1995 in paleontology

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils . ^[1] This includes the study of body fossils, tracks ( ichnites ), burrows , cast-off parts, fossilised feces ( coprolites ), palynomorphs and chemical residues . Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science . This article records significant discoveries and events related to paleontology that occurred or were published in the year 1995.

Plants

Conifers

Conifer research

• Phipps, Osborne, & Stockey detail permineralized Pinus pollen cones from the Allenby Formations Princeton Chert site. The description is the first to include in-situ pollen ultrastructure and the cones are the oldest Pinus pollen cones that had been described to date. Affiliation with the Princeton chert organ taxa Pinus similkameenensis (leaves) and Pinus arnoldii (seed cones) was suggested. ^[2]

Arthropods

Newly named arachnids

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Uintascorpio [3]	gen et sp nov	valid	Perry	Middle Eocene	Green River Formation	USA	only scorpion from the Green River Formation

Newly named insects

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aphaenogaster amphioceanica [4]	Sp nov	Valid	De Andrade	Burdigalian	Dominican amber	Dominican Republic	A myrmicine ant	Aphaenogaster amphioceanica
Aphaenogaster praerelicta [4]	Sp nov	Valid	De Andrade	Burdigalian	Mexican amber	Mexico	A myrmicine ant
Baltimartyria [5]	gen et comb nov	valid	Skalski	Early Eocene	Baltic amber		A micropterigid moth moved from Micropterix proavitella (1936)
Dryinus grimaldii [6]	sp nov	valid	Olmi	Burdigalian	Dominican amber	Dominican Republic	A Dryinuslamellatus group wasp	Dryinus grimaldii
Electrinocellia [7]	subfam, gen et sp nov	Valid	Engel	Eocene	Baltic amber		sister taxon to the rest of Inocelliidae
Exocryptocerus jansei [8]	sp nov	Jr synonym	Vierbergen & Scheven	Burdigalian	Dominican amber	Dominican Republic	A myrmicinae ant, moved to Cephalotes jansei (1999) [9]	Cephalotes jansei
Exocryptocerus elevatus [8]	sp nov	Jr synonym	Vierbergen & Scheven	Burdigalian	Dominican amber	Dominican Republic	A myrmicinae ant, jr synonym of Cephalotes serratus (1999) [9]
Exocryptocerus serratus [8]	sp nov	Jr synonym	Vierbergen & Scheven	Burdigalian	Dominican amber	Dominican Republic	A myrmicinae ant, moved to Cephalotes serratus (1999) [9]	Cephalotes serratus
Exocryptocerus truncatus [8] [9]	sp nov	Jr synonym	Vierbergen & Scheven	Burdigalian	Dominican amber	Dominican Republic	A myrmicinae ant, jr synonym of Cephalotes serratus (1999) [9]	Cephalotes serratus
Zacryptocerus alveolatus [8]	sp nov	Jr synonym	Vierbergen & Scheven	Burdigalian	Dominican amber	Dominican Republic	A myrmicinae ant, moved to Cephalotes alveolatus (1999) [9]	Zacryptocerus alveolatus

Brachiopods

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Boucotia argentina [10]	Sp nov	Valid	Herrera	Lochkovian	Talacasto Formation	Argentina	A species of Boucotia

Molluscs

Bivalves

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Camya [11]	gen et sp nov	valid	Hinz-Schallreuter	Early Cambrian	Bornholm	Denmark	The type species is C. asy
Pojetaia ostseensis [11]	sp nov	nomen dubium	Hinz-Schallreuter	Early Cambrian	Bornholm	Denmark	Probable synonym of P. runnegari

Fish

Newly named bony fish

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Asiamericana [12]	Gen. et sp. nov	dubious	Nesov	Turonian	Bissekty Formation	Uzbekistan	A possible ichthyodectid. Originally described as possible spinosaurid teeth.

