Burmese python

Burmese python
Conservation status
Vulnerable  (IUCN 3.1) [1]
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Squamata
Suborder:	Serpentes
Family:	Pythonidae
Genus:	Python
Species:	P. bivittatus
Binomial name
Python bivittatus (Kuhl, 1820)
Native distribution in green
Synonyms [2]
Python molurus bivittatus Kuhl, 1820

The Burmese python (Python bivittatus) is one of the largest species of snakes. It is native to a large area of Southeast Asia and is listed as Vulnerable on the IUCN Red List. ^[1] Until 2009, it was considered a subspecies of the Indian python, but is now recognized as a distinct species. ^[3] It is an invasive species in Florida as a result of the pet trade. ^[4]

Description

The Burmese python is a dark-colored non-venomous snake with many brown blotches bordered by black down the back. In the wild, Burmese pythons typically grow to 5 m (16 ft), ^{[5] [6]} while specimens of more than 7 m (23 ft) are unconfirmed. ^[7] This species is sexually dimorphic in size; females average only slightly longer, but are considerably heavier and bulkier than the males. For example, length-weight comparisons in captive Burmese pythons for individual females have shown: at 3.47 m (11 ft 5 in) length, a specimen weighed 29 kg (64 lb), a specimen of just over 4 m (13 ft) weighed 36 kg (79 lb), a specimen of 4.5 m (15 ft) weighed 40 kg (88 lb), and a specimen of 5 m (16 ft) weighed 75 kg (165 lb). In comparison, length-weight comparisons for males found: a specimen of 2.8 m (9 ft 2 in) weighed 12 kg (26 lb), 2.97 m (9 ft 9 in) weighed 14.5 kg (32 lb), a specimen of 3 m (9.8 ft) weighed 7 kg (15 lb), and a specimen of 3.05 m (10.0 ft) weighed 18.5 kg (41 lb). ^{[8] [9] [10] [11] [12]} In general, individuals over 5 m (16 ft) are rare. ^[13] The record for maximum length of Burmese pythons is 5.79 m (19 ft 0 in) and was caught 10 July 2023 in South Florida's Big Cypress National Preserve. ^[14] Widely published data of specimens reported to have been several feet longer are not verified. ^[7] At her death, a Burmese named "Baby" was the heaviest snake recorded in the world at the time at 182.8 kg (403 lb), ^[7] much heavier than any wild snake ever measured. ^[15] Her length was measured at 5.74 m (18 ft 10 in) circa 1999. ^[7] The minimum size for adults is 2.35 m (7 ft 9 in). ^[16] Dwarf forms occur in Java, Bali, and Sulawesi, with an average length of 2 m (6 ft 7 in) in Bali, ^[17] and a maximum of 2.5 m (8 ft 2 in) on Sulawesi. ^[18] Wild individuals average 3.7 m (12 ft) long, ^{[5] [6]} but have been known to reach 5.79 m (19 ft 0 in). ^[14]

Diseases

In both their native and invasive range they suffer from Raillietiella orientalis (a pentastome parasitic disease). ^[19]

Distribution and habitat

The Burmese python occurs throughout Southern and Southeast Asia, including eastern India, southeastern Nepal, western Bhutan, southeastern Bangladesh, Myanmar, Thailand, Laos, Cambodia, Vietnam, northern continental Malaysia, and southern China in Fujian, Jiangxi, Guangdong, Hainan, Guangxi, and Yunnan. ^[20] It also occurs in Hong Kong, and in Indonesia on Java, southern Sulawesi, Bali, and Sumbawa. ^[21] It has also been reported in Kinmen. ^[22]

It is an excellent swimmer and needs a permanent source of water. It lives in grasslands, marshes, swamps, rocky foothills, woodlands, river valleys, and jungles with open clearings. It is a good climber and has a prehensile tail. It can stay in water for 30 minutes but mostly stays on land.^{[ citation needed ]}

