | Fagales |
Family: | Fagaceae |
Genus: | Quercus |
Subgenus: | Quercus subg. Quercus |
Section: | Quercus sect. Lobatae |
Species: | Q. velutina |
Binomial name | |
Quercus velutina | |
Generalized natural range | |
Synonyms [3] | |
List
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Quercus velutina (Latin 'velutina', "velvety") , the black oak, is a species of oak in the red oak group (Quercus sect. Lobatae), native and widespread in eastern and central North America. It is sometimes called the eastern black oak. [4]
Quercus velutina was previously known as yellow oak due to the yellow pigment in its inner bark. It is a close relative of the California black oak (Quercus kelloggii) found in western North America.
In the northern part of its range, Quercus velutina is a relatively small tree, reaching a height of 20–25 metres (66–82 feet) and a diameter of 90 centimetres (35 inches), but it grows larger in the south and center of its range, where heights of up to 42 m (138 ft) are known.
The leaves of the black oak are alternately arranged on the twig and are 10–20 cm (4–8 in) long with 5–7 bristle-tipped lobes separated by deep U-shaped notches. The upper surface of the leaf is a shiny deep green, and the lower is yellowish-brown. There are also stellate hairs on the underside of the leaf that grow in clumps. [5] Some key characteristics for identification include that leaves grown in the sun have very deep U-shaped sinuses and that the buds are velvety and covered in white hairs. [5] [6]
Black oak is monoecious. The staminate flowers develop from leaf axils of the previous year and the catkins emerge before or at the same time as the current leaves in April or May. The pistillate flowers are borne in the axils of the current year's leaves and may be solitary or occur in two- to many-flowered spikes. The fruit, an acorn that occurs singly or in clusters of two to five, is about one-third enclosed in a scaly cup and matures in 2 years. Black oak acorns are brown when mature and ripen from late August to late October, depending on geographic location.
The fruits or acorns of the black oak are medium-sized and broadly rounded. [7] The cap is large and covers almost half of the nut. [5]
Black oak is found in all the coastal states from Maine to Texas, inland as far as Michigan, Ontario, Minnesota, Nebraska, Kansas, Oklahoma, and eastern Texas. [8]
It grows on all aspects and slope positions. It grows best in coves and on middle and lower slopes with northerly and easterly aspects. It is found at elevations up to 1,200 m (3,900 ft) in the southern Appalachians. In southern New England, black oak grows on cool, moist soils. Elsewhere it occurs on warm, moist soils. The most widespread soils on which black oak grows are the udalfs and udolls. These soils are derived from glacial materials, sandstones, shales, and limestone and range from heavy clays to loamy sands with some having a high content of rock or chert fragments. Black oak grows best on well drained, silty clay to loam soils.
The most important factors determining site quality for black oak are the thickness and texture of the A horizon, texture of the B horizon, aspect, and slope position. Other factors may be important in localized areas. For example, in northwestern West Virginia increasing precipitation to 1,120 millimetres (44 in) resulted in increased site quality; more than 1,120 mm (44 in) had no further effect. In southern Indiana, decreasing site quality was associated with increasing slope steepness.
Near the limits of its range, topographic factors may restrict its distribution. At the western limits black oak is often found only on north and east aspects where moisture conditions are most favorable. In southern Minnesota and Wisconsin it is usually found only on ridge tops and the lower two-thirds of south- and west-facing slopes.
