Vespula vulgaris, known as the common wasp, is a species found in regions that include the United Kingdom, Ireland, Germany, India, China, New Zealand[1] and Australia. It is sometimes known in English as the European wasp, but the same name is used for the species Vespula germanica or German wasp. In 2010, the ostensible Vespula vulgaris wasps in North America were found to be a different species, Vespula alascensis.[2]
Vespula vulgaris is a eusocialvespid that builds its tan paper nest in or on a structure capable of supporting it. A founding queen searches for a hollow tree, wall cavity, rock crevice, or even a hole made by other animals to build a nest. One colony cycle lasts for about 6–11 months and each colony cycle produces around 3,000–8,000 larvae.[citation needed]
The extraordinary adaptation skills of V. vulgaris enable it to live in a wide range of habitats, from very humid areas to artificial environments such as gardens and human structures. This species, along with other wasp species such as V. germanica, has impacted the ecosystem, especially those in New Zealand and Australia, where they were imported by humans, and frequently cause damage to fruit crops and endanger humans.[3]
Taxonomy
The name Vespula vulgaris comes from the origin of the word vulgaris, which means "common" in Latin, therefore giving the name "common wasp". This species has many synonyms such as "common wasp", "European wasp", Paravespula vulgaris, or "common yellow jacket".[4] However, a study from 2010 revealed that Vespula vulgaris and the American common yellow jacket are actually two different species, the latter now known as Vespula alascensis.[2] It is closely related to another wasp species, Vespula austriaca, and is considered a sister taxon.[5]
In 2009, the species Vespula rufosignataEck, 1998 was synonymised with Vespula vulgaris.[6]
Common wasps are colloquially called "jaspers" in some English regions (Dorset, Lincolnshire, and elsewhere in the English Midlands); whether this comes from the Latin vespa or from the abdomen resembling the striped mineral jasper, is not clear.[7]
Description and identification
Worker in flight
Adult workers of V. vulgaris measure about 12–17mm (0.5–0.7in) from head to tip of abdomen,[8] and weigh 84.1 ± 19.0mg,[9] whereas the queen is about 20mm (0.8in) long. It has aposematic colors of black and yellow; yellow pronotal bands which are almost parallel to each other and black dots and rings on its abdomen. The queens and workers appear very similar to Vespula germanica, except when they are seen head on, as the V. vulgaris face lacks the three black dots of V. germanica. Instead, each has only one black mark on its clypeus, which is usually anchor or dagger-shaped. This applies to queens and workers only. In addition, identification of this species may be difficult because the black mark on its clypeus can sometimes appear broken, making it again look similar to V. germanica.[7] It is prudent to use multiple identifying characteristics and if in any doubt to consult experts.
Still more difficult to distinguish between species are the males. Almost undetectable with the naked eye, the only confident identification of V. vulgaris males is to seek the distinct aedeagus tip shapes and lateral processes of their genitalia.[3]
Distribution and habitat
V. vulgaris is a Palearctic species.[2] It has been discovered in a wide range of countries, including the United Kingdom, Germany, India, and China. It is invasive in New Zealand and Australia, and South America.[7] Until 2010, it was thought to be in North America as well, but molecular and morphological data showed that specimens identified as V. vulgaris there were Vespula alascensis, which had previously been considered a taxonomic synonym, but is now considered a separate species.[2]
V. vulgaris has high adaptation skills for environments. It flourishes in most types of habitats, including prairie, grassland, natural and planted forests, shrub lands, and even in urban zones such as gardens, orchards, and buildings. However, it does require the temperature to be moderately warm, because its foraging activity is temperature-dependent (above 2°C [36°F]).[4]
Northern Hemisphere colony cycle
Five distinguishable stages occur in V. vulgaris colony cycle in the temperate parts of the Northern Hemisphere. Each lasts for similar periods, around 30–35 days. The colony starts in April as the queen begins the foundation of the nest and degenerates around October when the temperature drops and the queen dies. The size of the colony ranges from 3000 larvae up to a maximum of 8000 larvae, with the queen producing around 200–300 eggs per day for 24 days.[10]
Stage 1 – solitary stage: the queen builds the nest, provisions the cells and rears the first brood of workers.