Archosauromorphs

Newly named dinosaurs

• Fossil hunters working on behalf of the Royal Saskatchewan Museum discover a large coprolite from a theropod dinosaur in Maastrichtian strata. In 1997 it is sent to coprolite specialist Karen Chin, who determines that this specimen of fossilized feces was attributable to Tyrannosaurus rex . One year later, in 1998, Karen Chin and others publish a joint paper in Nature announcing the finding.
• Paul Sereno lead an expedition to the Kem Kem region of southeastern Morocco. Among the fossils discovered is a partial skull of Carcharodontosaurus saharicus . Significantly, it preserves a "complete and undistorted braincase" which would later be described in detail along with the structure of the inner ear of C. saharicus by Hans C. E. Larsson in 2001. ^[13]

Data courtesy of George Olshevsky's dinosaur genera list. ^[14]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Achelousaurus [15]	Gen et sp nov	Valid	Sampson	Campanian	Two Medicine Formation	USA ( Montana)	A centrosaurin ceratopsian	Achelousaurus
Ampelosaurus [16]	Gen et sp nov	Valid	Le Loeuff	Maastrichtian	Marnes Rouges Inferieures Formation	France	A French titanosaur	Ampelosaurus
"Brontoraptor"		nomen nudum	Redman
Bugenasaura	Gen et sp nov	Junior synonym of Thescelosaurus [17]	Galton	Lancian	Hell Creek	USA
Chindesaurus [18]	Gen et sp nov	Valid	Long & Murry			USA ( Arizona, New Mexico and Texas)	A herrerasaurid	Chindesaurus
"Dinotyrannus" [19]	Gen et sp nov	Junior synonym of Tyrannosaurus	Olshevsky	Lancian
Einiosaurus [15]	Gen et sp nov	Valid	Sampson	Campanian	Two Medicine Formation	USA ( Montana)	A centrosaurin ceratopsian	Einiosaurus
Genusaurus [20]	Gen et sp nov	Valid	Accarie et al.			France
Giganotosaurus [21]	Gen et sp nov	Valid	Coria & Salgado			Argentina	A carcharodontosaurid	Giganotosaurus
Jainosaurus [22]		Valid	Hunt, Lockley, Lucas, & Meyer			India
"Jenghizkhan" [23]		Junior Synonym	Olshevsky				Junior synonym of Tarbosaurus	Tarbosaurus .
Jingshanosaurus [24]		Valid	Zhang & Yang			China		"Jingshanosaurus"
Kulceratops [12]		Valid	Nesov
"Liassaurus" [25]		nomen nudum	Welles, Powell, & Pickering
"Merosaurus" [25]		nomen nudum	Welles, Powell, & Pickering
Niobrarasaurus [26]		Valid	Carpenter, Dilkes, & Weishampel	Coniacian-Santonian	Niobrara Formation	United States ( Kansas)	A nodosaurid
Saurophaganax [27]		Valid	Chure	Kimmeridgian-Tithonian	Morrison Formation	United States ( Oklahoma)	An allosaurid
"Stygivenator" [19]	Gen nov	junior synonym	Olshevsky	Lancian	Hell Creek Formation		Junior synonym of Tyrannosaurus
Texasetes [28]	Gen et sp nov	Valid	Coombs	late Albian-early Cenomanian	Paw Paw Formation	United States ( Texas)	A nodosaurid
"Walkersaurus" [25]		nomen nudum	Welles, Powell & Pickering vide: Pickering	Bajocian			Material later named Duriavenator

