Calcaires du Bou Dahar

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Calcaires du Bou Dahar
Stratigraphic range: late Sinemurian–earliest Toarcian 194–183  Ma
Jbel Bou Dahar early Mesozoic map.jpg
Shape, lithology and location within Morocco of the Bou Dahar platform
Type Geological formation
Underlies Tagoudite Formation
Overlies Central Atlantic Magmatic Province igneous rocks
Area Bni Tadjite
Thickness>450 m
Lithology
Primarylimestone
Location
LocationHigh Atlas
Coordinates 32°18′N3°30′W / 32.3°N 3.5°W / 32.3; -3.5
Approximate paleocoordinates 25°36′N1°18′W / 25.6°N 1.3°W / 25.6; -1.3
Country Morocco
Type section
Thickness at type section~360 m (1,180 ft)
Morocco relief location map.jpg
Blue pog.svg
Calcaires du Bou Dahar (Morocco)

The Calcaires du Bou Dahar (also known as Djebel Bou Dahar Paleoshoal, Djebel Bou Dahar, Calcaires du Bou Dahar Formation, or Bou Dahar Formation) is a geological formation or a sequence of formations of Late Sinemurian to Pliensbachian-Toarcian boundary (Early Jurassic) age in Bni Tadjite, the Central High Atlas, Morocco. This unit represents an excepcional record of an evolving reef complex (mountain laterals), platform slopes (Steep slopes between 20° and 35° on various sides) and a emerged shoal (nearly horizontal limestone layers on the top flat Bou Dahar plateau) developed inside a carbonate platform, recording the evolutionary cycles of this environment with notorious precision, also yielding what is considered one of the greatest/most diverse marine biotas of the entire Jurassic Tethys Ocean. [1] The Bou Dahar carbonate platform shoal stands prominently and structurally above surrounding plains, spanning 35-40 km in length and 4-15 km in width, with a relief of 100-450 m. This carbonate formation originated on metamorphosed Silurian to Ordovician siliciclastic rocks and tholeiitic volcanic layers tied to Central Atlantic Magmatic Province basalts, forming a corridor oriented WSW to ENE. Surrounding alluvial plains expose green marls, shales, and dark lime-mudstones representing basinal deposits contemporaneous or subsequent to the platform. [2] [3] It has been considered to be a sequence of different coeval inner geological formations, including the Foum Zidet Formation, the Aganane Formation and Ouchbis Formation, but is usually interpreted as a single major unit due to it´s unique preservation. [2]

Contents

Economic importance

Lead, Zinc, and Baryte mining in the Bou Dahar district relies solely on artisanal methods managed by the public "CADETAF" cooperative. [4] The cooperative’s claim area, spanning 60,000 km2 across the central and eastern High Atlas, along with exploration licenses held by ManagemGroup, together constitute North Africa’s largest calamine region. Data from CADETAF and ManagemGroup reveal over 1 million tons of zinc ore with more than 30% ZnO content extracted to date, including 559,403 tons of 16% Zn ore produced by ManagemGroup from 2012 to 2019. [4]

Sequence evolution

Succession of depositional sequences in the Bou Dahar platform Jbel Bou Dahar sequences.png
Succession of depositional sequences in the Bou Dahar platform

The Bou Dahar platform is a well-preserved example of carbonate platform development within a tectonically active rift-basin environment. This region formed as part of the Jurassic High Atlas seaway, where tectonic activity shaped the basin, with influences from rifting, faulting, and transtensional forces. [5] Early studies mapped the area’s Jurassic marine environments, with carbonate platform margins situated alongside deeper basin limestones. This platform records two primary periods of shallow-water carbonate growth: one during the Lower to Middle Liassic (Sinemurian to Domerian) and a subsequent phase spanning the Upper Liassic to Lower Dogger (Aalenian to Bajocian). However, only the earlier phase is preserved in Djebel Bou Dahar. [6] This phase was abruptly interrupted at the Domerian-Toarcian transition, marked by the onlap of Toarcian shales and Aalenian lime mudstones over the platform, signaling a shift to deeper marine conditions. [5]

The seismic modeling of Bou Dahar platform highlighted large-scale structures, such as the platform's form and the boundary marking its submergence, across various frequencies and modeling approaches. Lower-frequency wavelets emphasized truncation of layers and outward shifts in onlap points. [3] In contrast, internal platform facies zones were less distinct at frequencies below 50 Hz, but with a geometry still discernible through disturbances in reflections, especially near the lower boundaries, with image-ray migration models proving more responsive to these subtle variations even at lower frequencies. Seismic contrast, driven by differences in acoustic impedance, tends to decline with deeper burial and subsequent diagenesis, with peak resolution typically present at initial deposition stages before significant burial effects. [3]

Sequences I-II overlie CAMP basalts, spanning over 126 & 150 meters, marked by a dolomite-rich boundary featuring fenestral and stromatolitic structures, which indicate transgressive-regressive (T-R) cycles. Dominance of subtidal carbonates suggests extensive shallow marine conditions across the platform. [7] Early dolomitization preserved the fine microstructures within these sequences, aligning with the Hettangian-Lower Sinemurian Idikel Formation. [2] [8]

Sequence III developed as a low-relief carbonate platform and varies in slope angle between northern and southern margins, with thickness peaking in northeastern synclines at around 100 meters. Thinning toward the center, it comprises laterally continuous skeletal and packstone deposits interspersed with sponge mounds, mirroring the Foum Zidet Formation. [2] [7] [8]

Sequence IV with a wedge-shaped geometry, reveals steep slopes and retrogradational stacking, indicating subaerial exposure at its base with ammonite-rich wackestone beds. [7] This sequence thickens at the southern margin, where coral and sponge boundstones, extending to 140 meters, transition into grainstone and packstone accumulations along the slope. The eastern platform edge hosts cross-bedded grainstone layers, lacking intertidal deposits, characteristic of open-marine conditions. [2] [7] [8]