Stage 2 – rapid increase: the workers replace the queen as the foraging force. The queen is now a nurse and egg producer. There is very rapid buildup of worker population.
Stage 3 – slow increase: the rate of population growth is not as rapid as in stage 2.
Stage 4 – climax of colony: worker population no longer increases. Cells built are all queen cells. Males emerge, as do unmated queens, and foraging rate per worker increases. Drones and queens mate, and the queens hibernate to emerge in the spring, when the cycle repeats with stage 1.
Stage 5 – decline: probably caused by death or sickness of queen. Colony cohesion breaks down, cannibalism sets in, and the foraging becomes erratic.[10]
Brood production
The general incubation period is five days, but it can occasionally be seven, due to cannibalism, scavenging or replacement. The larval period can depend on the trophic condition inside and outside the nest. If there are abundant resources of food being brought into the nest, the larval period is short, but if the resource is short, the period can lengthen. The first adult emerges after around 23 days of hatching and the queen-pupated brood is completed in about a month (30 days).[10]
The first brood of workers are all of normal size, but the next generation of them is generally smaller, due to a decrease in the queen's foraging activity after producing the first batch of larvae. As the queen's foraging decreases, the second group of workers does not get so much food, often starves, and so spends more time in the larval stage.[10]
Nest foundation by the queen
After coming out of hibernation in the spring, the queen forages for flowers or shrubs and begins looking for nesting sites. She flies low on the ground, searching for any round, dark object or depression. If it is a hole, she flies in to inspect the details if it is suitable and if not moves on to the next hole. What constitutes a "perfect" nest is yet to be discovered, but it is mainly speculated that the queen's physiological state is the main deciding factor of the time and the place of nesting. When she has found a suitable site for nesting, she performs dance-like wing movements which allow her to locate the site even after she has left the area.[11]
After she has fixed the position of the nest, she begins to build it. She repeats the procedure of bringing in a pulp, attaching it to the nest to form a spindle and flying back out to bring in more pulp. During this process, the queen clings to the nest with her two posterior pairs of legs and attaches the pulp using her anterior legs and mouth parts.[10] She builds 20–30 cells before initial egg-laying, fashioning a petiole and producing a single cell at the end of it. Six further cells are then added around this to produce the characteristic hexagonal shape of the nest cells. One egg is laid in each cell, and as it hatches, each larva holds itself in the vertical cell by pressing its body against the sides.[8]
When the nest is completed, the queen is replaced by the workers as the foraging force and instead is now concerned only with nursing and egg producing. Her ovaries develop, her abdomen becomes distended with eggs, and she loses the ability to fly.[10]
After the founding phase has been completed, the colony encounters a change where the workers begin to build queen cells. Once the workers start building on the queen cells, no more worker cells are built, but those that still have brood growing in them are retained. The majority of the food resource brought in by the workers is fed to the queen larvae, known as gynes and the lack of feeding for other larvae causes the prolongation of their larval periods.[11] To ensure that only the queen's eggs are reared to adulthood, female workers remove worker-laid eggs in a process known as worker policing.[citation needed]
When the queen has completed her job of producing daughter queen larvae, she dies, leaving a crop of virgin queens which will leave the nest, mate, hibernate, and reproduce in the following spring. After the queen's death, the coordination of the colony breaks down and the workers begin to lay eggs. The rate and amount of foraging decrease drastically after the queen's death, so it is unable to support all the workers and their brood. This is when cannibalism occurs among the workers and furthermore, the workers tear off their cells and carry them out of the nest, dropping the temperature of the entire nest. Once this stage is reached, the remaining workers die of cold or starvation.[12]
Nest
Nest composed of chewed wood fibers
A V. vulgaris nest is made from chewed wood fibers mixed with worker saliva. It is generally made of brittle tan coloured paper. It has open cells and a cylindrical column known as a petiole attaching the nest to the substrate. The wasps produce a chemical which repels ants and secrete it around the base of the petiole, to avoid ant predation.[13]