Newly named birds

Name	Status	Novelty	Authors	Age	Type locality	Country	Notes	Images
Ajaia chione [29]	Sp. nov.	Valid	Steven D. Emslie	Pleistocene	Early Irvingtonian, Bermont Formation	USA: Florida	A threskiornithid.
Ameripodius silvasantosi [30]	Gen. et sp. nov.	Valid	Herculano M. F. de Alvarenga	Late Oligocene-Early Miocene	Tremembé Formation	Brazil	A quercymegapodiid craciform
Anhinga malagurala [31]	Sp. nov.	Valid	Brian Mackness	Early Pliocene	Allingham Formation	Australia: Queensland	An anhingid
Avisaurus gloriae [32]	Sp. nov.	Valid	David J. Varricchio Luis M. Chiappe	Late Cretaceous	Maastrichtian, Hell Creek Formation	USA: Montana	An avisaurid enantiornithean
Boluochia zhengi [33]	Gen. et sp. nov.	Valid	Zhou Zhonghe	Early Cretaceous	Valanginian, Jiufotang Formation	China	A longipterygid enantiornithean
Confuciusornis sanctus [34]	Gen. et sp. nov.	Valid	Hou Lianhai Zhou Zhonghe Gu Yucai Zhang He	Early Cretaceous	Yixian Formation	China	A confuciusornithid
Corvus moravicus [35]	Sp. nov.	Valid	Jiri Mlikovsky	Early Pleistocene	Stránská Skála	Czech Republic	A corvid
Eudocimus leiseyi [29]	Sp. nov.	Valid	Steven D. Emslie	Pleistocene	Early Irvingtonian, Bermont Formation	USA: Florida	A threskiornithid
Grus pagei [36]	Sp. nov.	Valid	Kenneth E. Campbell, jr.	Late Pleistocene	Rancho la Brea	USA: California	A gruid
Hinasuri nehuensis [37]	Gen. et sp. nov.	Valid	Claudia P. Tambussi	Middle-Late Pliocene	Monte Hermoso Formation	Argentina	A rheid
Idiornis tuberculata [38]	Sp. nov.	Valid	Dieter S. Peters	Middle Eocene	Messel pit: MP 11	Germany: Hessen	An idiornithid gruiform; the genus was synonymized with the genus Dynamopterus Milne-Edwards, 1892 by Mourer-Chauviré, 2013 [39]
Itardiornis hessae [40]	Gen. et sp. nov.	Valid	Cécile Mourer-Chauviré	Late Eocene and Middle Eocene	MP 21 and MP 23	France	A messelornithid gruiform
Lagopus balcanicus [41]	Sp. nov.	Valid	ppZlatozar N. Boev	Late Pliocene	MN 17, Middle Villafranchian	Bulgaria	A phasianid
Larus lacus [42]	Sp. nov.	Valid	Steven D. Emslie	Late Pliocene	Late Blancan	USA: Florida	A larid
Larus perpetuus [42]	Sp. nov.	Valid	Steven D. Emslie	Late Pliocene	Late Blancan	USA: Florida	A larid
Meganhinga chilensis [43]	Gen. et sp. nov.	Valid	Herculano M. F. de Alvarenga	Miocene	Cura-Mallín Formation	Chile	An anhingid
Menura tyawanoides [44]	Sp. nov.	Valid	Walter E. Boles	?Early Miocene	Riversleigh	Australia: Queensland	A menurid
Messelornis russelli [40]	Sp. nov.	Valid	Cécile Mourer-Chauviré	Late Paleocene	MP 6	France	A messelornithid gruiform
Phalacrocorax filyawi [42]	Sp. nov.	Valid	Steven D. Emslie	Late Pliocene	Late Blancan	USA: Florida	A phalacrocoracid
Qinornis paleocenica [45]	Gen. et sp. nov.	Valid	Xue Xiangxu	Early-Middle Paleocene	Shimen Basin	China	An indeterminate neognath

Pterosaurs

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Plataleorhynchus	Gen. et sp. nov	Valid	Howse and Milner	Early Cretaceous	Purbeck Limestone	UK		Plataleorhynchus

Synapsids

Mammals

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Asiavorator [46]	Gen. et sp. nov	Valid	Spassov and Lange-Badré	Late Eocene to early Oligocene	Hsanda Gol Formation	Mongolia	Type species is A. altidens, though this later became a junior synonym of A. gracilis.	Asiavorator limb bones