Sequence V, along Sequences IV to VI correlate with the Aganane Formation and Ouchbis Formations, being a progradational wedge, is thickest along the platform's southern and eastern edges, forming a concave-up clinoform structure, reaching 50-70 meters in height and indicating alternating subtidal and intertidal lithofacies and evidence of exposure events. Moving basinward, finer basinal sediments with green marls and peloidal wackestones emerge, suggesting periodic sediment starvation during the Early Pliensbachian. [2] [7] [8]

The final sequence, Sequence VI, fills the last stage of platform development with angular unconformity over older layers and CAMP basalts. [7] This sequence encompasses varied facies belts surrounding a central paleo-high with subtidal and intertidal deposits, coated-grain bars, and coastal-plain sediments like red shales and calcretes. Clinoforms along the lower slope show a concave-up profile that flattens basinward, with deposits consolidating into a fan along the southern margin, evidence of sediment redistribution likely from mass wasting. [2] [8]

Paleoenvironment

Patch reef in French Bay (San Salvador Island, Bahamas) 1 (15897790627).jpg
20130118-HighbormeCay-Stromatolite-03.JPG
Patch reef (left) and microbialite (right) in the Bahamas. The Bou Dahar sediments record multiple environments within a carbonate platform: from patch reefal areas to shallow microbialites, like the modern Bahamas.

The paleoenvironment of Bou Dahar is characterized by a dynamic evolution through distinct stages: pre-drowning, drowning, and post-drowning, each marked by specific sedimentological and ecological changes that reflect shifts in energy conditions and depositional environments. [8] [9]

During the pre-drowning phase, the carbonate platform was dominated by tranquil lagoonal settings, indicative of low-energy environments. These conditions fostered the accumulation of microsparite carbonates, which are essential in maintaining the stable conditions found in lagoons. [8] The presence of bioturbation, mud cracks, and tidal flats illustrates the periodic exposure of the substrate, suggesting a complex interplay of biogenic and abiogenic processes. The platform margin transitioned into coarser zones where higher-energy environments were characterized by sorted carbonate sands and natural barriers. This transitional buffer from the lagoon to slope facilitated the formation of intertidal mounds, which played a crucial role in the sediment dynamics of the region. [8]

As the platform underwent drowning, it shifted towards high-energy conditions. Sediments from this phase exhibited dense biosparites with significant skeletal deposits, indicating heightened wave action and the erosion of previously accumulated lagoonal deposits. The transition to these high-energy environments underscores a significant change in the depositional dynamics, where sediment transport was dominated by stronger currents. The emergence of tepee structures in the platform carbonates reflects this period of increased energy. [5] Formed from early lithification and sediment infill in intertidal zones, these structures were characterized by cracks filled with fine sediments and stromatolitic algae, highlighting rapid infilling processes shortly after deposition. Unlike their recent counterparts in the Persian Gulf, the Moroccan tepees developed in fully lithified rocks, shaped by thermal expansion and contraction driven by temperature variations in intertidal settings. [10]

Microscopic examination of the sediments reveals a rich diversity of biogenic and non-biogenic components, categorized into nine distinct microfacies types. [5] These include laminated pelmicrites, biopelsparites, and various intraclasts and bioclasts, showcasing the complex interplay of environmental factors influencing sediment composition. Foraminifers along with corals, indicate thriving marine ecosystems during the deposition of Bou Dahar. [9] The various microfacies illustrate shifts in sedimentation dynamics, with microfacies types reflecting different depositional environments-from tranquil lagoonal settings to more energetic reef environments. [11]

Local carbonate factories present a unique contrast with other coeval units in the Tethys, with siliceous sponge microbial boundstone mounds, which were more prevalent in Middle and Upper Jurassic systems. Despite their earlier emergence, these mounds show similarities to younger Jurassic formations in texture and biological components. They thrived in deep-water environments, particularly in areas that experienced low detrital input. [12] Siliceous sponge mounds record distinct rare earth element and yttrium patterns among carbonate components, indicating that Early Jurassic seawater resembled modern distributions. Microbialites contained higher rare earth element concentrations, likely influenced by organic matter and diagenetic processes. These mounds thrived in well-oxygenated environments, emphasizing the need to examine individual carbonate types to understand their unique geochemical signatures. [13] The coral calcareous sponge microbial boundstone aligns with the coral-microbolite-debris reef classification, indicating a high-energy, shallow-water habitat. [12]

Fossil content

Color key
Taxon Reclassified taxonTaxon falsely reported as presentDubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Foraminifera

GenusSpeciesLocationMaterialNotesImages
Agerina [11]
  • A. martana
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Cornuspiridae family
Ammobaculites [11]
  • A. coprolithiformis
  • A. spp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the family Ammomarginulininae.
Bosniella [11]
  • B. oenensis
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Biokovinidae family
Bullopora [11]
  • B. spp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Webbinellinae family
Everticyclammina [11]
  • E. praevirguliana
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Everticyclamminidae family.
Kurnubia [11]
  • K. sp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the family Kurbuniinae
Glomospira [11]
  • G. sp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the family Ammodiscidae.
Haurania [11]
  • H. amiji
  • H. deserta
  • H. ssp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the family Hauraniinae.
Involutina [1]
  • I. farinacciae
  • I. liassica
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Involutinidae family
Lituosepta [1] [11]
  • L. recoarensis
  • L. compressa
  • L. ssp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Mesoendothyridae family.
Mesoendothyra [11]
  • M. sp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Mesoendothyridae family.
Mayncina [11]
  • M. termieri
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Mayncinidae family
Nodosaria [11]
  • N. sexcostata
  • N. sp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the family Nodosariinae.
Orbitopsella [1] [11]
  • O. primaeva
  • O. preacursor
  • O. dubari
  • O. spp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Mesoendothyridae family.
Ophtalmidium [11]
  • O. concentricum
  • O. sp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the family Ophthalmidiidae.
Pseudocyclammina [11]
  • P. liasica
  • P. spp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Pfenderinidae family.
Planisepta [1]
  • P. compressa
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Planiseptinae family.
Reophax [11]
  • R. sp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Reophacinae family
Siphovalvulina [1]
  • S. gibraltarensis
  • S. variabilis
  • S. sp.A
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Pfenderinidae family.
Tetrataxis [11]
  • T. spp.
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Tetrataxidae family.
Trochammina [11]
  • T. sp
  • Bou Dahar
Calcareous SkeletonsA foraminifer of the Trochamminidae family