Many variations are seen in the characteristics of the nests within the species. Aerial nests and nests that are very close to the surface of the ground have much thicker envelopes than those under the ground. The wasps build thick envelopes to prevent heat from escaping. Also, smaller nests have thicker envelopes than larger nests. This is mainly due to the fact that the amount of heat produced is proportional to the volume of the nest. The larger a nest is the better it will be in conserving the nest warmth. The optimal temperature of the nest is around 32°C (90°F). However, when the temperature rises above the optimal temperature (during hot days), the workers use their wings as fans to cool the nest down.[14]
V. vulgaris wasps recognize their nests by making pheromone trails from the entrance of their nest to the site of foraging. Those "footprints" are not colony specific, but species specific. Although many other insects such as ants produce such pheromone trails as well, Vespula vulgaris generate pheromone either by special abdominal glands[15] or from the gut.[16]
Although the two species Vespula vulgaris and Vespula germanica have extremely similar biological features, characteristics of their nests are distinctive.[17]Vespula germanica nests generally survive the winter, while those of Vespula vulgaris do not. This difference is due mainly to the variation in prey. Vespula vulgaris food sources are greatly affected by temperature, while those of Vespula germanica are not, providing them with a higher chance of survival even during the winter.[17] Because many Vespula germanica are able to overwinter and most Vespula vulgaris are not, except for the queen, this affects the nest size of the two species. Vespula germanica generally have slightly bigger nest size (3500–9000 larvae in one colony cycle) than Vespula vulgaris (3000–8000 larvae in one colony cycle).[18]
Behaviors
Dance-like wing movement of queen
When the queen has found the appropriate nesting site, she orients in a similar way as the workers and flies back out of the hole. She flies frontwards and backwards in front of the nest with a slow, hovering flight and repeats this movement until its distance is further away from the hole. When the arc of the flight increases up to about 6ft (183cm), the pathway makes a figure of a sideways "8" and when the queen is about 60ft (18 m) away from the nest site, she then flies in a straight line. This is how the queen Vespula vulgaris marks the land, in respect to the landmarks around the nesting site.[10]
Defensive behaviors
Only wasps leaving the nest notice a disturbance and defend or fight off an intruder. The ones returning to the nest do not detect any disturbance and try to enter the nest. The workers that detect danger show a certain gesture – they rise onto the tips of their tarsi, put forward their heads, turn down their abdomens and constantly vibrate their wings in high frequencies and short beats. This behavior signals other workers to fly to the entrance of the nest and defend. However, if the nest is disturbed enough times, the workers stop defending the nest and instead grow tolerant to such attacks.[10] However, when they detect life-threatening level of danger, the Vespula vulgaris workers will vigorously defend their nest. Unlike honey bees, which die after stinging, Vespula vulgaris can sting multiple times. This makes its sting viable for personal defense when away from the colony, and the common wasp is therefore more apt to sting. However, it will usually not sting without being provoked by sudden movement or other violent behavior.[citation needed]
Research indicates the wasps use odor to identify and attack rival wasps from other colonies, and nest odor frequently changes.[14]Vespula vulgaris wasps have been observed aggressively competing with honey bees for the honeydew secreted by the scale insect Ultracoelostoma brittini in New Zealand's South Island black beech forests.[19]
Over-wintering behaviors
After mating, the queen overwinters in a hole or other sheltered location, sometimes in buildings. Wasp nests are not reused from one year to the next, but in rare instances wasps have been seen to re-nest in the footprint of a removed nest or even begin building a new nest within an old nest. In the mild climate of New Zealand and Australia, a few of the colonies may survive the winter, although this is much more common with the German wasp.[13][19]
The foraging activity of Vespula vulgaris is dependent on the light and temperature of the surrounding area. Generally, if the temperature of the surrounding area falls below 2°C, the wasps will not proceed with foraging even if the temperature of the nest itself is optimal. When the temperature is above 2°C, light intensity comes in to play a role. If the light intensity is high enough, the wasps will fly off to forage. However, there is a wide variance within the species, and even within individuals what that threshold light intensity would be – i. e. when would be the best light intensity for a wasp to go out to forage.[21]
