Thylacinidae | |
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Thylacine | |
Non-Thylacinus thylacinids, including Nimbacinus (top right) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Infraclass: | Marsupialia |
Order: | Dasyuromorphia |
Family: | † Thylacinidae C.L. Bonaparte, 1838 |
Genera | |
Thylacinidae is an extinct family of carnivorous marsupials from the order Dasyuromorphia. The only species to survive into modern times was the thylacine (Thylacinus cynocephalus), which became extinct in 1936.
The consensus of authors prior to 1982 was that the thylacinid family were related to the Borhyaenidae, a group of South American predators, also extinct, that exhibited many similar characteristics of dentition. A review published in 1982 compared the skeletal structure of these groups, concluding the tarsal bones show greater affinity with the dasyurmorphs, strongly supporting the later theory that any dental similarities emerged independently via convergent evolution. [1] Thylacinidae is currently considered to be the most basal and earliest diverging member of Dasyuromorpha, estimated to have split from other dasyuromorphs around 42-36 million years ago. [2]
The earliest thylacinid, Badjcinus from the Late Oligocene around 28 million years ago, is estimated to have been around 1.7 to 3.1 kilograms (3.7 to 6.8 lb) in weight, comparable to a living tiger quoll. [3] Early thylacinids were unspecialised faunivores. The family exhibited its greatest diversity during the Miocene epoch, when there were several separate lineages of contemporaneous thylacinids, most of which were considerably smaller than the living thylacine. The genus Thylacinus emerged during the Early Miocene around 16 million years ago, and after the beginning of the Pliocene was the only living thylacinid genus. Over time members of the genus Thylacinus showed an increase in body and increased adaptation to hypercarnivory. [2]
Cladogram after Rovinsky et al. (2019): [2]
The thylacine, also commonly known as the Tasmanian tiger or Tasmanian wolf, is an extinct carnivorous marsupial that was native to the Australian mainland and the islands of Tasmania and New Guinea. The thylacine died out in New Guinea and mainland Australia around 3,600–3,200 years ago, prior to the arrival of Europeans, possibly because of the introduction of the dingo, whose earliest record dates to around the same time, but which never reached Tasmania. Prior to European settlement, around 5,000 remained in the wild on Tasmania. Beginning in the nineteenth century, they were perceived as a threat to the livestock of farmers and bounty hunting was introduced. The last known of its species died in 1936 at Hobart Zoo in Tasmania. The thylacine is widespread in popular culture and is a cultural icon in Australia.
Diprotodontia is the largest extant order of marsupials, with about 155 species, including the kangaroos, wallabies, possums, koala, wombats, and many others. Extinct diprotodonts include the hippopotamus-sized Diprotodon, and Thylacoleo, the so-called "marsupial lion".
Dasyuromorphia is an order comprising most of the Australian carnivorous marsupials, including quolls, dunnarts, the numbat, the Tasmanian devil, and the extinct thylacine. In Australia, the exceptions include the omnivorous bandicoots and the marsupial moles. Numerous South American species of marsupials are also carnivorous, as were some extinct members of the order Diprotodontia, including extinct kangaroos and thylacoleonids, and some members of the partially extinct clade Metatheria and all members of the extinct superorder Sparassodonta.
The order Peramelemorphia includes the bandicoots and bilbies. All members of the order are endemic to Australia-New Guinea and most have the characteristic bandicoot shape: a plump, arch-backed body with a long, delicately tapering snout, very large upright ears, relatively long, thin legs, and a thin tail. Their size varies from about 140 grams up to 4 kilograms, but most species are about one kilogram.
Thylacinus is a genus of extinct carnivorous marsupials in the family Thylacinidae. The only recent member was the thylacine, commonly also known as the Tasmanian tiger or Tasmanian wolf. The last known Tasmanian tiger was in the Beaumaris Zoo in Tasmania, eventually dying in 1936. The earliest known member of the genus, Thylacinus macknessi appeared during the Early Miocene, around 16 million years ago, and was smaller than the modern thylacine, with a body mass of about 6.7–9.0 kilograms (14.8–19.8 lb). Thylacinus represented the only extant genus of the family after the beginning of the Pliocene around 5 million years ago. Over time members of the genus saw an increase in body mass and a greater adaption to hypercarnivory in their dental morphology.