As an invasive species

Main article: Burmese pythons in Florida

United States range in 2007

A captured Burmese python in the Florida Everglades

Python invasion has been particularly extensive, notably across South Florida, where a large number of pythons can now be found in the Florida Everglades. ^{[23] [24]} Between 1996 and 2006, the Burmese python gained popularity in the pet trade, with more than 90,000 snakes imported into the U.S. ^[25] The current number of Burmese pythons in the Florida Everglades may have reached a minimum viable population and become an invasive species. Hurricane Andrew in 1992 was deemed responsible for the destruction of a python-breeding facility and zoo, and these escaped snakes spread and populated areas into the Everglades. ^[26] More than 1,330 have been captured in the Everglades. ^[27] A genetic study in 2017 revealed that the python population is composed of hybrids between the Burmese python and Indian python. The species also displays cytonuclear discordance which has made phylogenetic studies of its origin more complicated. ^[28]

By 2007, the Burmese python was found in northern Florida and in the coastal areas of the Florida Panhandle. The importation of Burmese pythons was banned in the United States in January 2012 by the U.S. Department of the Interior. ^[29] A 2012 report stated, "in areas where the snakes are well established, foxes, and rabbits have disappeared. Sightings of raccoons are down by 99.3%, opossums by 98.9%, and white-tailed deer by 94.1%." ^[30] Road surveys between 2003 and 2011 indicated an 87.3% decrease in bobcat populations, and in some areas rabbits have not been detected at all. ^[31] Experimental efforts to reintroduce rabbit populations to areas where rabbits have been eliminated have mostly failed "due to high (77% of mortalities) rates of predation by pythons." ^[32] Bird and coyote populations may be threatened, as well as the already-rare Florida panther. ^[30] In addition to this correlational relationship, the pythons have also been experimentally shown to decrease marsh rabbit populations, further suggesting they are responsible for many of the recorded mammal declines. They may also outcompete native predators for food. ^[33]

For example, Burmese pythons also compete with the native American alligator, and numerous instances of alligators and pythons attacking—and in some cases, preying on—each other have been reported and recorded.

By 2011, researchers identified up to 25 species of birds from nine avian orders in the digestive tract remains of 85 Burmese pythons found in Everglades National Park. ^[34] Native bird populations are suffering a negative impact from the introduction of the Burmese python in Florida; among these bird species, the wood stork is of specific concern, now listed as federally endangered. ^[34]

Numerous efforts have been made to eliminate the Burmese python population in the last decade.^{[ when? ]} Understanding the preferred habitat of the species is needed to narrow down the python hunt. Burmese pythons have been found to select broad-leafed and low-flooded habitats. Broad-leafed habitats comprise cypress, overstory, and coniferous forest. Though aquatic marsh environments would be a great source for prey, the pythons seem to prioritize environments allowing for morphological and behavioral camouflage to be protected from predators.^{[ clarification needed ]} Also, the Burmese pythons in Florida have been found to prefer elevated habitats, since this provides the optimal conditions for nesting. In addition to elevated habitats, edge habitats are common places where Burmese pythons are found for thermoregulation, nesting, and hunting purposes. ^[25]

One of the Burmese python eradication movements with the biggest influence was the 2013 Florida Python Challenge. This was a month-long contest wherein a total of 68 pythons were removed. The contest offered incentives such as prizes for longest and greatest number of captured pythons. The purpose of the challenge was to raise awareness about the invasive species, increase participation from the public and agency cooperation, and to remove as many pythons as possible from the Florida Everglades. ^[35] The challenge has run a few times again since then and is now an annual event over the duration of ten days. ^[36] Recently, in 2023, it resulted in 209 pythons removed by 1,050 participants. ^[37]

A study from 2017 introduced a new method for identifying the presence of Burmese pythons in southern Florida; this method involves the screening of mosquito blood. Since the introduction of the Burmese python in Florida, mosquito communities use the pythons as hosts even though they are recently introduced. ^[38]

Invasive Burmese pythons also face certain physiological changes. Unlike their native South Asian counterparts who spend long periods fasting due to seasonal variation in prey availability, pythons in Florida feed year-round due to the constant availability of food. They are also vulnerable to cold stress, with winter freezes resulting in mortality rates of up to 90%. Genomic data suggests natural selection on these populations favors increased thermal tolerance as a result of these high-mortality freezes. ^[39]