Sponges

GenusSpeciesStratigraphic positionMaterialNotesImages
Arhoussia [14]
  • A. calyx
  • Jebel Bou Arhous Srhir
Colonial ImprintsA Glass sponge, member of the family Euplectellidae
Chaetetes [15]
  • C. (Pseudoseptifer) zignoi
  • C. (Pseudoseptifer) murciana
  • C. (Pseudoseptifer) liasica
  • Bou Dahar
Colonial ImprintsA Demosponge, member of the family Chaetetinae. Monospecific assamblages with encrusting and symbiont forms
Specimen of the same genus Chaetetes (8109235590).jpg
Specimen of the same genus
Palaeopora [15]
  • P. ramulosa
  • P. liasica
  • Bou Dahar
Colonial ImprintsA Calcareous sponge, Incertade sedis affinities
Stromatomorpha [16]
  • S. californica
  • S. cf. californica
  • S. liassica
  • S. (Cylicopsis) atlantis
  • Bou Dahar
  • IttoFezzou
  • Ait-Ani
Colonial ImprintsA Demosponge, member of the family Anthaspidellidae
Tillichtia [14]
  • T. aedificator
  • T. spp.
  • Kadoussa
Colonial ImprintsA Glass sponge, member of the family Euplectellidae

Hydrozoa

GenusSpeciesStratigraphic positionMaterialNotesImages
Spongiomorpha [15] [17]
  • S. ramosa
  • S. gibbosa
  • S. (Heptastylopsis) sp. 1
  • Bou Dahar
Colonial ImprintsA Hydrozoan, member of the family Spongiomorphidae

Anthozoa

GenusSpeciesStratigraphic positionMaterialNotesImages
Actinastrea [18]
  • A. minima
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Actinastreidae
Allocoeniopsis [18]
  • A. dendroidea
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Actinastreidae
Alloiteausmilia [18]
  • A. boudaharensis
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Zardinophyllidae
Araiophyllum [19]
  • A. liasicum
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Comoseridae
Archaeosmilia [18]
  • A. menchikoffi
  • A. beata
  • A. sp.
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Archaeosmiliidae
Brachyseris [18]
  • B. walfordi
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Latomeandridae
Chomatoseris [18]
  • C. orbulites
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Comoseridae
Cladophyllia [18]
  • C. dresnayi
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Cladophylliidae
Coenastraea [18]
  • C. cf.maccoyi
  • C. sp.
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Actinastreidae
Cyclophyllopsis [18]
  • C. cornutiformis
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Cyclophyllopsiidae
Dimorphastraea [18]
  • D. menchikoffi
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Latomeandridae
Eocomoseris [19]
  • E. minima
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Comoseridae
Epismilia [18] [19]
  • E. mauretaniensis
  • E.? sp.  
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Zardinophyllidae
Epismiliopsis [18]
  • E. liasica
  • E. sp.
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Zardinophyllidae
Epistreptophyllum [18]
  • E. vetusum
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Dermosmiliidae
Epistreptophyllum specimen (from Germany) Epistreptophyllum commune.jpg
Epistreptophyllum specimen (from Germany)
Funginella [18]
  • F. domeriensis
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Funginellidae
Goniocora [18]
  • G. concinna
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Stylinidae
Goniocora specimens Goniocora pumila, Sl. Gottwald.jpg
Goniocora specimens
Haplaraea [18]
  • H. spp.
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Astraraeidae
Margarosmilia [18] [19]
  • M. gemminata
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Margarophylliidae
Margarosmilia specimens Margarosmilia confluens - MUSE.jpg
Margarosmilia specimens
Mesomorpha [18]
  • M. gracilis
  • M. bussoni
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Thamnasteriidae
Neorylstonia [20]
  • N. pseudocolumellata
  • N. spp.
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Volzeioidea
Omphalophylliopsis [20]
  • O. lobatus
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Conophyllidae
Plesiophyllum [18]
  • P. cylindricum
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Zardinophyllidae
Proleptophyllia [18]
  • P. multiradiata
  • P. rejuvenescens
  • P. subcylindrica
  • P. bilobata
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Acrosmiliidae
Rodinosmilia [18] [19]
  • R. elegantula
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Pachycoeniidae
Stylosmilia [19]
  • S. dresnayi
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Stylinidae
Stylophyllopsis [18] [19]
  • S. vetusum
  • S. cf. rudis
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Stylophyllidae
Trigerastrea [18]
  • T. minima
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Latomeandridae
Volzeia [18]
  • V. badiotica
  • Bou Dahar
Imprints & isolated specimensA Stony coral member of the family Volzeiidae

