Tree trunk spider Temporal range: | |
---|---|
Male Hersilia okinawaensis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Class: | Arachnida |
Order: | Araneae |
Infraorder: | Araneomorphae |
Family: | Hersiliidae Thorell, 1869 |
Diversity | |
16 genera, 206 species | |
Hersiliidae is a tropical and subtropical family of spiders first described by Tamerlan Thorell in 1869, [1] which are commonly known as tree trunk spiders. They have two prominent spinnerets that are almost as long as their abdomen, earning them another nickname, the "two-tailed spiders". They range in size from 10 to 18 mm (0.4 to 0.7 in) long. Rather than using a web that captures prey directly, they lay a light coating of threads over an area of tree bark and wait for an insect to stray onto the patch. When this happens, they encircle their spinnerets around their prey while casting silk on it. When the insect is immobilized, they can bite it through the shroud.
Hersiliidae is an entelegyne family (characterized primarily by the nature of the female genital system), and together with the family Oecobiidae traditionally formed the superfamily Oecobioidea. [2] The family consists of about 206 species divided into sixteen genera. [3] It has a global distribution in tropical and subtropical regions, with only a few species being found north of the 40°N parallel. All members are ecribellate (lack the cribella or perforated plates which produce multiple, exceptionally fine strands of silk) and are recognizable by the pair of exceptionally long spinnerets set at the tip of the abdomen. [2] They have eight eyes, set in two curved rows. They are small to medium-sized spiders and are active day and night. They are very well camouflaged when stationary on the trunk of a tree and aligned with the bark markings. [4]
As of April 2019 [update] , the World Spider Catalog accepts the following genera: [3]
The family Dipluridae, known as curtain-web spiders are a group of spiders in the infraorder Mygalomorphae, that have two pairs of booklungs, and chelicerae (fangs) that move up and down in a stabbing motion. A number of genera, including that of the Sydney funnel-web spider (Atrax), used to be classified in this family but have now been moved to Atracidae.
Ground spiders comprise Gnaphosidae, the seventh largest spider family with over 2,000 described species in over 100 genera distributed worldwide. There are 105 species known to central Europe, and common genera include Gnaphosa, Drassodes, Micaria, Cesonia, Zelotes and many others. They are closely related to Clubionidae. At present, no ground spiders are known to be seriously venomous to humans.
Oonopidae, also known as goblin spiders, is a family of spiders consisting of over 1,600 described species in about 113 genera worldwide, with total species diversity estimated at 2000 to 2500 species. The type genus of the family is OonopsKeyserling, 1835.
Long-jawed orb weavers or long jawed spiders (Tetragnathidae) are a family of araneomorph spiders first described by Anton Menge in 1866. They have elongated bodies, legs, and chelicerae, and build small orb webs with an open hub with few, wide-set radii and spirals with no signal line or retreat. Some species are often found in long vegetation near water.
Scaffold web spiders or cave cobweb spiders (Nesticidae) are a family of araneomorph spiders closely allied with tangle-web spiders (Theridiidae). Like the Theridiidae, these spiders have a comb of serrated bristles on the hind tarsi that are used to pull silk bands from the spinnerets. Nesticidae contains 16 genera and about 300 species, many of which are associated with caves or overhangs. The genus Nesticus is the type for the family and is found throughout the world. The related Eidmannella has speciated considerably in Texas caves and includes some extremely localized species that are considered threatened. One species, Eidmannella pallida, is found in caves and under overhangs, but also in agricultural fields and other habitats away from such restricted areas. The genus Carpathonesticus is found in central Eurasia.
Theridiosomatidae, commonly known as Ray Spiders, are a family of araneomorph spiders first described by Eugène Simon in 1881. The family includes 137 species divided between 20 genera. They are most recognizable for their construction of cone-shaped webs.
Prodidomidae is a family of spider, sometimes called long-spinneret ground spiders. It was formerly regarded as a subfamily of Gnaphosidae, but was raised to a family in 2022.
Hersilia, also known as long-spinnereted bark spiders and two-tailed spiders, is a genus of tree trunk spiders that was first described by Jean Victoire Audouin in 1826. Their nicknames are a reference to their greatly enlarged spinnerets.
Scytodes is a genus of spitting spiders that occur all around the world. The most widely distributed species is Scytodes thoracica, which originally had a palearctic distribution, but has been introduced to North America, Argentina, India, Australia, and New Zealand. The genus was first described by Pierre André Latreille in 1804. Spitting spiders have pale yellow bodies with black spots on their cephalothorax, and legs that are characterized by black bands.
Tamopsis is a genus of tree trunk spiders that was first described by B. Baehr & M. Baehr in 1987. Like other members of the family, they may be called two-tailed spiders, referring to two elongated spinnerets. The name is derived from the genus Tama and the Ancient Greek ὄψις, meaning "resembling".
Neotama is a genus of tree trunk spiders that was first described by M. Baehr & B. Baehr in 1993.
Duninia is a genus of Asian tree trunk spiders that was first described by Yuri M. Marusik & Victor R. Fet in 2009.