funnel-web spiders Temporal range: | |
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Hexathele hochstetteri , female | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Class: | Arachnida |
Order: | Araneae |
Infraorder: | Mygalomorphae |
Clade: | Avicularioidea |
Family: | Hexathelidae Simon, 1892 |
Diversity | |
7 genera, 47 species |
Hexathelidae is a family of mygalomorph spiders. [1] It is one of a number of families and genera of spiders known as tunnelweb or funnel-web spiders. In 2018, the family was substantially reduced in size by genera being moved to three separate families: Atracidae, Macrothelidae and Porrhothelidae.
These spiders are medium to large in size, with body lengths ranging from 1 to 5 cm (0.5 to 2 in). The body is typically three times longer than it is wide. They are darkly colored, ranging from black to brown, with a glossy carapace covering the front part of the body. Like the related diplurid spiders, the hexathelids have generally long spinnerets.
Their moderately long posterior spinnerets and other features make the Hexathelidae appear similar to the Dipluridae, and were considered a subfamily of the latter until 1980. [1]
Like other Mygalomorphae (also called the Orthognatha, an infraorder of spiders which includes the true tarantulas), these spiders have fangs which point straight down and do not cross each other (see also Araneomorphae). They have ample venom glands that lie entirely within their chelicerae. Their chelicerae and fangs are large and powerful.
A molecular phylogenetic study in 2018 showed that Hexathelidae, as then circumscribed, was not monophyletic and hence split off genera into the new families Atracidae, Macrothelidae and Porrhothelidae. The following cladogram shows the relationships found between these families (former Hexathelidae families marked with boxes). [2]
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As of April 2019 [update] , the World Spider Catalog accepts the following genera: [3]
As of May 2018 [update] , some genera formerly placed in Hexathelidae have been moved to other families:
Five of the seven genera are found in Australia and New Zealand. Two genera are from Argentina and Chile in South America. [7]
Hexathelids typically live in burrows, which are constructed in the ground or in tree hollows. An elaborately constructed burrow entrance is common. These spiders construct a funnel-shaped web and lurk for prey in the small end of the funnel. They frequently search for a place to nest under human dwellings, or under nearby rocks, logs, or other similar objects. They are most active at night. Some build in rain forests, both in the soil and in hollows on trees; others build entirely in sand, e.g., Fraser Island, southeastern Queensland. In Australia, these spiders tend to prefer cooler climates, hence they are found primarily in rain forests. [8]
The Mygalomorphae, or mygalomorphs, are an infraorder of spiders, and comprise one of three major groups of living spiders with over 3,000 species, found on all continents except Antarctica. Many members are known as trapdoor spiders due to their creation of trapdoors over their burrows. Other prominent groups include Australian funnel web spiders and tarantulas, with the latter accounting for around one third of all mygalomorphs.
The family Dipluridae, known as curtain-web spiders are a group of spiders in the infraorder Mygalomorphae, that have two pairs of booklungs, and chelicerae (fangs) that move up and down in a stabbing motion. A number of genera, including that of the Sydney funnel-web spider (Atrax), used to be classified in this family but have now been moved to Atracidae.
Atracidae is a family of mygalomorph spiders, commonly known as Australian funnel-web spiders or atracids. It has been included as a subfamily of the Hexathelidae, but is now recognised as a separate family. All members of the family are native to Australia. Atracidae consists of three genera: Atrax, Hadronyche, and Illawarra, comprising 35 species. Some members of the family produce venom that is dangerous to humans, and bites by spiders of six of the species have caused severe injuries to victims. The bites of the Sydney funnel-web spider and northern tree-dwelling funnel-web spider are potentially deadly, but no fatalities have occurred since the introduction of modern first-aid techniques and antivenom.
The Mesothelae are a suborder of spiders. As of April 2024, two extant families were accepted by the World Spider Catalog, Liphistiidae and Heptathelidae. Alternatively, the Heptathelidae can be treated as a subfamily of a more broadly circumscribed Liphistiidae. There are also a number of extinct families.
The family Cyrtaucheniidae, known as wafer-lid trapdoor spiders, are a widespread family of Mygalomorphae spiders.
Atrax is a genus of venomous Australian funnel web spiders that was first described by O. Pickard-Cambridge in 1877 from the type species Atrax robustus. As of May 2019 it contains only three species: A. robustus, A. sutherlandi, and A. yorkmainorum. Originally placed with the curtain web spiders, it was moved to the Hexathelidae in 1980, then to the Australian funnel-web spiders in 2018.
Hadronyche is a genus of venomous Australian funnel-web spiders that was first described by L. Koch in 1873. Originally placed with the curtain web spiders, it was moved to the Hexathelidae in 1980, then to the Atracidae in 2018.
Microstigmatidae is a small family of spiders with about 38 described species in eleven genera. They are small ground-dwelling and free-living spiders that make little use of silk.
Nemesiidae is a family of mygalomorph spiders first described by Eugène Simon in 1889, and raised to family status in 1985. Before becoming its own family, it was considered part of "Dipluridae". The family is sometimes referred to as wishbone spiders due to the shape of their burrows.
Barychelidae, also known as brushed trapdoor spiders, is a spider family with about 300 species in 39 genera.
Macrothele is a genus of mygalomorph spiders in the family Macrothelidae, and was first described by A. Ausserer in 1871. Most of the species in the genus occur in Asia, from India to Japan, and Java, with five found in Africa, and two in Europe. The name is derived from Ancient Greek μακρός ("makro-"), meaning "big", and θηλή ("thele"), referring to the spinnerets.
Porrhothele is a genus of mygalomorph spiders endemic to New Zealand. They are the only members of the family Porrhothelidae. They were first described by Eugène Louis Simon in 1892. Originally placed with the curtain web spiders, it was moved to the Hexathelidae in 1980, they were placed in their own family in 2018.
Hexathele is a genus of tunnelweb spiders endemic to New Zealand that was first described by Anton Ausserer in 1871, though most others have been described by Raymond Robert Forster. Originally placed with the curtain web spiders, it was moved to the Hexathelidae in 1980.
Scotinoecus is a genus of South American funnel-web spiders that was first described by Eugène Louis Simon in 1892. Originally placed with the curtain web spiders, it was moved to the Hexathelidae in 1980.
Entypesa is a genus of African mygalomorph spiders in the family Entypesidae. It was first described by Eugène Louis Simon in 1902. Originally placed with the curtain-web spiders, it was transferred to the funnel-web trapdoor spiders in 1985, then to the Entypesidae in 2020. It is a senior synonym of Pseudohermacha.
Macrothelidae is a family of mygalomorph spiders, split off from the family Hexathelidae in 2018. It contains three genera, Macrothele,Vacrothele and the extinct Promacrothele.
Avicularioidea is a clade of mygalomorph spiders, one of the two main clades into which mygalomorphs are divided. It has been treated at the rank of superfamily.
Pycnothelidae is a family of mygalomorph spiders first described in 1917. It was downgraded to a subfamily of the funnel-web trapdoor spiders in 1985, but returned to family status in 2020.
Euagridae is a family of mygalomorph spiders. The group was first described as a tribe in 1979 by Robert Raven, who in 1985 elevated it to a subfamily. In 2020, Opatova et al. elevated it further to a family.
Afropesa is a genus of southern African mygalomorph spiders in the family Entypesidae. It was first described by S. L. Zonstein and D. Ríos-Tamayo in 2021, and it has only been found in South Africa. As of November 2021 it contains only three species: A. gauteng, A. schoutedeni, and A. schwendingeri. The type species was originally described under the name "Entypesa schoutedeni.