Brachiopoda

GenusSpeciesStratigraphic positionMaterialNotesImages
Aulacothyris [21] [22]
  • A. semiplana
  • Bou Dahar
Isolated ShellsA Terebratulidan, member of the family Zeilleriidae
Dictyothyris [23]
  • D. drepanensis
  • Bou Dahar
Isolated ShellsA Terebratulidan, member of the family Dictyothyrididae
Fimbriothyris [22]
  • F. itoensis
  • F. tenuiplicata
  • F. brevicostata
  • Bou Dahar
Isolated ShellsA Terebratulidan, member of the family Zeilleriidae
Linguithyris [24]
  • L. aspasia
  • L. adnethensis
  • Bou Dahar
Isolated ShellsA Terebratulidan, member of the family Nucleatidae
Rhynchonella [21] [22]
  • R. quadrata
  • R. spp.
  • Bou Dahar
Isolated ShellsA Rhynchonellida n, member of the family Rhynchonellidae
Prionorhynchia [25]
  • P. msougari
  • P. subcynocephala
  • P. rubriaxensis
  • Bou Dahar
Isolated ShellsA Rhynchonellida n, member of the family Wellerellidae
Prototegulithyris [26]
  • P. khadijae
  • Bou Dahar
Isolated ShellsA Terebratulidan, member of the family Dictyothyrididae
Pseudogibbirhynchia [27]
  • P. jurensis
  • P. moorei
  • P. waehneri
  • Bou Dahar
Isolated ShellsA Rhynchonellida n, member of the family Basiliolidae
Stolmorhynchia [28]
  • S. limata
  • Bou Dahar
Isolated ShellsA Rhynchonellida n, member of the family Wellerellidae
Suessia [21] [22]
  • S. liasina
  • Bou Dahar
Isolated ShellsA Terebratulida n, member of the family Zeilleriidae

