Paleobiota of the Maotianshan Shales

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This is a list of fossils found at Maotianshan Shales, whose most famous assemblage of organisms are referred to as the Chengjiang biota. [1]

Contents

The Maotianshan Shales are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation, [2] famous for their Konservat Lagerstätten , deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale of British Columbia, Canada.

Ecdysozoa

GenusSpeciesMaterialNotesImages

Acosmia

  • A. maotiania
  • 9 nearly complete specimens

A small ecdysozoan worm that was originally thought to have been a priapulid worm.

Fossil of Acosmia RCCBYU 10235 Acosmia maotiania.png
Fossil of Acosmia

Laojieella

  • L. thecata
  • One complete and one incomplete specimen

An ecdysozoan worm considered close to the priapulids, it has a spiny proboscis, an elliptical theca on the posterior part of the trunk, and a caudal appendage.

Scalidophora

Palaeoscolecidomorpha

GenusSpeciesMaterialNotesImages

Cricocosmia

  • C. jinningensis

A genus of palaeoscolecid worm. Fossils have been found with smaller worms seemingly attached to them.

Reconstruction of Cricocosmia with parasitic Inquicus Cricocosmia with Inquicus.jpg
Reconstruction of Cricocosmia with parasitic Inquicus

Mafangscolex

  • M. sinensis
  • M. yunnanensis

Maotianshania

  • M. cylindrica

An extinct palaeoscolecid worm closely related to Cricocosmia.

Fossil of Maotianshania Maotianshania detail CRF 02.jpg
Fossil of Maotianshania

Tabelliscolex

  • T. chengjiangensis
  • T. hexagonus
  • T. maanshanensis

A genus of palaeoscolecid worm

Tylotites

  • T. petiolaris

Priapulida

GenusSpeciesMaterialNotesImages

Corynetis

  • C. brevis
  • C. fortis

An archaeopriapulid worm known from the biota.

Reconstruction of Corynetis Corynetis.jpg
Reconstruction of Corynetis

Ercaivermis

  • E. sparios

A single specimen.

A stem-group priapulid.

Eximipriapulus

  • E. globocaudatus

A carnivorous priapulid-like worm that has been found living in the discarded shells of hyoliths.

Reconstruction of Eximipriapulus Eximipriapulus.jpg
Reconstruction of Eximipriapulus

Paraselkirkia

  • P. sinica

An archaeopriapulid worm that resembles another worm, Selkirkia , from the Burgess shale

Reconstruction of Paraselkirkia Paraselkirkia.jpg
Reconstruction of Paraselkirkia

Paratubiluchus

  • P. bicaudatus
Reconstruction of Paratubiluchus Paratubiluchus.jpg
Reconstruction of Paratubiluchus

Palaeopriapulites

  • P. parvus
Reconstruction of Palaeopriapulites Palaeopriapulites.jpg
Reconstruction of Palaeopriapulites

Selkirkia

  • S. elongata

A burrowing archaeopriapulid worm also known from the burgess shale, alongside the Fezouata Shale.

Fossil of the relatedSelkirkia Columbia Selkirkia columbia ROM63131.jpg
Fossil of the related Selkirkia Columbia

Sicyophorus

  • S. rarus

A genus of archaeopriapulid worm.

Reconstruction of Sicyophorus Sicyophorus.jpg
Reconstruction of Sicyophorus

Xiaoheiqingella

  • X. peculiaris

A priapulid worm thought to belong to the crown group.

Reconstruction of Xiaoheiqingella Xiaoheiqingella.jpg
Reconstruction of Xiaoheiqingella

Panarthropoda

Lobopodia

GenusSpeciesMaterialNotesImages

Antennacanthopodia

  • A. gracilis
  • Known from two specimens, making the species very rare in the Chengjiang.

An unarmoured lobopodian, apparently closely related to Onychophora. The long, paired, cirriform structures at the end of the body could have been sensory, or perhaps represent viscera not externally present in life.

Reconstruction of Antennacanthopodia Antennacanthopodia.jpg
Reconstruction of Antennacanthopodia

Cardiodictyon

  • C. catenulum

A highly elongate armoured lobopodian, with saddle-shaped sclerites and a lightly sclerotized head.

Reconstruction of Cardiodictyon 20210926 Cardiodictyon catenulum diagrammatic reconstruction.png
Reconstruction of Cardiodictyon

Diania

  • D. cactiformis

A heavily armoured lobopodian.

Reconstruction of Diania 20210429 Diania cactiformis diagrammatic reconstruction.png
Reconstruction of Diania

Lenisambulatrix

  • L. humboldti
  • A single specimen.

A simple, unarmoured lobopodian.

Reconstruction of Lenisambulatrix 20210928 Lenisambulatrix humboldti diagrammatic reconstruction.png
Reconstruction of Lenisambulatrix

Onychodictyon

  • O. ferox
  • O. gracilis
  • Several complete specimens.

An enigmatic genus of lobopodian, potentially related to either the Luolishaniidae or to Tardigrada. Not always monophyletic.

Reconstruction of Onychodictyon ferox 20210822 Onychodictyon ferox diagrammatic reconstruction.png
Reconstruction of Onychodictyon ferox

Paucipodia

  • P. inermis
  • Known from well-preserved body fossils.

A large unarmoured lobopodian, commonly found in association with mass death-beds of legless worms and Eldonia. Thought to have been a scavenger.