They have carried Raillietiella orientalis, a pentastome parasitic disease, with them from Southeast Asia. Other reptiles in Florida have become infested, and the parasite appears to have become endemic. ^[19]

In April 2019, researchers captured and killed a large Burmese python in Florida's Big Cypress National Preserve. It was more than 5.2 m (17 ft) long, weighed 64 kg (140 lb), and contained 73 developing eggs. ^[40] In December 2021, a Burmese python was captured in Florida that weighed 98 kg (215 lb) and had a length of 5.5 m (18 ft); it contained a record 122 developing eggs. ^[41] In July 2023, local hunters captured and killed a 5.8 m (19 ft) long Burmese python that weighed 57 kg (125 lb) in Florida's Big Cypress National Preserve. ^[14]

Behavior

Burmese pythons are mainly nocturnal rainforest dwellers. ^[42] When young, they are equally at home on the ground and in trees, but as they gain girth, they tend to restrict most of their movements to the ground. They are also excellent swimmers, being able to stay submerged for up to half an hour. Burmese pythons spend the majority of their time hidden in the underbrush. In the northern parts of its range, the Burmese python may brumate for some months during the cold season in a hollow tree, a hole in the riverbank, or under rocks. Brumation ^[43] is biologically distinct from hibernation. While the behavior has similar benefits, allowing organisms to endure the winter without moving, it also involves the preparation of both male and female reproductive organs for the upcoming breeding season. The Florida population also goes through brumation. ^[44]

They tend to be solitary and are usually found in pairs only when mating. Burmese pythons breed in the early spring, with females laying clutches of 12–36 eggs in March or April. They remain with the eggs until they hatch, wrapping around them and twitching their muscles in such a way as to raise the ambient temperature around the eggs by several degrees. Once the hatchlings use their egg tooth to cut their way out of their eggs, no further maternal care is given. The newly hatched babies often remain inside their eggs until they are ready to complete their first shedding of skin, after which they hunt for their first meal. ^[45]

Parthenogenesis

The Burmese python is able to reproduce asexually when in captivity. ^[12] Offspring are clones of their mother and reproduction appears to be by a parthenogenetic mechanism that involves a modification of the meiotic process. ^[12]

Diet

Burmese python photographed in Bardiya National Park, Nepal

Like all snakes, the Burmese python is carnivorous. Its diet consists primarily of birds and mammals, but also includes amphibians and reptiles. It is a sit-and-wait predator, meaning it spends most of its time staying relatively still, waiting for prey to approach, then striking rapidly. ^[46] The snake grabs a prey animal with its sharp teeth, then wraps its body around the animal to kill it through constriction. ^[47] The python then swallows its prey whole. It is often found near human habitation due to the presence of rats, mice, and other vermin as a food source. However, its equal affinity for domesticated birds and mammals means it is often treated as a pest. In captivity, its diet consists primarily of commercially available appropriately sized rats, graduating to larger prey such as rabbits and poultry as it grows. As an invasive species in Florida, Burmese pythons primarily eat a variety of small mammals including foxes, rabbits, and raccoons. Due to their high predation levels, they have been implicated in the decline and even disappearance of many mammal species. ^{[4] [33]} In their invasive range, pythons also eat birds and occasionally other reptiles. Exceptionally large pythons may even require larger food items such as pigs or goats, and are known to have attacked and eaten alligators and adult deer in Florida. ^[48]