The foraging activities of Vespula vulgaris are generally highest in early mornings. There are three known reasons for such behavior. First is to satisfy the larval requirements. The workers fly off early in the morning to forage, as soon as their optimal light intensity is reached. They want to feed the larvae which have been starving during the night as soon as possible to reduce their chance of dying. The second reason is to maximize fluid foraging. Because dews and nectars are most abundant early in the morning and are the best sources of fluids, the workers fly off early in the morning before the competition becomes severe. Last is the necessity to release excretory products such as trophallactic secretions from the larvae.[10]
Generally, wasps are unable to predict heavy rainfalls. They continue their foraging activities even in the rain, but cease when the rain becomes torrential. When the rainfall becomes extremely heavy near the nesting site, the workers rapidly return to their nests. Those whose foraging activities were interrupted by heavy rainfall become reluctant to forage again in the future, even when the weather is dry and calm. Those who continue to forage do so mainly for fluid.[citation needed]
The average life span of an adult Vespula vulgaris is known to be around two weeks (14 days). There was a negative correlation found between the age of the worker Vespula vulgaris and their respective foraging speeds, and a positive correlation between their age and the amount of time each worker spent in the nest between each foraging trip. The foraging speed decreased as the workers aged and the time a worker spent in the nest between foraging trips increased as they got older. Wasps that are over 30 days, in fact, generally cease from foraging trips and spend all of their time guarding the entrance of the nest.[10]
Diet
Similarly to other Vespulas, Vespula vulgaris feed on animal preys such as caterpillars to feed their developing larvae and carbohydrates, such as nectar and sweet fruits, to satisfy their own energy requirements.[22] Their usual food sources are freshly killed insects such as Hymenoptera, lepidopteran larvae and Diptera and spiders.[23] Common wasps will also attempt to invade honey bees' nests to steal their honey.[citation needed]
Although the types of prey Vespula vulgaris and Vespula germanica forage are almost the same, that of Vespula germanica are generally 2–3.5 times bigger and heavier than that of Vespula vulgaris. This is mainly due to the size difference of the two species. Because the Vespula germanica foragers are bigger in morphology than those of Vespula vulgaris, and they both transport the prey by carrying them, it would be advantageous for the wasp to be bigger to be able to hold larger prey.[22]
Caste system
In the early stage of the colony founding, the queen Vespula vulgaris does most of the work, both building and foraging. After completing the nest, she diversifies her foraging to include not just pulp, but also fluids and insect flesh. The labor divides as the first batch of larvae is hatched. However, during the very beginning of the hatching, the workers do not take part of any labor. Rather, they remain motionless with their heads inside a cell. Workers actually start foraging actively after seven days. They both forage and take care of the broods by feeding the larvae, breaking down the insect flesh, dividing fluids collected by the foragers, removing trophallactic secretion from the larvae, and fixing the nests.[10]
Queen
One possible factor that determines a female's social caste is known to be its nest position. The location of each cell is directly related to the amount of food a larva can receive, and the queen cell, being located at the very bottom of the nest, encounters the foragers before any other cell. Therefore, it gets most of the food and naturally, yields the biggest female – the queen.[10]
As mentioned above, the cell location can alter the size of the larva and eventually determine which female Vespula vulgaris is going to be queen. However, when several candidates of queens arise, there is competition among them. Certain variations among them rank the hierarchy of the queens and thus select the ultimate queen. Although the precise causes of the variations in the queens are still unknown, one possible factor is the amount of fat stored in the queen's body. Smaller developing queens have less fat stored, and thus have relatively higher mortality rates in winter than larger queens. The bigger the size and larger the amount of fat stored, the "higher" the queen is considered in quality, and she is able to out-compete other queens in finding a fit mate. She will then have an advantage in establishing a nest in the following spring.[12]
In order for the Vespula vulgaris queen to dominate all the workers – usually numbering more than 3000 in a colony –she possesses a special chemical that sends signals of dominance. The workers were discovered to regularly lick the queen while feeding her, and the air-borne pheromone from the queen's body alerts those workers of her dominance.[10]
Pest status