Reconstruction of Paucipodia 20210829 Paucipodia inermis diagrammatic reconstruction.png
Reconstruction of Paucipodia
Eoconchariidae
GenusSpeciesMaterialNotesImages

Microdictyon

  • M. sinicum

Microdictyon is known from sclerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Fossil of Microdictyon sinicum Microdictyon Chengjiang.JPG
Fossil of Microdictyon sinicum
Hallucishaniids
GenusSpeciesMaterialNotesImages

Luolishania

  • L. longicruris

Type genus of Luolishaniidae. The species Miraluolishania haikouensis is considered a juvenile synonym of Luolishania.

Reconstruction of Luolishania 20210902 Luolishania longicruris Miraluolishania haikouensis diagrammatic reconstruction.png
Reconstruction of Luolishania

Facivermis

  • F. yunnanicus

Derived Luolishaniid lobopodian with five setiferous sieving appendages, and lacking any walking limbs. Sometimes discovered living in constructed tubes, buried into the sediment.

Reconstruction of Facivermis Facivermis.png
Reconstruction of Facivermis

Hallucigenia

  • H. fortis

A hallucigeniid lobopodian with a bulbous sclerotized head and two pairs of small feeding appendages, borne on a short neck.

Reconstruction of Hallucigenia fortis 20210831 Hallucigenia fortis diagrammatic reconstruction.png
Reconstruction of Hallucigenia fortis
Siberiidae
GenusSpeciesMaterialNotesImages

Megadictyon

  • M. haikouensis
  • Posterior-most section of the trunk is unknown.

A poorly known siberiid.

Reconstruction of Megadictyon cf. haikouensis 20191118 Megadictyon cf. haikouensis.png
Reconstruction of Megadictyon cf. haikouensis

Jianshanopodia

  • J. decorata
  • Known from multiple large individuals, all from a singular exposure of the Chengjiang.

A giant siberiid lobopodian related to Megadictyon.

Reconstruction of Jianshanopodia 20191215 Jianshanopodia decora.png
Reconstruction of Jianshanopodia
Gilled Lobopodians
GenusSpeciesMaterialNotesImages

Omnidens

  • O. amplus
  • O. qiongqii
  • Numerous well preserved oral cones and portions of the frontal appendage.

A giant gilled lobopodian, closely related to Pambdelurion. Also present in the Xiaoshiba Biota.

Diagrammatic reconstruction of Omnidens 20210222 Omnidens Pambdelurion size.png
Diagrammatic reconstruction of Omnidens

Radiodonta

Radiodonta are a group of highly successful panarthropods. The Chengjiang Fauna includes a large number of Radiodont species, primarily from the clade Amplectobeluidae. While hurdiids are poorly known from the formation and surrounding region, several specimens attributable to the family have been discovered, but remain unnamed. Some radiodonts from the Chengjiang defy easy classification, or are otherwise debated in their exact affinity.

GenusSpeciesMaterialNotesImages

Houcaris

  • H. saron
  • H. consimilis
  • Isolated appendages.

Previously described as "Anomalocaris" saron.

Appendage of Houcaris saron 20191221 Radiodonta frontal appendage Houcaris saron.png
Appendage of Houcaris saron
Innovatiocaris
  • I. maotianshanensis
  • I. multispiniformis
  • I. sp.
  • Complete fossil and isolated appendages
Previously assigned to the Anomalocaris genus, and one of the most completely known radiodonts.
Reconstruction of I. maotianshanensis 20210624 Innovatiocaris maotianshanensis.png
Reconstruction of I. maotianshanensis

Laminacaris

  • L. chimera
  • Giant isolated appendages, up to nearly 30 centimetres long.

A giant radiodont combining features from different clades.

Appendage of Laminacaris 20191221 Radiodonta frontal appendage Laminacaris chimera.png
Appendage of Laminacaris
Amplectobeluidae
GenusSpeciesMaterialNotesImages

Amplectobelua

  • A. trispinata
  • Dozens of isolated appendages, and at least one complete body of a juvenile.

A large Amplectobeluid radiodont with long tail furcae. The frontal appendages formed a distinctive claw-like shape.

Reconstruction of Amplectobelua 20191201 Amplectobelua symbrachiata.png
Reconstruction of Amplectobelua

Lyrarapax

  • L. trilobus
  • L. unguispinus
  • Known from appendages and complete body fossils, including neural anatomy.

Small Amplectobeluid radiodont. Some specimens of this genus represent some of the smallest known radiodonts.

Reconstruction of Lyrarapax unguispinus (left) and Lyrarapax trilobus (right) 20210611 Lyrarapax.png
Reconstruction of Lyrarapax unguispinus (left) and Lyrarapax trilobus (right)

Ramskoeldia

  • R. platyacantha
  • R. consimilis?
  • Known from appendages, often associated with GLS plates, sclerites, and swimming flaps.

Large Amplectobeluid radiodonts, with crushing gnathobases. The species R. consimilis may belong to Houcaris instead. [3]

Appendages of Ramskoeldia 20191221 Radiodonta frontal appendage Ramskoeldia.png
Appendages of Ramskoeldia
Anomalocarididae
GenusSpeciesMaterialNotesImages

Anomalocaris

  • A. cf. canadensis
  • Isolated appendages.
Appendage of A. cf. canadensis Anomalocaris cf. canadensis JS-1880.png
Appendage of A. cf. canadensis

Lenisicaris

  • L. lupata
  • Isolated appendages.

A second species is included in the genus, Lenisicaris pennsylvanica, from the Kinzers Formation.