Gastropoda

GenusSpeciesStratigraphic positionMaterialNotesImages
Actaeonina [29]
  • A. radius
  • Bou Dahar N7
  • Bou Dahar "Carriere"
Isolated ShellsAn Architectibranchian, member of the family Acteonidae
Amphitrochilia [29]
  • A. (Aristerella) undata
  • A. (Aristerella) rorata
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N5
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Trochoidean, member of the family Proconulidae
Ampullina [29]
  • A. (Ampullospira) dominans
  • A. sp. nov
  • A. (Pseudamura) sp. nov
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N6
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Caenogastropodan, member of the family Ampullinidae
Ampullina specimens Ampullina parisiensis 01.jpg
Ampullina specimens
Aphanoptyxis [29]
  • A. cossmanni
  • Bou Dahar N7
  • Bou Dahar N9
Isolated ShellsA Nerineoidean, member of the family Ceritellidae
Aptyxiella [29]
  • A. praevia
  • A. fimbriata
  • A. subgradata
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N5
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Nerineoidean, Incertade sedis
Ataphrus [29]
  • A. cf.piettei
  • A. (Pleuratella) cf.normaniensis
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar "Carriere"
  • Tizi N´Oumzour
Isolated ShellsA Trochoidean, member of the family Ataphridae
Boehmia [29]
  • B. speciosa
  • B. costata
  • Bou Dahar N3
  • Bou Dahar N6
Isolated ShellsA Architectibranchian, member of the family Acteonidae
Brachytrema [29]
  • B. despujolsi
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N5
  • Bou Dahar N7
  • Bou Dahar N12
Isolated ShellsA Caenogastropodan, member of the family Brachytrematidae
Buvignieria [29]
  • B. sp.
  • Bou Dahar
Isolated ShellsA Littorinimorphan, member of the family Palaeorissoinidae
Cerithiella [29]
  • C. africana
  • C. sarrazini
  • C. aff.nudae
  • Bou Dahar N1
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar N8
  • Bou Dahar "Carriere"
Isolated ShellsA Triphoroidean, member of the family Newtoniellidae
Cerithiella specimen Cerithiella austrina 001.jpg
Cerithiella specimen
Cerithinella [29]
  • C. termieri
  • C. cf.italica
  • C. gratiosa
  • C. miliare
  • C. gemmifera
  • C. margaritata
  • C. baccata
  • C. cf.baccata
  • C. aff.cerithiiformis
  • C. (Laevibaculus) sp.
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N5
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N11
  • Bou Dahar N12
Isolated ShellsA Caenogastropodan, member of the family Procerithiidae
Climacina [29]
  • C. daharensis
  • C. cf.mariae
  • C. sp.
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N7
Isolated ShellsA Littorinimorphan, member of the family Tornidae
Climacina [29]
  • C. daharensis
  • C. cf.mariae
  • C. sp.
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N7
Isolated ShellsAn Architectibranchian, member of the family Cylindrobullinidae
Coelostylina [29]
  • C. chartroni
  • C. choffati
  • C. rigida
  • C. (Cosmostylina) darestei
  • C. (Cosmostylina) lima
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N6
  • Bou Dahar N7
  • Tizi N´Oumzour
  • Aguelmane-Azougar
Isolated ShellsA Caenogastropodan, member of the family Coelostylinidae
Crossostoma [29]
  • C. atlantis
  • Bou Dahar N7
  • Tizi N´Oumzour
Isolated ShellsA Trochoidean, member of the family Colloniidae
Cryptaulax [29]
  • C. angulodentatum
  • C. album
  • C. cf.contortum
  • C. cf.densestritum
  • C. aff.tetrataeniatum
  • C. cf.tetrataeniatum
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N11
Isolated ShellsA Cerithiimorphan, member of the family Cryptaulacinae
Diarthema [29]
  • D. sp.
  • Bou Dahar N3
Isolated ShellsA Littorinimorphan, member of the family Aporrhaidae
Diatinostoma [29]
  • D. sp.
  • Tizi N´Oumzour
Isolated ShellsA Caenogastropodan, member of the family Eustomatidae
Dicroloma [29]
  • D. sp.
  • Bou Dahar N7
Isolated ShellsA Littorinimorphan, member of the family Aporrhaidae
Discocirrus [29]
  • D. crispus
  • D. armatus
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N5
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Seguenziidan, member of the family Cirridae
Discohelix [29]
  • D. (Colpomphalus) spledens
  • D. (Colpomphalus) cf.glauca
  • D. (Colpomphalus) sp. A
  • Bou Dahar N3
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Seguenziidan, member of the family Eucyclidae
Discohelix specimen Discohelix tunisiensis apical.jpg
Discohelix specimen
Exelissa [29]
  • E. pusilla
  • E. douxamii
  • E. spissa
  • E. galenae
  • E. menchikoffi
  • E. cf.prealpina
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N8
  • Bou Dahar N9
  • Bou Dahar N12
  • Ait Daoud-ou-Azzi
Isolated ShellsA Cerithiimorphan, member of the family Cryptaulacinae
Emarginula [29]
  • E. henrici
  • E. cf.orthogonia
  • E. (Rimula) pileata
  • E. (Rimula) sp. A
  • E. (Rimula) sp. B
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
  • Aït Daoud
Isolated ShellsA Lepetellidan, member of the family Fissurellidae
Emarginula specimens Emarginula phrygium (10.3897-afrinvertebr.61.51989) Figure 8.jpg
Emarginula specimens
Eucyclus [29]
  • E. lateclathratus
  • Bou Dahar N3
Isolated ShellsA Seguenziidan, member of the family Eucyclidae
Fibulella [29]
  • F. afra
  • F. nicolai
  • F. sp.
  • Bou Dahar, all the levels
Isolated ShellsA Nerineoidean, member of the family Ceritellidae
Gymnocerithium [29]
  • G. sp.
  • Bou Dahar "Carriere"