Digestion

The digestive response of Burmese pythons to such large prey has made them a model species for digestive physiology. Its sit-and-wait hunting style is characterized by long fasting periods in between meals, with Burmese pythons typically feeding every month or two, but sometimes fasting for as long as 18 months. ^[46] As digestive tissues are energetically costly to maintain, they are downregulated during fasting periods to conserve energy when they are not in use. ^[49] A fasting python has a reduced stomach volume and acidity, reduced intestinal mass, and a 'normal' heart volume. After ingesting prey, the entire digestive system undergoes a massive re-modelling, with rapid hypertrophy of the intestines, production of stomach acid, and a 40% increase in mass of the ventricle of the heart to fuel the digestive process. ^[50] During digestion, the snake's oxygen consumption rises drastically as well, increasing with meal size by 17 to 40 times its resting rate. ^[46] This dramatic increase is a result of the energetic cost of restarting many aspects of the digestive system, from rebuilding the stomach and small intestine to producing hydrochloric acid to be secreted in the stomach. Hydrochloric acid production is a significant component of the energetic cost of digestion, as digesting whole prey items requires the animal to be broken down without the use of teeth, either for chewing or tearing into smaller pieces. To compensate, once food has been ingested, Burmese pythons begin producing large amounts of acid to make the stomach acidic enough to turn the food into a semi-liquid that can be passed through to the small intestine and undergo the rest of the digestive process.^{[ citation needed ]}

The energy cost is highest in the first few days after eating when these regenerative processes are most active, meaning Burmese pythons rely on existing food energy storage to digest a new meal. ^{[46] [51]} Overall, the entire digestive process from food intake to defecation lasts 8–14 days. ^[49]

Conservation

Leather goods and skins of Burmese pythons and reticulated pythons (Malayopython reticulatus) at a local shop at Mandalay, Myanmar

The Burmese python is listed on CITES Appendix II. ^[1] It has been listed as vulnerable on the IUCN Red List since 2012, as the wild population is estimated to have declined by at least 30% in the first decade of the 21st century due to habitat loss and over-harvesting. ^[1]

To maintain Burmese python populations, the IUCN recommends increased conservation legislation and enforcement at the national and international levels to reduce harvesting across the snake's native range. The IUCN also recommends increased research into its population ecology and threats. In Hong Kong, it is a protected species under Wild Animals Protection Ordinance Cap 170. It is also protected in Thailand, Vietnam, China, and Indonesia. However, it is still common only in Hong Kong and Thailand, with rare to very rare statuses in the rest of its range.^{[ citation needed ]}

In captivity

Audience volunteers holding an adult Burmese python

An amelanistic Burmese python at a zoo in Japan

Burmese pythons are often sold as pets, and are made popular by their attractive coloration and apparently easy-going nature. However, they have a rapid growth rate, and can exceed 2.1 m (6 ft 11 in) in length in a year if power fed. However this may cause health issues in the future. By age four, they will have reached their adult size, though they continue growing very slowly throughout their lives, which may exceed 20 years.

Although the species has a reputation for docility, they are very powerful animals – capable of inflicting severe bites and even killing by constriction. ^{[52] [53] [54] [55] [56] [57]} They also consume large amounts of food, and due to their size, require large, often custom-built, secure enclosures. As a result, some are released into the wild, and become invasive species that devastate the environment. For this reason, some jurisdictions (including Florida, due to the python invasion in the Everglades ^[58] ) have placed restrictions on the keeping of Burmese pythons as pets. Violators could be imprisoned for more than seven years or fined $500,000 if convicted.

Burmese pythons are opportunistic feeders; ^[59] they eat almost any time food is offered, and often act hungry even when they have recently eaten. As a result, they are often overfed, causing obesity-related problems to be common in captive Burmese pythons.

Like the much smaller ball python, Burmese pythons are known to be easygoing or timid creatures, which means that if cared for properly, they can easily adjust to living near humans. ^[60]

Handling

Although pythons are typically afraid of people due to their great stature, and generally avoid them, special care is still required when handling them. Given their adult strength, multiple handlers (up to one person per meter of snake) are usually recommended. ^[61] Some jurisdictions require owners to hold special licenses, and as with any wild animal being kept in captivity, treating them with the respect an animal of this size commands is important. ^[62]

Variations

Caramel Burmese python

The Burmese python is frequently captive-bred for color, pattern, and more recently, size. Its amelanistic form is especially popular and is the most widely available morph. This morph is white with patterns in butterscotch yellow and burnt orange. Also, "labyrinth" specimens with maze-like patterns, khaki-colored "green", and "granite" with many small angular spots are available. Breeders have recently begun working with an island lineage of Burmese pythons. Early reports indicate that these dwarf Burmese pythons have slightly different coloring and pattern from their mainland relatives and do not grow much over 2.1 m (6 ft 11 in) in length. One of the most sought-after of these variations is the leucistic Burmese. This particular variety is very rare, being entirely bright white with no pattern and blue eyes, and has only in 2008/2009 been reproduced in captivity as the homozygous form (referred to as "super" by reptile keepers) of the co-dominant hypomelanistic trait. The caramel Burmese python has a caramel-colored pattern with "milk-chocolate" eyes.