Appendage of Lenisicaris 20210513 Radiodonta frontal appendage Lenisicaris lupata.png
Appendage of Lenisicaris

Shucaris

  • S. ankylosskelos
  • Isolated appendages.
  • Oral cone
  • Partial body

An unusual radiodont seemingly transitional between anomalocaridids and amplectobeluids, with an oral cone similar to the former but gnathobase-like structures similar to the latter.

Appendage model of Shucaris 3D model of Shucaris ankylosskelos appendage.png
Appendage model of Shucaris
Hurdiidae (Peytoiidae)

Hurdiids (peytoiids [3] ) have historically been considered absent, or at least rare, from the Chengjiang Biota, an observation that researchers have found confusing given their regularity at other sites. In recent years, a number of fragmentary hurdiid fossils have been described, including 3 partial appendages which remain unnamed, [4] an isolated sclerite assigned to Cambroraster, [5] Zhenghecaris (the affinities of which, even its identity as a radiodont, remains questionable), and isolated lateral scelrites (also questionable). [6]

GenusSpeciesMaterialNotesImages

Cambroraster

  • C. sp.
  • Head sclerites.

A large nektobenthic hurdiid, first known from the Burgess Shale. Fossils of Cambroraster are also known from Mantou Formation of North China, [7] making this a very widespread genus. This genus was the first definitive peytoiid known from China.

Fossil of Cambroraster from the Burgess Shale. Cambroraster H-element ROMIP 65079.jpeg
Fossil of Cambroraster from the Burgess Shale.

Zhenghecaris

  • Z. shankouensis
  • Sclerites

Large-sized arthropod carapace, originally described as bivalved arthropod close to Thylacocephala, later reinterpreted as hurdiid radiodont close to Cambroraster , its classification is still discussed. Some fossils, apparently the lateral scelrites of a radiodont, connected anteriorly by a "beak" and featuring two-pronged, wing-like processes posteriorly (similar to the lateral processes of Zhenghecaris's central head sclerite), have also been attributed to Zhenghecaris. [6]

Fossil of Zhenghecaris 2019 Moysiuk & Caron f07 A-B.jpg
Fossil of Zhenghecaris

Arthropoda

GenusSpeciesMaterialNotesImages
Astutuscaris
  • A. bispinifer
  • A single specimen
A small arthropod of uncertain affinities superficially similar to fuxianhuiids.
Chuandianella
  • C. ovata
  • Several body fossils and carapaces
An enigmatc bivalved arthropod originally considered close to Waptia . It was then discovered that it lacked mandibles and therefore does not belong to Hymenocarina, the Waptiidae family, and even the larger mandibulata clade.
Reconstruction of Chuandianella Chuandianella life reconstruction.jpg
Reconstruction of Chuandianella

Combinivalvula

  • C. chengjiangensis

An enigmatic bivalved arthropod.

Cucumericrus

  • C. decorata
  • Multiple fragmentary specimens.

An enigmatic arthropod that has been tentatively classified as a possible radiodont, although a position closely related to Erratus as the most primitive deuteropod is more likely.

Reconstructed trunk appendage of Cucumericrus 20210207 Cucumericrus decoratus trunk appendage.png
Reconstructed trunk appendage of Cucumericrus

Cyathocephalus

  • C. bispinosus

Diplopyge

  • D. forcipatus
  • D. minutus

Dongshanocaris

  • D. foliiformis

A small arthropod that is currently "too poorly preserved to verify their identity as a valid taxa".

Erratus

  • E. sperare
  • Several carapaces
  • Eyes and other body parts

A bivalved arthropod currently placed as the most basal deuteropod, that shows the origins of lungs and legs in early arthropods.

Reconstruction of Erratus Erratus.png
Reconstruction of Erratus

Ercaia

  • E. miniscula

An arthropod that has been speculated to be an early crustacean.

Forfexicaris

  • F. valida

A small bivalved arthropod that somewhat resembles ostracods, but also has large, upward facing appendages with spines.

Reconstruction of Forfexicaris Forfexicaris valida.jpg
Reconstruction of Forfexicaris

Glossocaris

  • G. occulatus

Isoxys

  • I. auritus
  • I. curvirostratus
  • I. paradoxus

A basal nektonic arthropod that possessed a bivalved carapace, and large circular eyes, related to Surusicaris from the Burgess Shale. Also found in various other Cambrian aged sites.

Reconstruction of Isoxys curvirostratus Artistic reconstruction of Isoxys curvirostratus.jpg
Reconstruction of Isoxys curvirostratus

Jianshania

  • J. furcatus

A small, somewhat enigmatic arthropod known from the biota

Fossil of Jianshania Jianshania.png
Fossil of Jianshania

Kunmingocaris

  • K. bispinosus

Kylinxia

  • K. zhangi
  • Almost complete fossil
  • Isolated frontal appendages

An early arthropod that may show the "missing link" between early stem-arthropods, and more advanced true arthropods. It possessed frontal appendages similar to radiodonts.

Reconstruction of Kylinxia 20210310 Kylinxia zhangi.png
Reconstruction of Kylinxia

Mafangia

  • M. subscalaria

Mafangocaris

  • M. multinodus

Occacaris

  • O. oviformis

A predatory arthropod that had spiked frontal appendages similar to those of the megacheirans, and was previously thought to have belonged to that grouping

Reconstruction of Occacaris Occacaris oviformis.jpg
Reconstruction of Occacaris

Ovalicephalis

  • O. mirabilis

Parapaleomerus

  • P. sinensis

An arachnomorph arthropod of the group strabopida. Unlike the other members of strabopida, Parapaleomerus lacks dorsal eyes and only possesses ten trunk tergites. The telson has been described as trapezoidal in shape.