Common tree associates of black oak are white oak ( Quercus alba ), northern red oak ( Quercus rubra ), pignut hickory ( Carya glabra ), mockernut hickory (C. tomentosa), bitternut hickory (C. cordiformis), and shagbark hickory (C. ovata); American elm ( Ulmus americana ) and slippery elm (U. rubra); white ash ( Fraxinus americana ); black walnut ( Juglans nigra ) and butternut (J. cinerea); scarlet oak ( Quercus coccinea ), southern red oak (Q. falcata), and chinkapin oak (Q. muehlenbergii); red maple ( Acer rubrum ) and sugar maple (A. saccharum); black cherry ( Prunus serotina ); and blackgum ( Nyssa sylvatica ).[ citation needed ]
Common small tree associates of black oak include flowering dogwood ( Cornus florida ), sourwood ( Oxydendrum arboreum ), sassafras ( Sassafras albidum ), eastern hophornbeam ( Ostrya virginiana ), redbud ( Cercis canadensis ), pawpaw ( Asimina triloba ), downy serviceberry ( Amelanchier arborea ), and American bladdernut ( Staphylea trifolia ). Common shrubs include Vaccinium spp., mountain-laurel ( Kalmia latifolia ), witch-hazel ( Hamamelis virginiana ), beaked hazel ( Corylus cornuta ), spicebush ( Lindera benzoin ), sumac (Rhus spp.), and Viburnum spp. The most common vines are greenbrier ( Smilax spp.), grape ( Vitis spp.), poison-ivy ( Toxicodendron radicans ), and Virginia creeper ( Parthenocissus quinquefolia ).[ citation needed ]
Black oak is often a predominant species in the canopy of an oak–heath forest. [9] [10]
In forest stands, black oak begins to produce seeds at about age 20 and reaches optimum production at 40 to 75 years. It is a consistent seed producer with good crops of acorns every 2 to 3 years. In Missouri, the average number of mature acorns per tree was generally higher than for other oaks over a 5-year period, but the number of acorns differed greatly from year to year and from tree to tree within the same stand.
The number of seeds that become available for regenerating black oak may be low even in good seed years. Insects, squirrels, deer, turkey, small rodents, and birds consume many acorns. They can eat or damage a high percentage of the acorn crop in most years and essentially all of it in poor seed years.
Black oak acorns from a single tree are dispersed over a limited area by squirrels, mice, and gravity. The blue jay may disperse over longer distances.
Black oak is classed as intermediate in tolerance to shade. It is less tolerant than many of its associates such as white and chestnut oaks, hickories, beech ( Fagus grandifolia ), maples, elm, and blackgum. However, it is more tolerant than yellow-poplar (Liriodendron tulipifera), black cherry, and shortleaf pine ( Pinus echinata ). It is about the same as northern red oak and scarlet oak. Seedlings usually die within a few years after being established under fully stocked over stories. Most black oak sprouts under mature stands develop crooked stems and flat-topped or misshapen crowns. After the over story is removed, only the large stems are capable of competing successfully. Seedlings are soon overtopped. The few that survive usually remain in the intermediate crown class.
Even-aged silvicultural systems satisfy the reproduction and growth requirements of black oak better than the all-aged or uneven-aged selection system. Under the selection system, black oak is unable to reproduce because of inadequate light. Stands containing black oak that are managed under the selection system will gradually be dominated by more shade-tolerant species.
Dormant buds are numerous on the boles of black oak trees. These buds may be stimulated to sprout and produce branches by mechanical pruning or by exposure to greatly increased light, as by thinning heavily or creating openings in the stand. Dominant trees are less likely to produce epicormic branches than those in the lower crown class.
Wildfires seriously damage black oak trees by killing the cambium at the base of the trees. This creates an entry point for decay fungi. The result is loss of volume because of heart rot. Trees up to pole size are easily killed by fire and severe fires may even kill saw timber. Many of the killed trees sprout and form a new stand. However, the economic loss may be large unless at least some of it can be salvaged.
Oak wilt ( Bretziella fagacearum ) is a potentially serious vascular disease of black oak that is widespread throughout the eastern United States. Trees die within a few weeks after the symptoms first appear. Usually scattered individuals or small groups of trees are killed, but areas several hectares in size may be affected. The disease is spread from tree to tree through root grafts and over larger distances by sap-feeding beetles (Nitidulidae) and the small oak bark beetle.
Shoestring root rot ( Armillaria mellea ) attacks black oak and may kill trees weakened by fire, lightning, drought, insects, or other diseases. A root rot, Phytophthora cinnamomi , may kill seedlings in the nursery. Cankers caused by Strumella and Nectria species damage the holes of black oak but seldom kill trees. Foliage diseases that attack black oak are the same as those that typically attack species in the red oak group and include anthracnose ( Gnomonia quercina ), leaf blister ( Taphrina spp.), powdery mildews ( Phyllactinia corylea and Microsphaera alni ), oak-pine rusts ( Cronartium spp.), and leaf spots ( Actinopelte dryina ).
