Greenish warbler | |
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nominate race P. trochiloides trochiloides adult from Pangolakha Wildlife Sanctuary, Sikkim, India | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Passeriformes |
Family: | Phylloscopidae |
Genus: | Phylloscopus |
Species: | P. trochiloides |
Binomial name | |
Phylloscopus trochiloides (Sundevall, 1837) | |
Subspecies | |
5, and see text | |
Range of P. trochiloides Breeding Passage Non-breeding |
The greenish warbler (Phylloscopus trochiloides) is a widespread leaf warbler with a breeding range in northeastern Europe, and temperate to subtropical continental Asia. This warbler is strongly migratory and winters in India. It is not uncommon as a spring or early autumn vagrant in Western Europe and is annually seen in Great Britain. In Central Europe large numbers of vagrant birds are encountered in some years; some of these may stay to breed, as a handful of pairs does each year in Germany. [2]
Like all leaf warblers, it was formerly placed in the "Old World warbler" assemblage, but now belongs to the new leaf-warbler family Phylloscopidae. [3] The genus name Phylloscopus is from Ancient Greek phullon, "leaf", and skopos, "seeker" (from skopeo, "to watch"). The specific trochiloides is from Ancient Greek trokhalos, "bowed", and -oides "resembling", from the similarity to the willow warbler, P. trochilus. [4] The English name of this species provides a perfect argument in favour of the capitalisation of species names (i.e. treating them as proper nouns), a convention which is generally applied in scientific literature. The decapitalised "greenish warbler" is equally descriptive of many bird species across multiple families, whereas a capitalised "Greenish Warbler" shows unambiguously that Phylloscopus trochiloides is under discussion.
This is a typical leaf warbler in appearance, grayish-green above and off-white below. The single wing bar found in the southern and western populations distinguishes them from most similar species (except Arctic warbler P. borealis). It is slightly smaller than that species and has a thinner bill, without a dark tip to the lower mandible. A latitude-based analysis of wintering birds indicated that more northerly P. trochiloides are smaller, i.e. this species does not seem to follow Bergmann's rule. [5]
Its song is a high jerky trill, in some populations containing a sequence of down- and more rarely up slurred notes.
It breeds in lowland deciduous or mixed forest; non-breeding birds in the warmer parts of its range may move to montane habitat in summer. Individuals from southeast of the Himalayas are for example quite often seen in Bhutan during the hot months, typically in humid Bhutan Fir (Abies densa) forest up to about 3,800 meters ASL or more, but they do not breed there and return again to the adjacent subtropical lowlands in winter. [6]
The nest is on the ground in low shrub. Like its relatives, this small passerine is insectivorous.
It has a number of subspecies, of which P. t . viridianus is the most familiar in Europe. As it seems, it is a ring species, with populations diverging east- and westwards of the Tibetan Plateau, later meeting on the northern side. Their relationships are therefore fairly confusing: [7]
The groups' origin lies probably in the Himalayan region, where trochiloides is found. This taxon is close to the parapatric obscuratus, and to plumbeitarsus which is geographically separated from obscuratus; they all can (and in the case of the former two do naturally) hybridize. P. t. plumbeitarsus is often split as distinct species, as it does not hybridize with viridianus in the narrow zone in the western Sayan Mountains where their ranges overlap.
But phylogenetically, the western taxa are even more distinct. However, there is some gene flow between trochiloides and viridianus also, with their hybrids being especially common in Baltistan; which are now considered as a distinct subspecies ludlowi. The green warbler P. nitidus, now considered as a distinct species, is a mountain isolate that diverged from ancestral viridianus.
Song structure differs mainly between greenish warbler and two-barred warbler, which was formerly considered conspecific. The former has a fairly uniform, long, and warbling song. Around the Himalayas, song structure is similar, but songs are generally shorter. Two-barred warbler, on the other hand, has a long song that can be clearly divided into a warbling part, followed by a series of up- and downslurred notes. The songs of obscuratus and ludlowi, are short, but contain the downslur elements too; in the latter, they uniquely appear at the start of the song. [8]
Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. They are not closely related to the New World warblers. The family held over 400 species in over 70 genera, and were the source of much taxonomic confusion. Two families were split out initially, the cisticolas into Cisticolidae and the kinglets into Regulidae. In the past ten years they have been the subject of much research and many species are now placed into other families, including the Acrocephalidae, Cettiidae, Phylloscopidae, and Megaluridae. In addition some species have been moved into existing families or have not yet had their placement fully resolved. A smaller number of warblers, together with some babblers formerly placed in the family Timaliidae and the parrotbills, are retained in a much smaller family Sylviidae.
Leaf warblers are small insectivorous passerine birds belonging to the genus Phylloscopus.
The common chiffchaff, or simply the chiffchaff, is a common and widespread leaf warbler which breeds in open woodlands throughout northern and temperate Europe and the Palearctic.