Reconstructed carapace of Parapaleomerus Parapaleomerus sinensis restoration.png
Reconstructed carapace of Parapaleomerus

Pisinnocaris

  • P. subconigera

Pseudoiulia

  • P. cambriensis

A poorly known arthropod that, in 2013, was suggested to be a member of the family Kootenichelidae, alongside Kootenichela and Worthenella.

Pterotum

  • P. triacanthus

Rhombicalvaria

  • R. acanthi

Sunella

  • S. grandis

Syrrhaptis

  • S. intes

Tanglangia

  • T. caudata

Tauricornicaris

  • T. latizonae
  • T. oxygonae

A giant arthropod once misidentified as a peytoiid radiodont.

Wutingella

  • W. bindosa

Yiliangocaris

  • Y. ellipticus

Yunnanocaris

  • Y. megista
Hymenocarina
GenusSpeciesMaterialNotesImages

Branchiocaris

  • B. yunnanensis
  • Several isolated carapaces

Also known in the Burgess Shale by B. pretiosa, B. yunnanensis is only known from isolated carapaces, although it is thought that this may be because in this species the carapace covered the entire body. It also has two morphotypes.

Reconstruction of the related Branchiocaris pretiosa Branchiocaris.png
Reconstruction of the related Branchiocaris pretiosa

Canadaspis

  • C. laevigata

A hymenocarine arthropod, also known from the Burgess Shale by C. perfecta, although some authors believe that both species should be separated into different genera. They were mainly benthic animals.

Reconstruction of Canadaspis laevigata Canadaspis laevigata.png
Reconstruction of Canadaspis laevigata

Clypecaris

  • C. pteroidea
  • C. serrata

A hymenocarine arthropod. The species in the genus are primarily distinguished by the presence of a serrated edge on the front of the carapace of C. serrata. C. serrata is noted for the modification of an anterior pair of limbs into spined grasping appendages, indicating a predatory lifestyle.

Reconstruction of Clypecaris serrata Clypecaris serrata.png
Reconstruction of Clypecaris serrata

Ercaicunia

  • E. multinodosa

A hymenocarine arthropod that had a bivalved carapace which covered about a third of its total body-length, and had up to six serrations on its forward edge. The head had a pair of large uniramous antennae, as well as a smaller pair of secondary antennae, and a pair of mandibles and maxillae.

Reconstruction of Ercaicunia Ercaicunia multinodosa pennetta.png
Reconstruction of Ercaicunia

Erjiecaris

  • E. minusculo

A small euarthropod, and unlike other hymenocarines, it possibly had eyes directly on top of its carapace. It was most likely a nektobenthic animal.

Reconstruction of Erjiecaris Erjiecaris.png
Reconstruction of Erjiecaris

Pectocaris

  • P. spatiosa
  • P. eurypetala
  • P. inopinata

A bivalved arthropod of the hymenocarina grouping. The species P. spatiosa is regarded as a filter feeder, using the setae on the endites of their limbs to filter out matter from the water column

Reconstruction of Pectocaris spatiosa Pectocaris spatiosa.png
Reconstruction of Pectocaris spatiosa

Synophalos

  • S. xynos

A hymenocarine notable for fossils showing a bizarre chain of multiple individuals. Although the purpose for this behavior is unknown, it may have served a reproductive, migratory, or defensive purpose, with the authors of the describing paper of this genus considering migration as its most likely function.

Reconstruction of a Synophalos chain Synophalos (grupo).png
Reconstruction of a Synophalos chain
Tuzoia
  • T. sinensis
  • several carapaces and soft body parts
A large hymenocarine that possessed a semicircular bivalved carapace, and was one of the largest arthropods from the Cambrian
Reconstruction of Tuzoia sp. Tuzoia life restoration.jpg
Reconstruction of Tuzoia sp.
Jugatacaris A. agillis
  • several carapaces and soft body parts
A hymenocarine arthropod. The carapace of the animal was around 28 to 37 millimetres (1.1 to 1.5 in) in length, with a pronounced ridge at the top of the carapace separating the two valves, which formed a fin-like structure raised above the carapace. The head had a pair of stalked eyes, as well as a dumbbell shaped medial eye between them. The head also bore a pair of mandibles as well as at least one and possibly two pairs of antennules.
Reconstruction of Jugatacaris Jugatacaris.png
Reconstruction of Jugatacaris
Xiazhuangocaris X. chenggongensis
  • a single specimen (NIGP 172765), which only preserves the carapace and trunk regions
A hymenocarine with no known head fossils. The material known show that the carapace had a pronounced pair of notches at its front, as well as a posterior notch at its rear. The body had at least 13 tergite-pleurite rings, which terminate in a pair of rounded caudal rami, which are fringed with setae.
Reconstruction of Xiazhuangocaris Xiazhuangocaris.png
Reconstruction of Xiazhuangocaris
Fuxianhuiida
GenusSpeciesMaterialNotesImages

Chengjiangocaris

  • C. longiformis
  • C. kunmingensis

A Fuxianhuiid arthropod with numerous appendages. It is one of the most complete arthropods from the lower Cambrian.

Reconstruction of Chengjiangocaris 20211117 Chengjiangocaris kunmingensis.png
Reconstruction of Chengjiangocaris

Fuxianhuia

  • F. protensa

A large arthropod named after Fuxian Lake. Some adult specimens are found closely associated with numerous juveniles, indicating a level of parental care.