Isolated ShellsA Caenogastropodan, member of the family Campanilidae
Hesperidina [29]
  • H. striolata
  • Bou Dahar N7
  • Tizi N´Oumzour
  • Imi n´Ikis
Isolated ShellsA Caenogastropodan, member of the family Purpurinidae
Nerinella [29]
  • N. marucchiensis
  • N. cayeuxi
  • N. cf.atava
  • N. convoluta
  • N. sp.
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N8
  • Bou Dahar N11
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Nerineoidean, member of the family Nerinellidae
Neritodomus [29]
  • N. calaminae
  • N. (Sphaerochilus) daharensis
  • N. cf.tethys
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar "Carriere"
  • Bou Dahar Plateau
  • Tizi N´Oumzour
Isolated ShellsA Cycloneritida, member of the family Neridomidae
Neritopsis [29]
  • N. falloti
  • N. sp.
  • Bou Dahar N2
  • Bou Dahar "Carriere"
Isolated ShellsA Cycloneritida, member of the family Neritopsidae
Neritopsis specimens Neritopsis atlantica (MNHN-IM-2012-18255).jpeg
Neritopsis specimens
Ovacteonina [29]
  • O. sp.
  • Bou Dahar N7
Isolated ShellsAn Architectibranchian, member of the family Acteonidae
Paracerithium [29]
  • P. lacostei
  • P. echinatum
  • P. liasinum
  • P. merciaii
  • P. ingens
  • P. angulosum
  • P. cf.strueveri
  • P. cf.multiforme
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N11
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Caenogastropodan, member of the family Batillariidae
Pileolus [29]
  • P. laevis
  • Bou Dahar, all the levels
  • Tizi N´Oumzour
Isolated ShellsA Cycloneritida, member of the family Pileolidae
Platyacra [29]
  • P. (Asperilla) russoi
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N12
  • Amellago
Isolated ShellsA Trochoidean, member of the family Angariidae
Pleurotomaria [29]
  • P. cf.pisana
  • Bou Dahar
Isolated ShellsA Pleurotomariidan, member of the family Pleurotomariidae
Pleurotomaria specimens Pleurotomaria delagei (MNHN-IM-2000-4940).jpeg
Pleurotomaria specimens
Procerithium [29]
  • P. canescens
  • P. curtum
  • P. sp.
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar N12
Isolated ShellsA Caenogastropodan, member of the family Procerithiidae
Proconulus [29]
  • P. spinatus
  • P. sp. A
  • P. sp. B
  • Bou Dahar N3
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Trochoidean, member of the family Proconulidae
Protorcula [29]
  • P. fischeri
  • Bou Dahar N2
  • Bou Dahar N7
  • Bou Dahar N8
Isolated ShellsA Pseudozygopleuroidea, member of the family Protorculidae
Pseudomelania [29]
  • P. aristomacha
  • P. dealbata
  • P. chrysallis
  • P. (Gardetia) chouberti
  • P. (Oonia) marucchiensis
  • P. (Oonia) cf.turgidula
  • P. (Oonia) aff.pupoideae
  • P. (Oonia) sp.
  • P. (Rhabdoconcha) tenuestriata
  • P. cf.procera
  • P. cf.eulimoides
  • P. cf.rhea
  • P. cf.globosa
  • P. sp.
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N8
  • Bou Dahar N12
  • Bou Dahar "Carriere"
  • Tizi N´Oumzour
Isolated ShellsA Caenogastropodan, member of the family Pseudomelaniidae
Pseudomelania specimen Pseudomelania shell cropped.png
Pseudomelania specimen
Pseudonerinea [29]
  • P. terebra
  • P. liasica
  • P. sp. A
  • P. sp.
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N5
  • Bou Dahar N6
  • Bou Dahar "Carriere"
Isolated ShellsA Nerineoidean, member of the family Pseudonerineidae
Purpurina [29]
  • P. rudis
  • P. moghrabiensis
  • P. patroclus
  • P. berberica
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
  • Kasba d´Amellago
  • Tizi N´Oumzour
Isolated ShellsA Caenogastropodan, member of the family Purpurinidae
Example of Purpurina specimen Purpurina serrata side 1 CRF.jpg
Example of Purpurina specimen
Rigauxia [29]
  • R. baculus
  • Bou Dahar N2
  • Bou Dahar N7
  • Bou Dahar "Carriere"
Isolated ShellsA Streptacidoidea, Incertade sedis
Scurriopsis [29]
  • S. altissima
  • S. parma
  • S. (Scurria?) cf.nana
  • S. (Scurria?) sp.
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
  • Aït Daoud
  • El-Kansera
Isolated ShellsA Lottioidean, member of the family Acmaeidae
Scurria specimens Scurria variabilis 01.jpg
Scurria specimens
Teliochilus [29]
  • T. cf.deslongchampsi
  • T. clathratus
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
Isolated ShellsA Cerithiimorphan, member of the family Cryptaulacinae
Tectus [29]
  • T. (Dimorphotectus) chouberti
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar "Carriere"
Isolated ShellsA Trochoidean, member of the family Tegulidae
Tectus specimens Tectus niloticus 01.JPG
Tectus specimens
Teinostoma [29]
  • T. sp.
  • Bou Dahar N2
  • Bou Dahar "Carriere"
Isolated ShellsA Littorinimorphan, member of the family Teinostomatidae
Teinostoma specimens Teinostoma morlierei (MNHN-IM-2000-31235).jpeg
Teinostoma specimens
Trochotoma [29]
  • T. (Ditremaria) secans
  • T. (Ditremaria) pachyspira
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Pleurotomariidan, member of the family Trochotomidae
Tylostoma [29]
  • T. maurum
  • Bou Dahar N7
  • Bou Dahar "Carriere"
Isolated ShellsA Caenogastropodan, member of the family Tylostomatidae
Tylostoma specimen Tylostoma.jpg
Tylostoma specimen
Zygopleura [29]
  • Z. pectita
  • Z. appenninica
  • Z. paradisi
  • Z. tatia
  • Z. menchikoffi
  • Z. sp.
  • Bou Dahar N1
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Pseudozygopleuroidea, member of the family Zygopleuridae
Example of Zygopleura specimen Zygopleura corvaliana.jpg
Example of Zygopleura specimen