Australidelphia is the superorder that contains roughly three-quarters of all marsupials, including all those native to Australasia and a single species – the monito del monte – from South America. All other American marsupials are members of the Ameridelphia. Analysis of retrotransposon insertion sites in the nuclear DNA of a variety of marsupials has shown that the South American monito del monte's lineage is the most basal of the superorder.
Thylacoleo is an extinct genus of carnivorous marsupials that lived in Australia from the late Pliocene to the Late Pleistocene, often known as marsupial lions. They were the largest and last members of the family Thylacoleonidae, occupying the position of apex predator within Australian ecosystems. The largest and last species, Thylacoleo carnifex, approached the weight of a lioness. The estimated average weight for the species ranges from 101 to 130 kg.
Riversleigh World Heritage Area is Australia's most famous fossil location, recognised for the series of well preserved fossils deposited from the Late Oligocene to more recent geological periods. The fossiliferous limestone system is located near the Gregory River in the north-west of Queensland, an environment that was once a very wet rainforest that became more arid as the Gondwanan land masses separated and the Australian continent moved north. The approximately 100 square kilometres (39 sq mi) area has fossil remains of ancient mammals, birds, and reptiles of the Oligocene and Miocene ages, many of which were discovered and are only known from the Riversleigh area; the species that have occurred there are known as the Riversleigh fauna.
Thylacinus potens was the largest species of the family Thylacinidae, originally known from a single poorly preserved fossil discovered by Michael O. Woodburne in 1967 in a Late Miocene locality near Alice Springs, Northern Territory. It preceded the most recent species of thylacine by 4–6 million years, and was 5% bigger, was more robust and had a shorter, broader skull. Its size is estimated to be similar to that of a grey wolf; the head and body together were around 5 feet long, and its teeth were less adapted for shearing compared to those of the now-extinct thylacine.
Badjcinus is an extinct thylacinid marsupial. It is the earliest and most primitive known thylacinid, living 23 to 28 million years ago in the late Oligocene.
The genus Nimbacinus contains two species of carnivorous, quadrupedal marsupials in Australia both of which are extinct:
Ektopodon is an extinct genus of marsupial, and is the type genus of the family Ektopodontidae which occurred in forested environments in South Australia, Queensland and Victoria. The last species of this group went extinct in the early Pleistocene. Its body mass was estimated around 1300 grams. Scientists believe that ektopodontids were highly specialised seed-eating possums.
Thylacinus megiriani lived during the late Miocene, 8 million years ago; the area T. megiriani inhabited in the Northern Territory was covered in forest with a permanent supply of water.
Mutpuracinus archibaldi is an extinct carnivorous, quadrupedal marsupial that lived during the middle Miocene and is the smallest known thylacinid at approximately 1.1 kilograms, the size of a quoll, though, more closely related to the recently extinct thylacine.
Ngamalacinus timmulvaneyi lived during the early Miocene and has been found in Riversleigh.
Wabulacinus ridei lived during the early Miocene in Riversleigh. It is named after David Ride, who made the first revision of thylacinid fossils. The material was found in system C of the Camel Spurtum assembledge.
The Macropodidae are an extant family of marsupial with the distinction of the ability to move bipedally on the hind legs, sometimes by jumping, as well as quadrupedally. They are herbivores, but some fossil genera like Ekaltadeta are hypothesised to have been carnivores. The taxonomic affiliations within the family and with other groups of marsupials is still in flux.
Riversleigh fauna is the collective term for any species of animal identified in fossil sites located in the Riversleigh World Heritage Area.
Thylacinus yorkellus is a fossil species of carnivorous marsupial, a sister species of the recently extinct Thylacinus cynocephalus, the Tasmanian tiger, both of which existed on mainland Australia.
Dorcopsoides is a genus of extinct species of kangaroo from the Pliocene of Australia.