The willow warbler is a very common and widespread leaf warbler which breeds throughout northern and temperate Europe and the Palearctic, from Ireland east to the Anadyr River basin in eastern Siberia. It is strongly migratory, with almost all of the population wintering in sub-Saharan Africa.
The Arctic warbler is a widespread leaf warbler in birch or mixed birch forest near water throughout its breeding range in Fennoscandia and the northern Palearctic. It has established a foothold in North America, breeding in Alaska. This warbler is strongly migratory; the entire population winters in southeast Asia. It therefore has one of the longest migrations of any Old World insectivorous bird.
The green warbler, also known as green willow warbler or green leaf warbler, is a leaf warbler found in the Caucasus Mountains in southeastern Europe.
Pallas's leaf warbler or Pallas's warbler, is a bird that breeds in mountain forests from southern Siberia east to northern Mongolia and northeast China. It is named after the German zoologist Peter Simon Pallas, who first formally described it. This leaf warbler is strongly migratory, wintering mainly in south China and adjacent areas of southeast Asia, although in recent decades increasing numbers have been found in Europe in autumn.
The yellow-browed warbler is a leaf warbler which breeds in the east Palearctic. This warbler is strongly migratory and winters mainly in tropical South Asia and South-east Asia, but also in small numbers in western Europe. Like the rest of Phylloscopidae, it was formerly included in the Old World warbler assemblage.
Hume's leaf warbler or Hume's warbler is a small leaf warbler which breeds in the mountains of inner Asia. This warbler is migratory and winters mainly in India.
Radde's warbler is a leaf warbler which breeds in Siberia. This warbler is strongly migratory and winters in Southeast Asia. The genus name Phylloscopus is from Ancient Greek phullon, "leaf", and skopos, "seeker". The specific schwarzi commemorates German astronomer Ludwig Schwarz (1822–1894).
The western Bonelli's warbler is a warbler in the leaf warbler genus Phylloscopus. It was formerly regarded as the western subspecies of a wider "Bonelli's warbler" species, but as a result of modern taxonomic developments, they are now usually considered to be two species:
The Eastern Bonelli's warbler, sometimes known as Balkan warbler, is a "warbler" in the leaf warbler genus Phylloscopus. It was formerly regarded as the eastern subspecies of a wider "Bonelli's warbler" species, but as a result of modern taxonomic developments, they are now usually considered to be two species:
Tickell's leaf warbler is a leaf warbler found in Asia in the countries of Bangladesh, Bhutan, China, India, Myanmar, Nepal, Pakistan and Thailand. The species has a yellowish underside and supercilium. Like other leaf warblers it feeds mostly on insects by gleaning and short sallies. An active bird, it prefers the canopy and low shrubbery and can be difficult to track as it moves actively from branch to branch, acrobatically exploring the underside of leaves and twigs. The clear yellowish undersides and lack of a wing bar can be used to tell it apart from similar species. It has slim dark legs with largely pale lower mandible and grayish wing panel.
The eastern crowned warbler is a species of Old World warbler in the family Phylloscopidae. It inhabits boreal and temperate forests in the east Palearctic.
The Emei leaf warbler is a species of leaf warbler. It was formerly included in the "Old World warbler" assemblage.
Tytler's leaf warbler is a songbird species. Like all leaf warblers, it was formerly placed in the "Old World warbler" assemblage, but now belongs to the new leaf-warbler family Phylloscopidae.
The green-crowned warbler is a species of leaf warbler. It was formerly included in the "Old World warbler" assemblage.
Alström's warbler, or the plain-tailed warbler, is a species of Old World warbler in the family Phylloscopidae. It was first described in 1999. It breeds only in China and winters as far as Thailand, Laos, Cambodia and Vietnam. Its natural habitat is temperate forests.
The two-barred warbler is a bird of the leaf warbler family (Phylloscopidae). The species was first described by Robert Swinhoe in 1861. It was formerly included in the "Old World warbler" assemblage. It is closely related to the greenish warbler, to which it was formerly considered conspecific.
The limestone leaf warbler is a species of warbler in the family Phylloscopidae. When this species was first seen, beginning in 1994, it was mistaken for the similar sulphur-breasted warbler. It is smaller than the sulphur-breasted warbler, and has more rounded wings. The plumage is almost identical, with comparisons showing only a slightly colder yellow below and a greyer tinge above. Although smaller, the bill is proportionally larger than that of the sulphur-breasted warbler. Accurate measurements are not available; the holotype has a wing length of 5.2 cm (2.0 in); the paratype a tail length of 3.7 cm (1.5 in) and a bill length of 1.39 cm (0.55 in). The species is known to occur in northern Vietnam and Laos, and potentially also occurs in southern China as well. The species name, calciatilis, means "dwelling on limestone", which along with its common name is a reference to its natural habitat, which is broadleaved evergreen and semi-evergreen forest growing around limestone karst mountains. The bare-faced bulbul, described in 2009, was found in the karst of the same region.