Reconstruction of Fuxianhuia with associated juveniles 20211117 Fuxianhuia protensa.png
Reconstruction of Fuxianhuia with associated juveniles
Bradoriida
GenusSpeciesMaterialNotesImages

Comptaluta

  • C. inflata
  • C. kailiensis

Indiana

  • I. sp.
CT-scan of a fossil of Indiana sp. Indiana Bradoriida.jpg
CT-scan of a fossil of Indiana sp.

Jiucunella

  • J. paulula

Kunmingella

  • K. angustacostata
  • K. douvillei
  • K. guanshanensis
Reconstruction of Kunmingella douvillei Kunmingella.png
Reconstruction of Kunmingella douvillei

Kunyangella

  • K. cheni

Liangshanella

  • L. liangshanensis

Malongella

  • M. bituerculata

Parakunmingella

  • P. malongensis

Spinokunmingella

  • S. typica

Tsunyiella

  • T. daindongensis
Megacheira
GenusSpeciesMaterialNotesImages

Haikoucaris

  • H. ercaiensis
Reconstruction of Haikoucaris 20191020 Haikoucaris ercaiensis.png
Reconstruction of Haikoucaris
Leanchoiliidae
GenusSpeciesMaterialNotesImages

Alalcomenaeus

  • A. cambricus

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Reconstruction of Alalcomenaeus 20210422 Alalcomenaeus cambricus.png
Reconstruction of Alalcomenaeus

Leanchoilia

  • D. asiatica
  • L. illecebrosa

One of the several species of Leanchoilia, it has known juvenile specimens and the labrum morphology known.

Fossil of Leanchoilia illecebrosa Leanchoilia illecebrosa.jpg
Fossil of Leanchoilia illecebrosa
Jianfengiidae
GenusSpeciesMaterialNotesImages

Fortiforceps

  • F. foliosa
Reconstruction of Fortiforceps 20191025 Fortiforceps foliosa.png
Reconstruction of Fortiforceps
Sklerolibyon *S. maomima
Reconstruction of Sklerolibyon Sklerolibyon reconstruction.png
Reconstruction of Sklerolibyon

Jianfengia

  • J. multisegmentalis
Reconstruction of Jianfengia Aria et al. 2020 Jianfengia multisegmentalis reconstruction.png
Reconstruction of Jianfengia

Parapeytoia

  • P. yunnanensis

A jianfengiid once misidentified as a radiodont

Reconstruction of known elements from Parapeytoia 20191021 Parapeytoia yunnanensis ventral structures.png
Reconstruction of known elements from Parapeytoia
Marrellomorpha
GenusSpeciesMaterialNotesImages

Primicaris

  • P. larvaformis
Reconstruction of Primicaris Primicaris 3D.png
Reconstruction of Primicaris
Artiopoda
GenusSpeciesMaterialNotesImages

Acanthomeridion

  • A. serratum

Multiple Specimens.

Reconstruction of the carapace of Acanthomeridion Acanthomeridion.jpg
Reconstruction of the carapace of Acanthomeridion

Almenia

  • A. spinosa

Cindarella

  • C. eucalla
Reconstruction of Cindarella Cindarella.png
Reconstruction of Cindarella

Kuamaia

  • K. lata
  • K. muricata
Reconstruction of Kuamaia lata Kuamaia lata.svg
Reconstruction of Kuamaia lata

Panlongia

  • P. spinosa
  • P. tetranudosa
Fossil of Panlognia Panlongia spinosa CRF.jpeg
Fossil of Panlognia

Pygmaclypeatus

  • P. daiensis
CT-scan of a specimen of Pygmaclypeatus Pygmaclypeatus.tif
CT-scan of a specimen of Pygmaclypeatus

Retifacies

  • R. abnormalis
Reconstruction of Retifacies Retifacies.jpg
Reconstruction of Retifacies

Saperion

  • S. glumaceum

Sidneyia

  • S. minor
Reconstruction of Sidneyia minor Sidneyia minor.png
Reconstruction of Sidneyia minor

Sinoburius

  • S. lunaris
Reconstruction of Sinoburius lunaris Sinoburius 7.png
Reconstruction of Sinoburius lunaris

Skioldia

  • S. aldna

Squamacula

  • S. clypeata
Reconstruction of Squamacula S. clypeata.png
Reconstruction of Squamacula

Urokodia

  • U. aequalis

An arthropod that had segmentation which resembled that of a millipede, as well as head and tail shields with thorny spikes. Recently, it was recovered as the most basal artiopod.

Reconstruction of Urokodia Urokodia restoration.png
Reconstruction of Urokodia

Xandarella

  • X. spectaculum
Reconstruction of Xandarella Xandarella.png
Reconstruction of Xandarella
Nektaspida
GenusSpeciesMaterialNotesImages

Misszhouia

  • M. longicaudata
Reconstruction of Misszhouia Misszhouia.png
Reconstruction of Misszhouia

Naraoia

  • N. spinosa
Reconstruction of Naraoia spinosa Naraoia spinosa.jpg
Reconstruction of Naraoia spinosa

Kangacaris

  • K. shui
Reconstruction of Kangacaris zhangi, the type species from Cambrian Australia Kangacaris zhangi.jpg
Reconstruction of Kangacaris zhangi, the type species from Cambrian Australia
Trilobita
GenusSpeciesMaterialNotesImages