Polyplacophora

GenusSpeciesStratigraphic positionMaterialNotesImages
Allochiton [30]
  • A. sp. nov?
  • Bou Dahar
Isolated ShellsA Chitonidan, member of the family Mopaliidae
Heterochiton [30]
  • H. cf.giganteus
  • H. cf.zitteli
  • H. sp. nov?
  • Bou Dahar
Isolated ShellsA Chitonidan, member of the family Mopaliidae

Bivalvia

GenusSpeciesStratigraphic positionMaterialNotesImages
Anisocardia [31]
  • A. quadrata
  • Bou Dahar N4
Isolated ShellsAn Veneridan, member of the family Arcticidae
Astarte [31]
  • A. vendaeensis
  • A. marcaisi
  • A. (Theveninia) aff. tetragona
  • A. aff.interlineata
  • A. sp.
  • Bou Dahar N3
  • Bou Dahar N5
  • Bou Dahar N6
  • Bou Dahar N12
  • Bou Dahar Sup
Isolated ShellsA Carditidan, member of the family Astartidae
Astarte specimens Astarte elliptica 01.jpg
Astarte specimens
Barbatia [31]
  • B. metellaria
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N5
  • Bou Dahar N7
Isolated ShellsAn Arcidan, member of the family Arcidae
Barbatia specimens Barbatia amygdalumtostum 02.jpg
Barbatia specimens
Cardium [31]
  • C. pustulatum
  • C. galenae
  • C. (Protocardia) praecalva
  • C. sp.
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N7
  • Tizi N´Oumzour
Isolated ShellsA Pectinidan, member of the family Pectinidae
Cardium specimens Cardiidae - A. tuberculata (Europe); D. muricata (America).jpg
Cardium specimens
Camptonectes [31]
  • C. sp.
  • Bou Dahar "Carriere"
Isolated ShellsA Cardiidan, member of the family Cardiidae
Chlamys [31]
  • C. aff.uhligi
  • Bou Dahar Sup
Isolated ShellsA Pectinidan, member of the family Pectinidae
Cochlearites [1] [9]
  • C. loppianus
  • Bou Dahar "Carriere"
  • SE Azou
  • N Aït Oufella
  • Foum Kheneg
  • Tizi N´Oumzour
Isolated ShellsAn Ostreidan, member of the family Plicatostylidae
Fimbria [31]
  • F. (Corbis) alveolus
  • F. (Corbis) cochlearis
  • F. (Corbis) trulla
  • F. (Corbis) sp.
  • Bou Dahar N1
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Lucinidan, member of the family Lucinidae.
Fimbria specimens (from Hawaii) Fimbria fimbriata 02.jpg
Fimbria specimens (from Hawaii)
Gervillia [31]
  • G. (Gervilleia) spinicosta
  • G. (Gervilleia) sp.
  • Bou Dahar "Carriere"
  • Toutia
  • Foum Kheneg
Isolated ShellsAn Ostreidan, member of the family Bakevelliidae
Gervilleioperna [1] [31]
  • G. atlantis
  • G. buchi
  • G. termieri
  • G. timorensis
  • G. sp.
  • Bou Dahar "Carriere"
  • SE Azou
  • N Aït Oufella
  • Foum Kheneg
  • Tizi N´Oumzour
Isolated ShellsAn Ostreidan, member of the family Plicatostylidae
Lima [31]
  • L. (Plagiostoma) eucharis
  • Lima (Limea) aff.juliana
  • L. (Ctenostreon) cf.raricostata
  • L. (Ctenostreon) sp.
  • Bou Dahar N3
  • Bou Dahar "Carriere"
Isolated ShellsA Limidan, member of the family Limidae
Lima specimens Lima (YPM IZ 049863).jpeg
Lima specimens
Lithioperna [1] [32]
  • L. (Lithiopedalion) kuehni
  • L. scutata
  • L. spp.
  • Bou Dahar "Carriere"
  • SE Azou
  • N Aït Oufella
  • Foum Kheneg
  • Tizi N´Oumzour
Isolated ShellsAn Ostreidan, member of the family Plicatostylidae
Lucina [31]
  • L. (Phacoides) lunulicava
  • L. (Phacoides) laevis
  • L. (Phacoides) cf.zonaria
  • L. (Phacoides) sp. nov
  • Bou Dahar N2
  • Bou Dahar N4
  • Bou Dahar N7
  • Bou Dahar "Carriere"
  • Aguelmane-Azougar
Isolated ShellsA Lucinidan, member of the family Lucinidae.
Lucina specimens Lucina aurantia 01.jpg
Lucina specimens
Isoarca [31]
  • I. ssp.
  • Bou Dahar
Isolated ShellsA Nuculanidan, member of the family Isoarcidae
Isocyprina [31]
  • I. pulla
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar "Carriere"
Isolated ShellsA Veneridan, member of the family Isocyprinidae
Myoconcha [31]
  • M. linguaeformis
  • M. (Daharina) gentili
  • M. aff.corniculun
  • Bou Dahar N3
  • Bou Dahar N7
  • Bou Dahar "Carriere"
Isolated ShellsAn Imparidentian, member of the family Kalenteridae
Mytilus [31]
  • M. maurus
  • M. aff.furcatus
  • M. (Pachymytilus) superbus
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsA Mytilidan, member of the family Mytilidae
Mytilus specimens Mytilus edulis var. flavida 01.jpg
Mytilus specimens
Opis [31]
  • O. alata
  • O. caudata
  • O. bicristata
  • Bou Dahar
  • Aguelmane-Azougar
Isolated ShellsA Carditidan, member of the family Astartidae.
Opisoma [31]
  • O. excavatum
  • O. menchikoffi
  • O. curvidens
  • O. scalprum
  • O. rex
  • O. bourcarti
  • O. spp.
  • Bou Dahar N2
  • Bou Dahar N3
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N8
  • Bou Dahar N12
  • Bou Dahar "Carriere"
  • Khorchef
Isolated ShellsA Carditidan, member of the family Astartidae. Is considered a genus that evolved from shallow burrowing ancestors, becoming a secondarily semi-infaunal edgewise recliner adapted to photosymbiosis. [33]
Pachyerisma [31]
  • P. opimum
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar N7
Isolated ShellsA Megalodontidan, member of the family Megalodontidae
Pachymegalodon [31]
  • P. timorense
  • Bou Dahar "Carriere"
Isolated ShellsA Megalodontidan, member of the family Megalodontidae
Parallelodon [31]
  • P. (Beushausenia) daharensis
  • P. (Beushausenia) sp.
  • Bou Dahar N1
  • Bou Dahar N3
  • Bou Dahar N4
  • Bou Dahar "Carriere"
Isolated ShellsAn Arcidan, member of the family Parallelodontidae
Perna [31]
  • P. scutata
  • P. sp. nov
  • P. sp.
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar N12
  • Bou Dahar "Carriere"
Isolated ShellsAn Ostreidan, member of the family Pteriidae
Perna specimens Perna perna 06.jpg
Perna specimens
Pecten [31]
  • P. (Aequipecten) julianus
  • P. (Aequipecten) norigliensis
  • P. aff.bersaskensis
  • P. spp.
  • Bou Dahar N1
  • Bou Dahar N3
  • Bou Dahar N12
  • Bou Dahar "Carriere"
  • Foum Kheneg
Isolated ShellsA Pectinidan, member of the family Pectinidae
Pecten specimens Pecten maximus 03.jpg
Pecten specimens
Plicatula [31]
  • P. daharensis
  • Bou Dahar N3
  • Bou Dahar N6
  • Bou Dahar N7
  • Bou Dahar "Carriere"
Isolated ShellsA Pectinidan, member of the family Plicatulidae
Plicatula specimens Plicatula angolensis (MNHN-IM-2000-4055).jpeg
Plicatula specimens
Pinna [31]
  • P. (Pinnigena) sp.
  • Bou Dahar
Isolated ShellsAn Ostreidan, member of the family Pinnidae
Pinna specimens Pinna carnea 000.jpg
Pinna specimens
Pseudotrapezium [31]
  • P. spp.
  • Bou Dahar N7
Isolated ShellsA Veneridan, member of the family Trapezidae
Spondylus [31]
  • S. numidus
  • Bou Dahar Sup
Isolated ShellsA Pectinidan, member of the family Spondylidae
Spondylus specimens Spondylus variegatus 03.jpg
Spondylus specimens

Ammonites

GenusSpeciesStratigraphic positionMaterialNotesImages
Apoderoceras [34]
  • A. davoei
  • A. sp.
  • Bou Dahar N1
Isolated ShellsAn Ammonite, member of the family Coeloceratidae
Apoderoceras specimen Apoderoceras FMNH.jpg
Apoderoceras specimen
Dactylioceras [34]
  • D. tenuicostatum
  • Bou Dahar N1
Isolated ShellsAn Ammonite, member of the family Dactylioceratidae
Dactylioceras reconstruction Dactylioceras NT.jpg
Dactylioceras reconstruction
Hildoceratoides [34]
  • H. pectinatum
  • Bou Dahar N7
Isolated ShellsAn Ammonite, member of the family Hildoceratidae
Juraphyllites [1]
  • J. libertus
  • J. limatus
  • J. (Harpophylloceras) eximius
  • Bou Dahar
Isolated ShellsAn Ammonite, member of the family Juraphyllitidae
Paltechioceras [1]
  • P. boehmi
  • Bou Dahar
Isolated ShellsAn Ammonite, member of the family Echioceratidae
Tropidoceras [9]
  • T. flandrini
  • Bou Dahar N4
Isolated ShellsAn Ammonite, member of the family Polymorphitidae

Annelida

GenusSpeciesStratigraphic positionMaterialNotesImages
Serpula [35]
  • S. spp.
  • Bou Dahar
Tubiform StructuresA Serpulidan, member of the family Serpulidae
Serpula specimen Serpula columbiana.jpg
Serpula specimen