Along with the invasive German wasp (Vespula germanica) and three species of Polistes, the common wasp is considered a pest species in New Zealand. It predates native insects and competes with endemic species for food, such as insects and honeydew.[24][25][1] In some South Island beech forests it has been calculated that wasp densities are higher than anywhere else in the world,[13] and that the total weight of common wasps may exceed that of all birds.[8][1]
Predators
Vespula vulgaris is subject to predation by the honey buzzard, which excavates the nests to obtain the larvae. The hoverfly Volucella pellucens and some of its relatives lay their eggs in a wasp nest, and their larvae feed on the wasps' young and dead adults. Larvae of the ripiphorid beetle Metoecus paradoxus are a parasitoid of V. vulgaris larvae.[26]Robber fly and spiders are other predators of this and many other species.[12] A species of parasitic mite, Varroa destructor jacobsoni, was found on larvae of this species in Poland in 1988.[8]
In New Zealand a 2010 study found that an as yet unidentified predator/s is responsible for local extinction of Vespula (both vulgaris and germanica) on many offshore islands 5–7 years following eradication of introduced rodents. Studies including 36 islands have not identified the agent responsible. However, a 2020 study of 36 islands found, "no evidence for a link between rat presence and wasp abundance on islands", but that wasps were reduced by tree canopy cover by species not attracting honeydew-secreting scale insects.[27]
Yellowjacket or yellowjacket is the common name in North America for predatory social wasps of the genera Vespula and Dolichovespula. Members of these genera are known simply as "Wasps" in other English-speaking countries. Most of these are black and yellow like the eastern yellowjacket and the aerial yellowjacket ; some are black and white like the bald-faced hornet. Some have an abdomen with a red background color instead of black. They can be identified by their distinctive markings, their occurrence only in colonies, and a characteristic, rapid, side-to-side flight pattern prior to landing. All females are capable of stinging. Yellowjackets are important predators of pest insects.
Paper wasps are vespid wasps and typically refers to members of the vespid subfamily Polistinae, though it often colloquially includes members of the subfamilies Vespinae and Stenogastrinae, discussed elsewhere, which also make nests out of paper. Paper wasp nests are characterized by open combs with down pointing cells. Some types of paper wasps are also sometimes called umbrella wasps, due to the distinctive design of their nests.
The median wasp is a species of social wasp of the family Vespidae found throughout Europe and Asia. It builds aerial paper nests often in shrubs or trees, and occasionally under the eaves of buildings. It is most common to see this wasp between May and October during its 3.3 month colony cycle. Behaviours of this wasp include nest defense, curling which is believed to function in brood incubation, and gastral vibration which is involved in larval feeding. The median wasp has a halplodiploid sex determination system that results in a high level of relatedness within the colony. This species is not usually aggressive but will sting if they feel their nest is threatened. Most foraging in the nest is done by the workers once the first ones reach adulthood. These workers forage for insects, nectar, and wood for nest construction in temperatures as low as 7 °C (45 °F). The median wasp is known to be occasionally affected by the fungus Cordyceps sphecocephala and the Cricket paralysis virus.
Volucella pellucens, the pellucid fly, is a hoverfly.
Vespula germanica, the European wasp, German wasp, or German yellowjacket is a species of wasp found in much of the Northern Hemisphere, native to Europe, Northern Africa, and temperate Asia. It has spread and become well-established in many other places, including North America, South America, Australia, South Africa, and New Zealand. German wasps are part of the family Vespidae and are sometimes mistakenly referred to as paper wasps because they build grey paper nests, although strictly speaking, paper wasps are part of the subfamily Polistinae. In North America, they are also known as yellowjackets.
Polistes humilis, known as the Australian paper wasp, is a species of wasp in the family Vespidae that is found throughout Australia and which has been introduced to northern New Zealand. These paper wasps can be identified by their long thin legs and banded yellow and black coloring. They have been known to re-utilize old nests. While the species does not exhibit morphological class differences, there are distinct behavioral differences between queens and workers. In addition, the species is eusocial and benefits from relatedness between individuals. They are known for delivering a painful sting, especially when their nest is disturbed, a behavior that has been developed as a nest defense mechanism. While wasps are often viewed negatively, they play an important pollination role for many plants.