See also

• Inclusion body disease, a viral disease affecting pythons

References

1. Stuart, B.; Nguyen, T.Q.; Thy, N.; Grismer, L.; Chan-Ard, T.; Iskandar, D.; Golynsky, E. & Lau, M.W. (2019) [errata version of 2019 assessment]. "Python bivittatus". IUCN Red List of Threatened Species . 2019: e.T193451A151341916. Retrieved 6 April 2021.
2. Python bivittatus at the Reptarium.cz Reptile Database
3. Jacobs, H.J.; Auliya, M.; Böhme, W. (2009). "On the taxonomy of the Burmese Python, Python molurus bivittatus KUHL, 1820, specifically on the Sulawesi population". Sauria. 31 (3): 5–11.
4. Sarill, M. (2016). "Burmese Pythons in the Everglades". Berkeley Rausser College of Natural Resources. Retrieved 2021-03-17.
5. Smith MA (1943). "Python molurus bivittatus", pp. 108–109 in The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. III.—Serpentes. London: Secretary of State for India. Taylor and Francis.
6. Campden-Main SM (1970). A Field Guide to the Snakes of South Vietnam. Washington, District of Columbia. pp. 8–9.
7. Barker, D.G.; Barten, S.L.; Ehrsam, J.P. & Daddono, L. (2012). "The corrected lengths of two well-known giant pythons and the establishment of a new maximum length record for Burmese Pythons, Python bivittatus" (PDF). Bulletin of the Chicago Herpetological Society. 47 (1): 1–6. Retrieved 2020-03-02.
8. Van Mierop, L.H. & Barnard, S.M. (1976). "Observations on the reproduction of Python molurus bivittatus (Reptilia, Serpentes, Boidae)". Journal of Herpetology. 10 (4): 333–340. doi:10.2307/1563071. JSTOR   1563071.
9. Barker, D.G.; Murphy J.B. & Smith, K.W. (1979). "Social behavior in a captive group of Indian pythons, Python molurus (Serpentes, Boidae) with formation of a linear social hierarchy". Copeia. 1979 (3): 466–471. doi:10.2307/1443224. JSTOR   1443224.
10. Marcellini, D.L. & Peters, A. (1982). "Preliminary observations on endogeneous heat production after feeding in Python molurus". Journal of Herpetology. 16 (1): 92–95. doi:10.2307/1563914. JSTOR   1563914.
11. Jacobson, E.R.; Homer, B. & Adams, W. (1991). "Endocarditis and congestive heart failure in a Burmese python (Python molurus bivittatus)". Journal of Zoo and Wildlife Medicine. 22: 245–248.
12. Groot TV, Bruins E, Breeuwer JA. Molecular genetic evidence for parthenogenesis in the Burmese python, Python molurus bivittatus. Heredity (Edinb). 2003 Feb;90(2):130-5. doi: 10.1038/sj.hdy.6800210. PMID 12634818
13. Saint Girons, H. (1972). "Les serpents du Cambodge". Mémoires du Muséum national d'Histoire naturelle. Série A: 40–41.
14. Jones, Dustin (13 July 2023). "A record-breaking Burmese python — as long as a giraffe is tall — caught in Florida". NPR. Retrieved 14 July 2023.
15. Rivas, J.A. (2000). The life history of the green anaconda (Eunectes murinus), with emphasis on its reproductive Biology (PDF) (Ph.D. thesis). University of Tennessee. Archived from the original (PDF) on 2016-03-03.
16. Murphy, J.C. & Henderson, R.W. (1997). Tales of Giant Snakes: A Historical Natural History of Anacondas and Pythons. Krieger Pub. Co. pp. 2, 19, 37, 42, 55–56. ISBN   0-89464-995-7.
17. McKay, J.L. (2006). A Field Guide to the Amphibians and Reptiles of Bali. Krieger Publishing Company. pp. 13, 14, 18, 86. ISBN   1-57524-190-0.