References

• Paleontology portal

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3. Santiago-Blay, Jorge A.; Soleglad, Michael E.; Fet, Victor (2004). "A redescription and family placement of Uintascorpio Perry, 1995 from the Parachute Creek Member of the Green River Formation (middle Eocene) of Colorado, USA (Scorpiones: Buthidae)" (PDF). Revista Ibérica de Aracnología . 10: 7–16.
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10. Herrera, Zarela Angélica (1995). "The first notanopliid brachiopod from the South American Devonian sequence" . Geobios. 28 (3): 337–342. doi:10.1016/S0016-6995(95)80008-5.
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12. Nesov, L.A. (1995). "Dinosaurs of northern Eurasia: new data about assemblages, ecology and palaeogeography (in Russian)". University of Saint Petersburg, Saint Petersburg: 156.
13. "Introduction," Chure (2001). Pg. 20.
14. Olshevsky, George. "Dinogeorge's Dinosaur Genera List". Archived from the original on 2011-07-15. Retrieved 2008-08-07.
15. Sampson, S.D. (1995). "Two new horned dinosaurs from the Upper Cretaceous Two Medicine Formation of Montana, with a phylogenetic analysis of the Centrosaurinae (Ornithischia: Ceratopsidae)". Journal of Vertebrate Paleontology. 15 (4): 743–760. Bibcode:1995JVPal..15..743S. doi:10.1080/02724634.1995.10011259.
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20. Accarie, H., B. Beaudoin, J. Dejax, G. Fries, J.-C. Michard, and P. Taquet. 1995. Decouverte d’un Dinosaure Theropode nouveau (Genusaurus sisteronis n. g., n. sp.) dans l’Albien marin de Sisteron (Alpes de Haute-Provence, France) et extension au Cretace inferieur de la lignee ceratosaurienne. Compte rendu hebdomadaire des séances de l’Académie des Sciences Paris, tomo 320, 2nd series : pp. 327-334.
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22. Hunt, A.P., M.G. Lockley, S.G. Lucas, and C.A. Meyer. 1995 [George Olshevsky notes 1995, not 1994]. The global sauropod fossil record. In:Aspects of sauropod paleobiology (M.G. Lockley, V.F. dos Santos, C.A. Meyer, and A. Hunt, eds,). Revista de Geociencias, Gaia 10: pp. 261-279.
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24. Zhang, Y., and Z. Yang. 1994 [George Olshevsky notes 1995, not 1994]. A new complete ostology of Prosauropoda in Lufeng Basin Yunnan China, Jingshanosaurus. Yunnan Publishing House of Science and Technology, Kunming, China: pp. 1-100.
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26. Carpenter, K.; Dilkes, D.; Weishampel, D.B. (1995). "The Dinosaurs of the Niobrara Chalk Formation (Upper Cretaceous, Kansas)". Journal of Vertebrate Paleontology. 15 (2): 275–297. Bibcode:1995JVPal..15..275C. doi:10.1080/02724634.1995.10011230.
27. Chure, D.J. 1995. A reassessment of the gigantic theropod Saurophagus maximus from the Morrison Formation (Upper Jurassic) of Oklahoma, USA. Sixth Symposium on Mesozoic Terrestrial Ecosystems and Biota: pp. 103-106.
28. Coombs, W.P. 1995. A nodosaurid ankylosaur (Dinosauria: Ornithischia) from the Lower Cretaceous of Texas. J. Vertebr. Paleontol. 15 (2):pp. 298-312.
29. Steven D. Emslie (1995). "An Early Irvingtonian Avifauna from Leisey Shell Pit". Bulletin of the Florida Museum of Natural History. 37 (1): 299–344.
30. Herculano M. F. de Alvarenga (1995). "Um Primitivo Membro da Ordem Galliformes (Aves) do Terciário Médio da Bacia de Taubaté, Estado de São Paulo, Brasil". Anais da Academia Brasileira de Ciências. 67: 33–44.