Vespula pensylvanica, the western yellowjacket, is a Nearctic species of wasp in the genus Vespula. It is native to regions of North America, largely in areas with northern temperate climates. Its reproductive behavior is constrained by cold weather, which successfully reduces the number of western yellowjackets in cold months. However, in the absence of cold weather, this wasp's population can explode. The western yellowjacket has become particularly invasive in the Hawaiian Islands, resulting in their label as a major pest.
The eastern yellow jacket or eastern yellowjacket is a wasp found in eastern North America. Although most of their nests are subterranean, they are often considered a pest due to their nesting in recreational areas and buildings. This yellow jacket is a social insect, living in colonies of hundreds to thousands of individuals. Along with their subfamily, Vespinae, this species demonstrates supportive parental care for offspring, separation of reproductive and sterile castes, and overlapping generations. They aggressively defend their hives from threats and are known to inflict painful stings.
Polistes chinensis is a polistine vespid wasp in the cosmopolitan genus Polistes, and is commonly known as the Asian, Chinese or Japanese paper wasp. It is found in East Asia, in particular China and Japan. The subspecies P. chinensis antennalis is an invasive species in New Zealand, having arrived in 1979.
Dolichovespula saxonica, also known as the Saxon wasp, is a common social wasp found in the Palearctic region, specifically in large parts of Europe and in northern and central Asia. Although originally from continental Europe, D. saxonica has since colonised Britain, mainly in the south and east, but has been recorded as far north as East Lothian, Scotland. Most of their nests are above ground in trees and bushes, but they can also be found in buildings. Due to the proliferation of nests in urban areas and near residential homes, D. saxonica can be a pest for people. As a result, many human interventions are in place to remove Saxon nests. D. saxonica has been found to use chemical signaling in a lot of behaviours, such as alarm calls, fertility cues, and chemical trails.
The tree wasp is a species of eusocial wasp in the family Vespidae, found in the temperate regions of Eurasia, particularly in western Europe. Despite being called the tree wasp, it builds both aerial and underground paper nests, and can be found in rural and urban habitats. D. sylvestris is a medium-sized wasp that has yellow and black stripes and a black dot in the center of its clypeus. It is most common to see this wasp between May and September during its 3.5 month colony cycle.
Dolichovespula adulterina is a species of parasitic social wasp found in the Palearctic region. D. adulterina feeds on a variety of foods, including insects, spiders, arthropods, meat, molluscs, fruit, nectar, and larval secretions. D. adulterina was formerly considered to be synonymous with D. arctica from the Holarctic region, but more recent research indicates that D. arctica is a separate species.
Vespula rufa, commonly known as the red wasp, is a social wasp species belonging to the genus Vespula. It is found in northern and central Europe, parts of Asia, and northern parts of North America. Vespula rufa is characterised by red-brown markings and body segmentation, with the appearance varying amongst the different roles of individuals in the species. These wasps build small nests in dry banks underground that are not far below the surface. The colony cycle begins in the fall. Vespula rufa feed on live insects. One interesting fact about Vespula rufa is that the queen policing occurs in the species, and that worker policing occurs at much lower rates than other species in the genus Vespula. There are predators and parasites of the species. The species goes through a series of events before leaving the nest.
Vespula acadica, also known as the Forest Yellowjacket, is a North American species of eusocial wasp which is part of the "rufa" group within the genus Vespula. It is a black and yellow wasp that is found in arboreal areas and builds its nests most often in decaying vegetation like logs, but has occasionally been found to build aerial nests. Due to its preference for forests V. acadica does not normally come into contact with humans; however, when colonies are disturbed, workers of this yellowjacket may be quite aggressive and persistent and sting repeatedly.
Vespula squamosa, or the southern yellowjacket, is a social wasp. This species can be identified by its distinctive black and yellow patterning and orange queen. This species is typically found in eastern North America, and its territory extends as far south as Central America. Within these territories, they create enormous, multiple-comb nests. The colonies may be either annual or perennial depending on the climate, and in many perennial nests, polygyny takes place. In addition, this species uses pheromones both as a sexual attractant and an alarm signal. This species feeds on insects and animal carcasses; it does not produce honey. V. squamosa, a social insect, has developed a parasitic relationship with the species V. vidua and V. maculifrons. Due to their painful, venomous stings, the species is considered a pest.