Echinodermata

GenusSpeciesStratigraphic positionMaterialNotesImages
Apiocrinus [36]
  • A. amalthei
  • Bou Dahar
StemsA Crinoid, member of the family Apiocrinidae
Atlasaster [37]
  • A. termieri
  • Bou Dahar
SpecimensAn Echinoidean, member of the group Irregularia
Diademopsis [37]
  • D. behtensis
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, member of the family Pedinidae
Diademopsis specimen Diademopsis micropora FOS 2400 - 1.JPG
Diademopsis specimen
Diplechinus [37]
  • D. hebbriensis
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, member of the family Stomechinidae
Diplocidaris [36] [37]
  • D. menchikoffi
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, member of the family Diplocidaridae
Diplocidaris specimen Diplocidaridae - Diplocidaris gigantea.JPG
Diplocidaris specimen
Firmacidaris [36] [37]
  • F. precincta
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, incertae sedis
Hemicidaris [36] [37]
  • H. termieri
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, member of the family Hemicidaridae
Hemicidaris specimen Hemicidaris crenularis FOS 2333 - 1.jpg
Hemicidaris specimen
Microdiadema [36] [37]
  • M. richeri
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, member of the family Pseudodiadematidae
Miocidaris [37]
  • M. dubari
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, member of the family Miocidaridae
Pseudopedina [37]
  • P. atlantis
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, member of the family Pedinidae
Pygaster [37]
  • P. daguini
  • Bou Dahar
Specimens & Isolated spinesAn Echinoidean, member of the family Pygasteridae

Microproblematic/Algae

GenusSpeciesLocationMaterialNotesImages
Aeolissacus [11]
  • A. spp.
  • Bou Dahar
Calcified ThalliA Possible Green Algae of the family Dasycladaceae.
Baccanella [11]
  • B. floriformis
  • Bou Dahar
Calcified specimensIncertade sedis
Bacinella [11]
  • B. irregularis
  • B. ordinata
  • Bou Dahar
Calcified specimensA Cyanobacterial Alga of the family Garwoodiaceae
Boueina [16]
  • B. hochstetteri
  • Bou Dahar
  • IttoFezzou
  • Ait-Ani
Calcified specimensA Green Algae of the family Halimedaceae.
Cayeuxia [11]
  • C. liasica
  • C. spp.
  • Bou Dahar
Calcified ThalliA green algae of the Halimedaceae or Udoteaceae family.
Modern Udotea Udotea conglutinata as Corallina conglutinata in Ellis & Solander 1786.jpg
Modern Udotea
Lithocodium [11]
  • L. aggregatum
  • Bou Dahar
Calcified specimensIncertade sedis
Muranella [11]
  • M. sphaerica
  • Bou Dahar
Calcified specimensIncertade sedis
Neomizzia [38]
  • N. elongata
  • Bou Dahar
Calcified specimensA Green Algae of the family Dasycladaceae.
Considered to be convergent with modern Cymopolia Cymopolia barbata in Harvey 1858.jpg
Considered to be convergent with modern Cymopolia
Thaumatoporella [11]
  • T. parvovesiculifera
  • Bou Dahar
Calcified ThalliA Green alga of the Thaumatoporellales group  
Palaeodasycladus [11]
  • P. fragilis
  • P. mediterraneus
  • P. dolomiticus
  • P. anae
  • P. spp.
  • Bou Dahar
Calcified ThalliA Green Algae of the family Dasycladaceae. A reefal algae usually found in carbonate settings along all the Mediterranean
Radiomura [11]
  • R. cautica
  • Bou Dahar
Calcified specimensIncertade sedis
Rivularia [11]
  • R. moesica
  • Bou Dahar
Calcified specimensA Cyanobacterial Alga of the family Rivulariaceae
Rivularia specimens Rivularia bullata.jpg
Rivularia specimens
Solenopora [11] [17]
  • S. liasica
  • Bou Dahar
Calcified specimensA Red Alga of the family Solenoporaceae
Tersella [39]
  • T. genotii
  • Bou Dahar
Calcified specimensA Green Algae of the family Dasycladaceae.
Toulaina [16]
  • T. liassica
  • Bou Dahar
  • IttoFezzou
  • Ait-Ani
Calcified specimensA Green Algae of the family Halimedaceae.


See also

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<span class="mw-page-title-main">Tafraout Group</span> Geological formations in Morocco

The Tafraout Group is a geological group of formations of Toarcian-Aalenian age in the Azilal, Béni-Mellal, Imilchil, Zaouiat Ahansal, Ouarzazate, Tinerhir and Errachidia areas of the High Atlas of Morocco. The Group represents the remnants of a local massive Siliciclastic-Carbonate platform, best assigned to succession W-E of alluvial environment occasionally interrupted by shallow marine incursions and inner platform to open marine settings, and marks a dramatic decrease of the carbonate productivity under increasing terrigenous sedimentation. Fossils include large reef biotas with richness in "lithiotid" bivalves and coral mounts, but also by remains of vertebrates such as the sauropod Tazoudasaurus and the basal ceratosaur Berberosaurus, along with several undescribed genera. While there have been attributions of its lowermost layers to the Latest Pliensbachian, the current oldest properly measured are part of the Earliest Toarcian regression ("MRST10"), part of the Lower-Middle Palymorphum biozone. This group is composed of the following units, which extend from west to east: the Azilal Formation ; the "Amezraï Formation" ; the Tafraout Formation the Tagoudite Formation & maybe the "Aït Athmane Formation". They are connected with the deeper shelf deposits of the Agoudim 1 Formation. Overall, this group represents a mixed carbonate-siliciclastic system of several hundred meters thick, dominated by deposits of shallow marine platforms linked to a nearby hinterland dominated by conglomerates. The strata of the group extend towards the central High Atlas, covering different anticlines and topographic features along the mountain range.

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