31. Brian Mackness (1995). "Anhinga malagurala, A New Pygmy Darter form the Early Pliocene Bluff Downs Local Fauna, North- Eastern Queensland" . Emu. 95 (4): 265–271. Bibcode:1995EmuAO..95..265M. doi:10.1071/mu9950265.
32. David J. Varricchio; Luis M. Chiappe (1995). "A New Enantiornithine Bird from the Upper Cretaceous Two Medicine Formation of Montana". Journal of Vertebrate Paleontology. 15 (1): 201–204. Bibcode:1995JVPal..15..201V. doi:10.1080/02724634.1995.10011219.
33. Zhou Zhonghe (1995). "Discovery of a New Enantiornithine Bird from the Early Cretaceous of Liaoning, China" (PDF). Vertebrata PalAsiatica. 33 (2): 99–113.
34. Hou Lianhai; Zhou Zhonghe; Gu Yucai; Zhang He (1995). "Confuciusornis sanctus, a New Late Jurassic Sauriurine Bird from China". Chinese Science Bulletin. 40: 1545–1551. Archived from the original on 2014-08-09.
35. Jiri Mlikovsky (1995). "Early Pleistoceen Birds of Stránska Skála Hill, Czech Republic: 1. Musil's Talus Cone" (PDF). In: R Musil (Ed): Stránka Skála Hill,. Excavation of Open-Air Sediments 1964-1972. Moravian Museum, Brno, Anthropos Series. 26: 111–126. Archived from the original (PDF) on 2014-05-02. Retrieved 2014-08-08.
36. Kenneth E. Campbell, jr. (1995). "A Review of the Cranes (Aves: Gruidae) of Rancho La Brea, with the Description of a New Species" (PDF). Museum of Natural History of Los Angeles County, Contributions in Science. 452: 1–13. Archived from the original (PDF) on 2016-06-29. Retrieved 2014-08-08.
37. Claudia P. Tambussi (1995). "The Fossil Rheiformes from Argentina". Courier Forschungsinstitut Senckenberg. 181: 121–129.
38. Dieter S. Peters (1995). "Idiornis tuberculata N. Spec. Ein Weiterer Ungewohnlicher Vogel aus der Grube Messel (Aves: Gruiformes: Cariamidae: Idiornithinae)". Courier Forschungsinstitut Senckenberg. 181: 107–119.
39. Cécile Mourer-Chauviré (2002). "Idiornis Oberholser, 1899 (Aves, Gruiformes, Cariamae, Idiornithidae): a Junior Synonym of Dynamopterus Milne-Edwards, 1892 (Paleogene, Phosphorites du Quercy, France)". Neues Jahrbuch für Geologie und Paläontologie. 270 (1): 13–22. doi:10.1127/0077-7749/2013/0355.
40. Cécile Mourer-Chauviré (1995). "The Messelornithidae (Aves: Gruiformes) from the Palaeocene of France". Courier Forschungsinstitut Senckenberg. 181: 95–105.
41. Zlatozar N. Boev (1995). "Middle Villafranchian Birds from Varshets (Western Balkan Range-Bulgaria)". Courier Forschungsinstitut Senckenberg. 181: 259–269.
42. Steven D. Emslie (1995). "A Catastrophic Death Assemblage of a New Species of Cormorant and Other Seabirds from the Late Pliocene of Florida". Journal of Vertebrate Paleontology. 15 (2): 313–330. Bibcode:1995JVPal..15..313E. doi:10.1080/02724634.1995.10011232.
43. Herculano M. F. de Alvarenga (1995). "A Large and Probably Flightless Anhinga from the Miocene of Chile". Courier Forschungsinstitut Senckenberg. 181: 149–161.
44. Walter E. Boles (1995). "A Preliminary Analysis of the Passeriformes from Riversleigh, Northwestern Queensland, Australia, with the description of a New Species of Lyrebird" (PDF). Courier Forschungsinstitut Senckenberg. 181: 163–170.
45. Xue Xiangxu (1995). "Qinornis paleocenica - a Palaeocene Bird Discovered in China". Courier Forschungsinstitut Senckenberg. 181: 89–93.
46. Spassov, Nikolay; Lange-Badré, B. (1995). "Asiavorator altidens gen et sp. nov., un mammifere carnivore nouveau de l'Oligocene superieur de Mongolie". Annales de Paléontologie (in French). 81 (3): 109–123.