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Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Infraorder: Araneomorphae
Family: Udubidae
Griswold & Polotow, 2015
4 genera, 19 species

Udubidae is a family of araneomorph spiders, most of whose members were formerly placed in the family Zorocratidae, which is no longer accepted. [1]



A study investigating the phylogenetic relationships of lycosoid spiders concluded that the genera formerly placed in the family Zorocratidae fell into two groups. The largest group formed the sister clade to the "grate-shaped tapetum clade" (see the cladogram below). The type genus, Zorocrates , grouped with Zoropsis inside the grate-shaped tapetum clade. [1] Some earlier studies had also cast doubt on the monophyly of the Zorocratidae. A 2003 study found that Raecius, Uduba, and Zorodictyna formed a clade somewhat separated from Zorocrates. [2] A 2014 study including Zorocrates and Raecius did not find that they grouped together. [3]

Moving the type genus to a different family meant that a new family name was needed for the remaining members. Griswold and Polotow proposed "Udubidae", with the type genus Uduba. [1] As of November 2015, the World Spider Catalog accepted this family. [4] A summary phylogeny is shown below. [1]

RTA clade

basal groups


Oval calamistrum clade

Udubidae (including Raecius, Uduba & Zorodictyna)

Grateshaped tapetum clade

Zoropsidae (including Zorocrates)


(Shading marks genera formerly placed in Zorocratidae.)


As of April 2019, the World Spider Catalog accepts the following genera: [4]

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Nephilinae Spider family

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RTA clade

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Zorodictyna is a genus of spiders in the family Udubidae native to Madagascar. It has been described as an intermediate genus between Zoropsidae and Dictynidae, though it is now placed in Udubidae. This genus was originally placed in the family Zoropsidae, but it has been reassigned several times since. In 1967, Lehtinen moved it to Miturgidae. In 1999, it was moved back to Zoropsidae, and in 2015, it was moved to Udubidae.

Campostichomma is a genus of spiders in the family Udubidae native to Sri Lanka. Many of its species were moved to either Griswoldia or Devendra. This genus was originally placed in the family Agelenidae. It was moved to Miturgidae in 1967, to Zoropsidae in 1999, then to Udubidae in 2015.

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Vulsor is a genus of araneomorph spiders in the family Viridasiidae, first described by Eugène Simon in 1889. Originally placed with the wandering spiders, it was moved to the Viridasiidae in 2015.

Manzuma is a genus of spiders in the jumping spider family Salticidae native to Africa.


  1. 1 2 3 4 Polotow, Daniele; Carmichael, Anthea & Griswold, Charles E. (2015). "Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae)". Invertebrate Systematics. 29 (2): 124–163. doi: 10.1071/IS14041 .
  2. Davila, Diana Silva (2003), "Higher-level relationships of the spider family Ctenidae (Araneae, Ctenoidea)", Bulletin of the American Museum of Natural History (274), retrieved 2015-10-17
  3. Ramírez, M. (2014), The morphology and phylogeny of Dionychan spiders (Araneae: Araneomorphae), Bulletin of the American Museum of Natural History, retrieved 2015-10-31, p. 289
  4. 1 2 "Family: Udubidae Griswold & Polotow, 2015". World Spider Catalog. Natural History Museum Bern. Retrieved 2019-04-26.