Eoredlichia

  • E. intermedia
Reconstruction of Eoredlichia Eoredlichia intermedia life restoration.jpg
Reconstruction of Eoredlichia

Kuanyangia

  • K. pustulosa

Palaeolenus

  • E. miniscula
Fossil pair of Palaeolenus Palaeolenus douvillei pair.jpg
Fossil pair of Palaeolenus

Tsunyidiscus

  • T. aclis
Line drawing of Tsunyidiscus niutitangensis Tsunyidiscus niutitangensis linedrawing.jpg
Line drawing of Tsunyidiscus niutitangensis

Wutingapsis

  • W. tingi

Yunnanocephalus

  • Y. yunnanensis
Fossil of Yunnanocephalus Yunnanocephalus S CRF.jpg
Fossil of Yunnanocephalus

Spiralia

Clade Gnathifera

GenusSpeciesMaterialNotesImages

Amiskwia

  • A. sinica

Inquicus

  • I. fellatus
Reconstruction of Inquicus Inquicus fellatus.jpg
Reconstruction of Inquicus

Chaetognatha

GenusSpeciesMaterialNotesImages

Ankalodous

  • A. ercainella

Eognathacantha

  • E. ercainella
Reconstruction of Eognathacantha Eognathacantha ercainella.jpg
Reconstruction of Eognathacantha

Protosagitta

  • P. spinosa
Reconstruction of Protosagitta Protosagitta spinosa.jpg
Reconstruction of Protosagitta

Lophotrochozoa

Annelida

GenusSpeciesMaterialNotesImages

Iotuba

  • I. chengjiangensis
Fossil of Iotuba Iotuba chengjiangensis.tif
Fossil of Iotuba

Mollusca

GenusSpeciesMaterialNotesImages

Orthrozanclus

  • O. elongata
Reconstruction of Orthrozanclus elongata Orthrozanclus elongata reconstruction.jpg
Reconstruction of Orthrozanclus elongata

Wiwaxia

  • W. papillo
Reconstruction of Wiwaxia papilio Lateral aspect of Wiwaxia papilio.png
Reconstruction of Wiwaxia papilio

Hyolitha

GenusSpeciesMaterialNotesImages

Ambrolinevitus

  • A. maximus
  • A. platypluteus
  • A. ventricosus

Burithes

  • B. yunnanensis

Glossolithes

  • G. magnus

Linevitus

  • L. billingsi
  • L. flabellaris
  • L. opimus

Phoronida

GenusSpeciesMaterialNotesImages

Eophonoris

  • E. chengjiangensis

Entoprocta

GenusSpeciesMaterialNotesImages

Cotyledion

  • C. tylodes
Reconstruction of Cotyledion Reconstruction of Cotyledion tylodes.jpg
Reconstruction of Cotyledion

Cornulitida

GenusSpeciesMaterialNotesImages

Cambrocornulitus

  • C. rarus

Brachiopoda

GenusSpeciesMaterialNotesImages

Diandongia

  • D. pista

Heliomedusa

  • H. orienta
Fossil of Heliomedusa orienta Heliomedusa orienta.jpg
Fossil of Heliomedusa orienta

Lingulella

  • L. orienta
Fossil of the related Lingulella caelata Botsfordia caelata 1.png
Fossil of the related Lingulella caelata

Lingulellotreta

  • L. malongensis

Longtancunella

  • L. chengjiangensis

Deuterostomia

GenusSpeciesMaterialNotesImages

Yunnanozoon

  • Y. lividium
A species of deuterostome animal with controversial taxonomy, with some authors placing it as an chordate, stem-vertebrate, hemichordate, etc.
Fossil of Yunnanozoon Yunnanozoon lividum RCCBYU 10310.jpg
Fossil of Yunnanozoon

Chordata

GenusSpeciesMaterialNotesImages

Cathaymyrus

  • C. diadexus
  • C. haikouensis
A chordate known from the Yunnan Province with potential relations to cephalochordates like lancelets.
Reconstruction of C. diadexus and Vetulocystis Cathaymyrus diacodexis.jpg
Reconstruction of C. diadexus and Vetulocystis

Haikouichthys

  • H. ercaicunensis
An early craniate (an animal with a distinct notochord and head) of the Myllokunmingiida order. Some authors consider it potentially synonymous with the contemporary Myllokunmingia. [8]
Reconstruction of Haikouichthys Haikouichthys NT2.jpg
Reconstruction of Haikouichthys

Myllokunmingia

  • M. fengjiaoa
A myllokunmingiid known for its wide body plan, sail-like fin on its dorsal plane, and a non-biomineralized skeleton.
Reconstruction of Myllokunmingia Myllokunmingia NT.jpg
Reconstruction of Myllokunmingia

Zhongjianichthys

  • Z. rostratus
Another myllokunmingiid known for lacking a dorsal fin, and having a more slim body compared to its two contemporary relatives.
Reconstruction of Zhongjianichthys Zhongjianichthys rostratus.jpg
Reconstruction of Zhongjianichthys

Zhongxiniscus

  • Z. intermedius
An early chordate thought to represent a transitional form between earlier chordates, and more derived craniates.
Reconstruction of Zhongxiniscus Zhongxiniscus.png
Reconstruction of Zhongxiniscus

Vetulicolia

GenusSpeciesMaterialNotesImages

Beidazoon

  • B. venustum
Reconstruction of Beidazoon Beidazoon venustum copy.jpg
Reconstruction of Beidazoon