Tunneling insects that attack the boles of black oak and cause serious lumber degrade include the carpenter worm ( Prionoxystus robiniae ), red oak borer ( Enaphalodes rufulus ), the twolined chestnut borer ( Agrilus bilineatus ), the oak timber worm ( Arrenodes minutus ), and the Columbian timber beetle ( Corthylus columbianus ).
The gypsy moth ( Lymantria dispar ) feeds on foliage and is potentially the most destructive insect. Although black oaks withstood a single defoliation, two or three defoliations in successive years kill many trees. Other defoliators that attack black oak and may occasionally be epidemic are the variable oak leaf caterpillar ( Heterocampa manteo ), the orange striped oakworm ( Anisota senatoria ), and the brown tail moth ( Euproctis chrysorrhoea ).
The nut weevils ( Curculio spp.), gall-forming cynipids ( Callirhytis spp.), filbertworm ( Melissopus latiferreanus ), and acorn moth ( Valentinia glandulella ) damage black oak acorns.
Black oak is well known to readily hybridize with other members of the red oak (Quercus sect. Lobatae) group, being one parent in at least a dozen different named hybrids. [5]
The inner bark of the black oak contains a yellow-orange coloring from the pigment quercitron, which was sold commercially in Europe until the 1940s, and lending the species its former common name of yellow oak. [11]
An oak is a hardwood tree or shrub in the genus Quercus of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisphere; it includes some 500 species, both deciduous and evergreen. Fossil oaks date back to the Middle Eocene. Molecular phylogeny shows that the genus is divided into Old World and New World clades, but many oak species hybridise freely, making the genus's history difficult to resolve.
Quercus kelloggii, the California black oak or Kellogg oak, is an oak in the red oak section native to western North America. Although genetically separated from them for more than 20 million years, its leaves are remarkably similar in appearance to several other members of the red oak section including the red oak and the black oak found in eastern and central North America.
Quercus rubra, the northern red oak, is an oak tree in the red oak group. It is a native of North America, in the eastern and central United States and southeast and south-central Canada. It has been introduced to small areas in Western Europe, where it can frequently be seen cultivated in gardens and parks. It prefers good soil that is slightly acidic. Often simply called red oak, northern red oak is so named to distinguish it from southern red oak (Q. falcata), also known as the Spanish oak. Northern red oak is sometimes called champion oak.
Quercus palustris, also called pin oak, swamp oak, or Spanish oak, is a tree in the red oak section of the genus Quercus. Pin oak is one of the most commonly used landscaping oaks in its native range due to its ease of transplant, relatively fast growth, and pollution tolerance.
Quercus muehlenbergii, the chinquapinoak, is a deciduous species of tree in the white oak group. The species was often called Quercus acuminata in older literature. Quercus muehlenbergii is native to eastern and central North America. It ranges from Vermont to Minnesota, south to the Florida panhandle, and west to New Mexico in the United States. In Canada it is only found in southern Ontario, and in Mexico it ranges from Coahuila south to Hidalgo.
Quercus falcata, also called southern red oak, spanish oak, bottomland red oak or three-lobed red oak is an oak. Native to the southeastern United States, it gets its name the "Spanish Oak" as these are the areas of early Spanish colonies, whilst "southern red oak" comes from both its range and leaf color during late summer and fall. The southern red oak is a deciduous angiosperm, so has leaves that die after each growing period and come back in the next period of growth.
The fox squirrel, also known as the eastern fox squirrel or Bryant's fox squirrel, is the largest species of tree squirrel native to North America. Despite the differences in size and coloration, it is sometimes mistaken for American red squirrels or eastern gray squirrels in areas where the species co-exist.
Quercus coccinea, the scarlet oak, is a deciduous tree in the red oak section Lobatae of the genus Quercus, in the family Fagaceae.
Quercus lyrata, the overcup oak, is an oak in the white oak group. The common name, overcup oak, refers to its acorns that are mostly enclosed within the acorn cup. It is native to lowland wetlands in the eastern and south-central United States, in all the coastal states from New Jersey to Texas, inland as far as Oklahoma, Missouri, and Illinois. There are historical reports of it growing in Iowa, but the species appears to have been extirpated there. It is a slow-growing tree that often takes 25 to 30 years to mature. It has an estimated lifespan of 400 years.