Synoeca cyanea, commonly known as the marimbondo-tatu in Brazil, is a swarm-founding eusocial wasp. Native to Brazil and Argentina, S. cyanea is one of the largest and most aggressive species of social wasps and is feared in many rural areas. It begins its colony cycle in the early spring and continues until nest abandonment. Throughout its life, S. cyanea forage sugary substances and animal carcasses for food and wood pulp for its nest. S. cyanea is also known for its strong venom, which is enough to cause haemolytic activity.
Vespula atropilosa, also known as the prairie yellowjacket, is a black and yellow social wasp that forms annual colonies. Vespula atropilosa was formerly a member of the Vespa family. Males have either xanthic (yellow) or melanic (black) abdominal coloring. The prairie yellowjacket can be found in the Western and Midwestern United States as well as parts of southern Canada. It builds its nests underground and prefers large open areas such as pastures and golf courses. Vespula atropilosa colonies are founded by a queen in the spring and grow most during mid-summer. It preys on other insects such as flies and grasshoppers but is not a scavenger like other wasps. Queens mate with multiple males, and males will fight each other for matings. V. atropilosa is a social wasp that practices altruism. Prairie yellowjackets are not considered pests but are commonly found in backyards in the Pacific Northwest. This species derives its common name from the prairies and grasslands where it can be found.
Protopolybia exigua is a species of vespid wasp found in South America and Southern Brazil. These neotropical wasps, of the tribe Epiponini, form large colonies with multiple queens per colony. P. exigua are small wasps that find nourishment from nectar and prey on arthropods. Their nests are disc-shaped and hang from the undersides of leaves and tree branches. This particular species of wasp can be hard to study because they frequently abandon their nests. P. exigua continuously seek refuge from phorid fly attacks and thus often flee infested nests to build new ones. The wasps' most common predators are ants and the parasitoid phorid flies from the Phoridae family.
Parachartergus fraternus is a neotropical, swarm founding, polistine wasp species that is distributed throughout Central and South America. They live in nests in second growth tropical dry forests, near pasture fields, roadside areas, and urban areas as well. These wasps eat insects, such as caterpillars of Lepidoptera. They also drink nectar, honeydew, and water. The workers capture their prey during foraging. They also use venom to paralyze their prey in order to consume it later. P. fraternus wasps are not very aggressive and they do not attack when the nest is approached.
Vespula infernalis is an obligate parasitic wasp, parasitizing the nests of other species in the genus Vespula. Its common host species is V. acadica in North America. It is sometimes called the cuckoo yellowjacket wasp due to its inquiline lifestyle. They differ from other parasitic wasps in their intensely aggressive behaviour during invasion and occupation of the host colony. Several morphological adaptations such as bigger body parts and highly curved stingers are present in these wasps to aid their aggressive parasitic behaviour.
↑ Carpenter, J. M. (1987). "Phylogenetic relationships and classification of the Vespinae (Hymenoptera: Vespidae)". Systematic Entomology. 12 (4): 413–43. doi:10.1111/j.1365-3113.1987.tb00213.x. S2CID9388017.
1 2 3 4 5 6 7 8 9 10 11 12 13 Potter, N.B. (1964). A Study of the Biology of the Common Wasp, Vespula vulgaris L., with Special Reference to the Foraging Behavior. Bristol: University of Bristol. pp.5–60.
1 2 M. V, Brian; A. D., Brian (1952). "The wasp Vespula sylvestris Scopoli: Feeding, foraging and colony development". Transactions of the Royal Entomological Society of London. 103: 1–26. doi:10.1111/j.1365-2311.1952.tb02261.x.
1 2 3 R. J., Harris; J. R., Beggs (1991). "Variation in the Quality of Vespula vulgaris (L.) Queens (Hymenoptera: Vespidae) and Its Significance in Wasp Population Dynamics". New Zealand Journal of Zoology. 22 (2): 131–142. doi:10.1080/03014223.1995.9518030.
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Microscope-magnified image of a queen wasp's stinger
Electron micrograph of antenna surface detail, V. vulgaris
Queen returning to nest
Face
Male