Didazoon

  • D. haoae
Reconstruction of Beidazoon Didazoon haoae.jpg
Reconstruction of Beidazoon

Pomatrum

  • P. ventralis
Reconstruction of Beidazoon Xidazoon stephanus e.jpg
Reconstruction of Beidazoon

Vetulicola

  • V. cuneata
  • V. gantoucunensis
  • V. rectangulata
Reconstruction of Vetulicola rectangulata Vetulicola rectangulata restoration.jpg
Reconstruction of Vetulicola rectangulata

Yuyuanozoon

  • Y. magnificissimi
Reconstruction of Yuyuanozoon Yuyuanozoon.jpg
Reconstruction of Yuyuanozoon

Ambulacraria

Cambroernida

GenusSpeciesMaterialNotesImages

Phlogites

  • P. brevis
  • P. longus

Senior synonym of Cheungkongella ancestralis.

Fossil of Phlogites Phlogites longus.png
Fossil of Phlogites

Rotadiscus

  • R. grandis
Reconstruction of Rotadiscus Rotadiscus grandis.jpg
Reconstruction of Rotadiscus

Stellostomites

  • S. eumorphus
Senior synonym of Eldonia eumorpha.
Diagram of Stellostomites Stellostomites diagrams.png
Diagram of Stellostomites

Echinodermata

GenusSpeciesMaterialNotesImages

Dianchicystis

  • D. jianshanensis

Vetulocystis

  • V. catenata
Reconstruction of Vetulocystis (bottom) Cathaymyrus diacodexis.jpg
Reconstruction of Vetulocystis (bottom)

Phylum Hemichordata

GenusSpeciesMaterialNotesImages

Cambrobranchus

  • C. pelagobenthos

Galeaplumosus

  • G. abilus

Shankouclava

  • S. anningense
  • S. shankouense

Likely the earliest known tunicate, alongside Palaeoikopleuria .

Reconstruction of Shankouclava anningense Shankouclava anningense.jpg
Reconstruction of Shankouclava anningense

Yuknessia

  • Y. simplex

Phylum Ctenophora

GenusSpeciesMaterialNotesImages

Maotianoascus

  • M. octanarius
Reconstruction of Maotianoascus octanarius Maotianoascus octanarius.JPG
Reconstruction of Maotianoascus octanarius

Sinoascus

  • S. paillatus
Reconstruction of Sinoascus papillatus Sinoascus papillatus.jpg
Reconstruction of Sinoascus papillatus
Stromatoveris
  • S. psygmoglena
Reconstruction of Stromatoveris psygmoglena Stromatoveris psygmoglena.jpg
Reconstruction of Stromatoveris psygmoglena

Trigoides

  • T. aclis

Phylum Porifera

GenusSpeciesMaterialNotesImages

Allantospongia

  • A. mica

Choia

  • C. xiaolantianensis

Choiaella

  • C. radiata
Hazelia
  • H.

Leptomitella

  • L. confusa
  • L. conica
  • L. metta

Leptomitus

  • L. teretiusculus
Paraleptomitella
  • Paraleptomitella dictyodroma
  • Paraleptomitella globula

Quadrolaminiella

  • Quadrolaminiella crassa
  • Quadrolaminiella diagonalis

Saetaspongia

  • S. densa
Sinfoflabrum
  • S. antiquum
Fossil of Sinfoflabrum Sinoflabrum antiquum.jpg
Fossil of Sinfoflabrum
Triticispongia
  • T. diagonata

Phylum Cnidaria

GenusSpeciesMaterialNotesImages

Archisaccophyllia

  • A. kunmingensis
Fossil of Archisaccophyllia Archisaccophyllia kunmingensis YKLP-10379.png
Fossil of Archisaccophyllia

Nailiana

  • N. elegans

Priscapennamarina

  • P. angusta

Xianguangia

  • X. sinica
Fossil of Xianguangia Xianguangia sinica YKLP-13830.png
Fossil of Xianguangia

Enigmatic

GenusSpeciesMaterialNotesImages

Allonnia

  • A. phrixothrix
Reconstruction of Allonnia sp. Allonnia pennetta.png
Reconstruction of Allonnia sp.

Anthrotum

  • A. robustus

Batofasciculus

  • B. ramificans

Calathites

  • C. spinalis

Cambrotentacus

  • C. sanwuia
Reconstruction of a pair of Cambrotentacus Cambrotentacus sanwuia.jpg
Reconstruction of a pair of Cambrotentacus

Conicula

  • C. straita

Daihua

  • D. sanqiong
Fossil of Daihua Daihua sanqiong YKLP-13401a.png
Fossil of Daihua

Dinomischus

  • D. venustus
Fossil of Dinomischus venustus Dinomischus venustus YKLP-13412b.png
Fossil of Dinomischus venustus

Discoides

  • D. abnormalis

Hippotrum

  • H. spinatus

Jiucunia

  • J. petalina

Maanshania

  • M. crusticeps

Macrocephalus

  • M. elongates

Petalilium

  • P. latus

Possible synonym of Nectocaris pteryx

Fossil of Petalilium Petalilium HK-MF-00001A 098.g2.jpg
Fossil of Petalilium

Parvulonoda

  • P. dubia

Phacatrum

  • P. tubifer

Phasangula

  • P. striata

Pristitoites

  • P. bifarius

Rhipitrus

  • R. calvifer

Miscellaneous macroalgae

Macroalgae rank fourth in species-level diversity behind arthropods, priapulids, and sponges, and account for 71.5% of total abundance from the biota, particularly dominated by unattached (floating, planktonic, and drifting) species. [9]

GenusSpeciesMaterialNotesImages

Enteromophites

  • E. intestinalis
Sessile, attached to the substrate.