The Appalachian mixed mesophytic forests is an ecoregion of the temperate broadleaf and mixed forests biome, as defined by the World Wildlife Fund. It consists of mesophytic plants west of the Appalachian Mountains in the Southeastern United States.
Quercus nigra, the water oak, is an oak in the red oak group, native to the eastern and south-central United States, found in all the coastal states from New Jersey to Texas, and inland as far as Oklahoma, Kentucky, and southern Missouri. It occurs in lowlands and up to 450 meters in elevation.
The Appalachian–Blue Ridge forests are an ecoregion in the Temperate broadleaf and mixed forests Biome, in the Eastern United States. The ecoregion is located in the central and southern Appalachian Mountains, including the Ridge-and-Valley Appalachians and the Blue Ridge Mountains. It covers an area of about 61,500 square miles (159,000 km2) in: northeast Alabama and Georgia, northwest South Carolina, eastern Tennessee, western North Carolina, Virginia, Maryland, and central West Virginia and Pennsylvania; and small extensions into Kentucky, New Jersey, and New York.
Carya glabra, the pignut hickory, is a common, but not abundant species of hickory in the oak-hickory forest association in the Eastern United States and Canada. Other common names are pignut, sweet pignut, coast pignut hickory, smoothbark hickory, swamp hickory, and broom hickory. The pear-shaped nut ripens in September and October, has a sweet maple like smell, and is an important part of the diet of many wild animals. The wood is used for a variety of products, including fuel for home heating. Its leaves turn yellow in the Fall.
Spitler Woods State Natural Area is a 202.5-acre (81.9 ha) state park located adjacent to Mount Zion, Illinois. The state park is located within the Decatur, Illinois metropolitan area. The eastern two-thirds of the state park is a listed Illinois Nature Preserve noted for its old-growth forest grove of white oak and hickory. The park is managed by the Illinois Department of Natural Resources (IDNR).
Carya laciniosa, the shellbark hickory, in the Juglandaceae or walnut family is also called kingnut, big, bottom, thick, or western shellbark, attesting to some of its characteristics. It is a slow-growing, long-lived tree, hard to transplant because of its long taproot, and subject to insect damage. The nuts, largest of all hickory nuts, are sweet and edible. Wildlife and people harvest most of them; those remaining produce seedling trees readily. The wood is hard, heavy, strong, and very flexible, making it a favored wood for tool handles. A specimen tree has been reported in Missouri with 117 cm (46 in) diameter at breast height, 36.9 m tall, and a spread of 22.6 m.
Carya tomentosa, commonly known as mockernut hickory, mockernut, white hickory, whiteheart hickory, hognut, bullnut, is a species of tree in the walnut family Juglandaceae. The most abundant of the hickories, and common in the eastern half of the United States, it is long lived, sometimes reaching the age of 500 years. A straight-growing hickory, a high percentage of its wood is used for products where strength, hardness, and flexibility are needed. The wood makes excellent fuel wood, as well. The leaves turn yellow in Autumn.
Carya myristiciformis, the nutmeg hickory, a tree of the Juglandaceae or walnut family, also called swamp hickory or bitter water hickory, is found as small, possibly relict populations across the Southern United States and in northern Mexico on rich moist soils of higher bottom lands and stream banks. Little is known of the growth rate of nutmeg hickory. Logs and lumber are sold mixed with other hickories. The nuts are an oil-rich food for wildlife.
An oak forest is a plant community with a tree canopy dominated by oaks. In terms of canopy closure, oak forests contain the most closed canopy, compared to oak savannas and oak woodlands.
Quercus pagoda, the cherrybark oak, is one of the most highly valued red oaks in the southern United States. It is larger and better formed than southern red oak and commonly grows on more moist sites. Its strong wood and straight form make it an excellent timber tree. Many wildlife species use its acorns as food, and cherrybark oak makes a fine shade tree. Cherrybark oak was formerly considered to be a subspecies of southern red oak, Quercus falcata, subsp pagodifolia.
The Southeastern mixed forests are an ecoregion of the temperate broadleaf and mixed forest biome, in the lower portion of the Eastern United States.