18. De Lang, R. & Vogel, G. (2005). The Snakes of Sulawesi: A Field Guide to the Land Snakes of Sulawesi with Identification Keys. Frankfurt Contributions to Natural History (Band 25 ed.). Chimaira. pp. 23–27, 198–201. ISBN   3-930612-85-2.
19. Waymer, J. (2019). "Bloodsucking worms in pythons are killing Florida snakes, study says". Florida Today . Retrieved 2021-12-16.
20. Barker, D.G.; Barker, T.M. (2010). "The Distribution of the Burmese Python, Python bivittatus, in China" (PDF). Bulletin of the Chicago Herpetological Society. 45 (5): 86–88. Archived from the original (PDF) on 2016-03-03. Retrieved 2012-06-26.
21. Barker, D.G.; Barker, T.M. (2008). "The distribution of the Burmese Python, Python molurus bivittatus" (PDF). Bulletin of the Chicago Herpetological Society. 43 (3): 33–38. Archived from the original (PDF) on 2014-05-20. Retrieved 2012-10-17.
22. Breuer, H.; Murphy, W.C. (2009–2010). "Python molurus bivittatus". Snakes of Taiwan. Archived from the original on 26 June 2012. Retrieved 17 October 2012.
23. "Top 10 Invasive Species". Time. 2010. Archived from the original on February 6, 2010. Retrieved 27 April 2010.
24. Species Profile - Burmese Python (Python molurus bivittatus). National Invasive Species Information Center, United States National Agricultural Library.
25. Walters TM, Mazzotti FJ, Fitz HC (2016). "Habitat selection by the invasive species Burmese python in Southern Florida". Journal of Herpetology. 50 (1): 50–56. doi:10.1670/14-098. S2CID   86327588.
26. "Democrats Hold Hearing on Administration's Plan to Constrict Snakes in the Everglades - House Committee on Natural Resources". Naturalresources.house.gov. 2010. Archived from the original on 16 September 2012. Retrieved 9 August 2012.
27. "(US National Park Service website - December 31, 2009)". nps.gov. Retrieved 8 April 2019.
28. Hunter, M.E.; Johnson, N.A.; Smith, B.J.; Davis, M.C.; Butterfield, John S.S.; Snow, R.W.; Hart, K.M. (2017). "Cytonuclear discordance in the Florida Everglades invasive Burmese python (Python bivittatus) population reveals possible hybridization with the Indian python (P. molurus)". Ecology and Evolution. 8 (17): 9034–9047. doi:10.1002/ece3.4423. PMC   6157680 . PMID   30271564.
29. "Salazar Announces Ban on Importation and Interstate Transportation of Four Giant Snakes that Threaten Everglades". doi.gov (Press release). 2012. Retrieved April 26, 2022.
30. Adams, G. (2012). "Pythons are squeezing the life out of the Everglades, scientists warn". The Independent. London.
31. Dorcas ME, Willson JD, Reed RN, Snow RW, Rochford MR, Miller MA, Meshaka WE, Andreadis PT, Mazzotti FJ, Romagosa CM, Hart KM (2012). "Severe mammal declines coincide with proliferation of invasive Burmese pythons in Everglades National Park". Proceedings of the National Academy of Sciences. 109 (7): 2418–2422. Bibcode:2012PNAS..109.2418D. doi: 10.1073/pnas.1115226109 . PMC   3289325 . PMID   22308381.
32. Willson, J. (2017). "Indirect effects of invasive Burmese pythons on ecosystems in southern Florida". Journal of Applied Ecology. 54 (4): 1251–1258. Bibcode:2017JApEc..54.1251W. doi: 10.1111/1365-2664.12844 .
33. McCleery RA, Sovie A, Reed RN, Cunningham MW, Hunter ME, Hart KM (2015). "Marsh rabbit mortalities tie pythons to the precipitous decline of mammals in the Everglades". Proceedings of the Royal Society B: Biological Sciences. 282 (1805): 20150120. doi:10.1098/rspb.2015.0120. PMC   4389622 . PMID   25788598.