Fuxianospira

  • F. gyrata
The most common species of macroalgae present, accounting for 52% of all macroalgae fossils reported from the Chengjiang.Unattached (planktonic, drifting, floating)

Liulingjitaenia

  • L. alloplecta
Unattached (planktonic, drifting, floating)

Longfengshania

  • L. stipitata
  • L. spheria
  • L. cordata
Sessile, attached to the substrate.

Morania

  • M. fragmenta
Unattached (planktonic, drifting, floating)
Fossil of Morania fragment from the Cambrian Wheeler Shale of Utah, USA. Morania fragmenta (fossil bacteria) (Wheeler Formation, Middle Cambrian; Millard County, Utah, USA) 2.jpg
Fossil of Morania fragment from the Cambrian Wheeler Shale of Utah, USA.

Paralongfengshania

  • P. sicyoides
Sessile, attached to the substrate.

Paradelesseria

  • P. sanguinea
Unattached (planktonic, drifting, floating)

Plantulaformis

  • P. sinensis
Sessile, attached to the substrate.

Punctariopsis

  • P. latifolia
  • P. simplexis
Sessile, attached to the substrate.

Sinocylindra

  • S. yunnanensis
  • S. linearis
Unattached (planktonic, drifting, floating)
Fossil of Sinocylindra yunnanensis Zhejiang Geological Museum 20240804 121334.jpg
Fossil of Sinocylindra yunnanensis

Tawuia

  • T. sp.
Unattached (planktonic, drifting, floating)
Fossil of Tawuia sp. Tawuia NGMC.jpg
Fossil of Tawuia sp.

Yunnanospirellus

  • Y. typica
  • Y. elegans
Unattached (planktonic, drifting, floating)

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Shucaris is a genus of radiodont of uncertain taxonomic placement from the Lower Cambrian Maotianshan Shales in Yunnan, South China. The type and only species is S. ankylosskelos, known from multiple specimens comprising frontal appendages, multiple endites, gnathobase‐like structures, a nearly complete body, a head carapace complex, and one body flap associated with setal blades.

References

  1. "Chengjiang Maotianshan Shales". www.fossilmuseum.net. Archived from the original on 9 March 2021. Retrieved 10 September 2015.
  2. Lipps, J. H.; Signor, P. W (1992). Origin and early evolution of the Metazoa. Springer. ISBN   978-0-306-44067-0.
  3. 1 2 McCall, Christian R. A. (September 2023). "A large pelagic lobopodian from the Cambrian Pioche Shale of Nevada". Journal of Paleontology. 97 (5): 1009–1024. Bibcode:2023JPal...97.1009M. doi:10.1017/jpa.2023.63. ISSN   0022-3360.
  4. Wu, Yu; Pates, Stephen; Ma, Jiaxin; Lin, Weiliang; Wu, Yuheng; Zhang, Xingliang; Fu, Dongjing (1 November 2022). "Addressing the Chengjiang conundrum: A palaeoecological view on the rarity of hurdiid radiodonts in this most diverse early Cambrian Lagerstätte". Geoscience Frontiers. 13 (6): 101430. Bibcode:2022GeoFr..1301430W. doi: 10.1016/j.gsf.2022.101430 . ISSN   1674-9871.
  5. Liu, Yu; Lerosey-Aubril, Rudy; Audo, Denis; Zhai, Dayou; Mai, Huijuan; Ortega-Hernández, Javier (July 2020). "Occurrence of the eudemersal radiodont Cambroraster in the early Cambrian Chengjiang Lagerstätte and the diversity of hurdiid ecomorphotypes". Geological Magazine. 157 (7): 1200–1206. Bibcode:2020GeoM..157.1200L. doi:10.1017/S0016756820000187. ISSN   0016-7568.
  6. 1 2 Zeng, Han; Zhao, Fangchen; Yin, Zongjun; Zhu, Maoyan (2 January 2018). "Morphology of diverse radiodontan head sclerites from the early Cambrian Chengjiang Lagerstätte, south-west China". Journal of Systematic Palaeontology. 16 (1): 1–37. Bibcode:2018JSPal..16....1Z. doi:10.1080/14772019.2016.1263685. ISSN   1477-2019.
  7. Sun, Zhixin; Zeng, Han; Zhao, Fangchen (September 2020). "Occurrence of the hurdiid radiodont Cambroraster in the middle Cambrian (Wuliuan) Mantou Formation of North China". Journal of Paleontology. 94 (5): 881–886. Bibcode:2020JPal...94..881S. doi:10.1017/jpa.2020.21. ISSN   0022-3360.
  8. Lerosey-Aubril, R.; Ortega-Hernández, J. (2024). "A long-headed Cambrian soft-bodied vertebrate from the American Great Basin region". Royal Society Open Science. 11 (7). 240350. Bibcode:2024RSOS...1140350L. doi: 10.1098/rsos.240350 . PMC   11267725 . PMID   39050723.
  9. Li, Gaorong; Wei, Fan; Guo, Jin; Cong, Peiyun (2024). "Macroalgae from the early Cambrian Chengjiang biota". Papers in Palaeontology. 10 (5): e1585. Bibcode:2024PPal...10E1585L. doi:10.1002/spp2.1585. ISSN   2056-2802.