34. Dove CJ, Snow RW, Rochford MR, Mazzotti FJ (2011). "Birds Consumed by the Invasive Burmese Python (Python molurus bivittatus) in Everglades National Park, Florida, USA". The Wilson Journal of Ornithology. 123 (1): 126–131. doi:10.1676/10-092.1. JSTOR   23033493. S2CID   55495469.
35. Mazzotti FJ, Rochford M, Vinci J, Jeffery BM, Eckles JK, Dove C, Sommers KP (2016). "Implications of the 2013 Python Challenge® for Ecology and Management of Python molorus bivittatus (Burmese python) in Florida". Southeastern Naturalist. 15 (sp8): 63–74. doi:10.1656/058.015.sp807. JSTOR   26454670. S2CID   90352897.
36. "Florida Python Challenge - Python Challenge". Florida Python Challenge. Retrieved 2024-03-29.
37. "ICYMI: Lieutenant Governor Jeanette Nuñez Announces Winners of the 2023 Florida Python Challenge®". Florida Fish And Wildlife Conservation Commission. Retrieved 2024-03-29.
38. Reeves LE, Krysko KL, Avery ML, Gillett-Kaufman JL, Kawahara AY, Connelly CR, Kaufman PE (2018-01-17). Paul R (ed.). "Interactions between the invasive Burmese python, Python bivittatus Kuhl, and the local mosquito community in Florida, USA". PLOS ONE. 13 (1): e0190633. Bibcode:2018PLoSO..1390633R. doi: 10.1371/journal.pone.0190633 . PMC   5771569 . PMID   29342169.
39. Card DC, Perry BW, Adams RH, Schield DR, Young AS, Andrew AL, Jezkova T, Pasquesi GI, Hales NR, Walsh MR, Rochford MR, Mazzotti FJ, Hart KM, Hunter ME, Castoe TA (2018). "Novel ecological and climatic conditions drive rapid adaptation in invasive Florida Burmese pythons". Molecular Ecology. 27 (23): 4744–4757. Bibcode:2018MolEc..27.4744C. doi:10.1111/mec.14885. PMID   30269397.
40. Mettler, K. (2019). "A 17-foot, 140-pound python was captured in a Florida park. Officials say it's a record". The Washington Post. Retrieved 23 May 2020.
41. Williams, A. B. "Caught! Record-breaking 18-foot Burmese python pulled from Collier County wilderness". The News-Press. Retrieved 2022-06-25.
42. Evans S (2003). "Python molurus, Burmese Python". The deep Scaly Project. Digital Morphology. Retrieved 2007-07-25.
43. "Glossary of reptile and amphibian terminology". Kingsnake.com. Retrieved 8 April 2019.^{[ permanent dead link ]}
44. Krusling, Lindsey A. "1.4 Invasive species Burmese python (Python bivittatus) and its effect in Florida". In Shaul, Travis R.; Shaul, Kylienne A.; Weaver, Ella M. (eds.). Environmental ScienceBites. Vol. 2. The Ohio State University.
45. Ghosh A (11 July 2012). "Burmese Python". AnimalSpot.net. Retrieved 27 December 2012.
46. Secor, S.M.; Diamond, J. (1995). "Adaptive responses to feeding in Burmese pythons: pay before pumping". The Journal of Experimental Biology. 198 (Pt 6): 1313–1325. doi: 10.1242/jeb.198.6.1313 . PMID   7782719.
47. Szalay, J. (2016). "Python Facts". Live Science. Retrieved 2021-03-17.
48. "Photo in the News: Python Bursts After Eating Gator (Update)". National Geographic News. 2006. Archived from the original on October 21, 2018. Retrieved 8 April 2019.
49. Starck, J.M.; Beese, K. (2001). "Structural flexibility of the intestine of Burmese python in response to feeding". The Journal of Experimental Biology. 204 (2): 325–335. doi:10.1242/jeb.204.2.325. PMID   11136618.
50. Secor, S.M. (2008). "Digestive physiology of the Burmese python: broad regulation of integrated performance". Journal of Experimental Biology. 211 (24): 3767–3774. doi: 10.1242/jeb.023754 . PMID   19043049. S2CID   5545174.
51. Secor, S.M. (2003). "Gastric function and its contribution to the postprandial metabolic response of the Burmese python Python molurus". The Journal of Experimental Biology. 206 (10): 1621–1630. doi: 10.1242/jeb.00300 . PMID   12682094.
52. "Python Kills Careless Student Zookeeper in Caracas" . The Telegraph. London. AP. 2008-08-26. ISSN   0307-1235. Archived from the original on 2022-01-12. Retrieved 2019-02-18.
53. Chiszar D, Smith HM, Petkus A, Doughery J (1993). "A Fatal Attack on a Teenage Boy by a Captive Burmese Python (Python molurus bivittatus) in Colorado" (PDF). The Bulletin of the Chicago Herpetological Society. 28 (#12). Chicago Herpetological Society: 261. ISSN   0009-3564. Archived from the original (PDF) on 2019-02-18.
54. Kaplan M (1994). "The Keeping of Large Pythons: Realities and Responsibilities". www.anapsid.org. Herp Care Collection. Retrieved 2019-02-18.
55. "Python Caused Death in Ontario Home in 1992 Case". Toronto News. CBC News. Toronto: Canadian Broadcasting Commission. Canadian Press. 2013-04-13. ISSN   0708-9392 . Retrieved 2019-02-17.
56. Davison J (2013-08-07). "Python-linked Deaths Raise Questions over Exotic Animal Laws". News. CBC News. Toronto: Canadian Broadcasting Corporation. ISSN   0708-9392 . Retrieved 2019-02-17.
57. "Dr. D. H. Evans, Coroner of Ontario, "Inquest into the Death of Mark Nevilles: Verdict of Coroner's Jury" (Brampton, Ontario: June 1992)". documentcloud.org. Retrieved 8 April 2019.
58. Burrage G (30 June 2010). "New law makes Burmese python illegal in Florida". Abcactionnews.com. Archived from the original on 2013-05-01. Retrieved 9 August 2012.
59. Orzechowski SC, Romagosa CM, Frederick PC (2019-07-01). "Invasive Burmese pythons (Python bivittatus) are novel nest predators in wading bird colonies of the Florida Everglades". Biological Invasions. 21 (7): 2333–2344. Bibcode:2019BiInv..21.2333O. doi: 10.1007/s10530-019-01979-x . S2CID   102350541.
60. "Python bivittatus (Kuhl, 1820)". www.gbif.org. Archived from the original on 22 October 2018. Retrieved 8 April 2019.
61. "Playing with the Big Boys: Handling Large Constrictors". www.anapsid.org. Retrieved 8 September 2017.
62. "Captive Animals - Most states have no laws governing captive wild animals". Animal Legal Defense Fund. Retrieved 8 April 2019.

Further reading

• Christy (2008). The Lizard King: The True Crimes and Passions of the World's Greatest Reptile Smugglers. New York: TWELVE. ISBN   978-0-446-58095-3.
• Mattison C (1999). Snake. DK Publishing. ISBN   978-0-7894-4660-2.
• Willson JD, Dorcas ME, Snow RW (2010-11-21). "Identifying plausible scenarios for the establishment of invasive Burmese pythons (Python molurus) in Southern Florida". Biological Invasions. 13 (7): 1493–1504. doi:10.1007/s10530-010-9908-3. S2CID   207096799.

External links

• Burmese python (Python molurus) - EDDMapS State Distribution - EDDMapS
• Species profile - Burmese Python (Python molurus), National Invasive Species Information Center, U.S. Department of Agriculture National Agricultural Library