Little owl | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Strigiformes |
Family: | Strigidae |
Genus: | Athene |
Species: | A. noctua |
Binomial name | |
Athene noctua (Scopoli, 1769) | |
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Range of the little owl Resident Extant & Introduced (resident) | |
Synonyms | |
Carine noctua |
The little owl (Athene noctua), also known as the owl of Athena or owl of Minerva, is a bird that inhabits much of the temperate and warmer parts of Europe, the Palearctic east to Korea, and North Africa. It was introduced into Britain at the end of the 19th century and into the South Island of New Zealand in the early 20th century.
This owl is a member of the typical or true owl family Strigidae, which contains most species of owl, the other grouping being the barn owls, Tytonidae. It is a small (approx. 22 cm long), cryptically coloured, mainly nocturnal species and is found in a range of habitats including farmland, woodland fringes, steppes and semi-deserts. It feeds on insects, earthworms, other invertebrates and small vertebrates. Males hold territories which they defend against intruders. This owl is a cavity nester and a clutch of about four eggs is laid in spring. The female does the incubation and the male brings food to the nest, first for the female and later for the newly hatched young. As the chicks grow, both parents hunt and bring them food, and the chicks leave the nest at about seven weeks of age.
Being a common species with a wide range and large total population, the International Union for Conservation of Nature has assessed its conservation status as "least concern".
The little owl was formally described in 1769 by the Italian naturalist Giovanni Antonio Scopoli under the binomial name Strix noctua. [3] The little owl is now placed in the genus Athene that was introduced by the German zoologist Friedrich Boie in 1822. [4] [5] The owl was designated as the type species of the genus by George Robert Gray in 1841. [6] [7] The genus name, Athene, commemorates the goddess Athena, whose original role as a goddess of the night might explain the link to an owl. The species name noctua has, in effect, the same meaning, being the Latin name of an owl sacred to Minerva, Athena's Roman counterpart. [8]
The little owl is probably most closely related to the spotted owlet (Athene brama). A number of variations occur over the bird's wide range and there is some dispute over their taxonomy. The most distinct is the pale grey-brown Middle-Eastern type known as the Syrian little owl (A. n. lilith). A 2009 paper in the ornithological journal Dutch Birding (vol. 31: 35–37, 2009) has advocated splitting the southeastern races as a separate species, Lilith's owl (Athene glaux), with subspecies A. g. glaux, A. g. indigena, and A. g. lilith. DNA evidence and vocal patterns support this proposal. [9]
Other forms include another pale race, the north African A. n. desertae, and three intermediate subspecies, A. n. indigena of southeast Europe and Asia Minor, A. n. glaux in north Africa and southwest Asia, and A. n. bactriana of central Asia. Differences in size of bird and length of toes, reasons put forward for splitting off A. n. spilogastra, seem inconclusive; A. n. plumipes has been claimed to differ genetically from other members of the species and further investigation is required. In general, the different varieties both overlap with the ranges of neighbouring groups and intergrade (hybridise) with them across their boundaries. [9]
Thirteen subspecies are recognised: [5]
The little owl is a small owl with a flat-topped head, a plump, compact body and a short tail. The facial disc is flattened above the eyes giving the bird a frowning expression. The plumage is greyish-brown, spotted, streaked and barred with white. The underparts are pale and streaked with darker colour. [10] It is usually 22 cm (8.7 in) in length with a wingspan of 56 cm (22 in) for both sexes, and weighs about 180 g (6.3 oz). [11]
The adult little owl of the most widespread form, the nominate A. n. noctua, is white-speckled brown above, and brown-streaked white below. It has a large head, long legs, and yellow eyes, and its white "eyebrows" give it a stern expression. Juveniles are duller, and lack the adult's white crown spots. This species has a bounding flight like a woodpecker. [10] Moult begins in July and continues to November, with the male starting before the female.
The call is a querulous kiew, kiew. Less frequently, various whistling or trilling calls are uttered. In the breeding season, other more modulated calls are made, and a pair may call in duet. Various yelping, chattering or barking sounds are made in the vicinity of the nest. [10]
The little owl is widespread across Europe, Asia and North Africa. Its range in Eurasia extends from the Iberian Peninsula and Denmark eastwards to China and southwards to the Himalayas. In Africa it is present from Mauritania to Egypt, the Red Sea and Arabia. It was introduced to the United Kingdom [12] in the 19th century, and has spread across much of England and the whole of Wales. It was introduced to Otago in New Zealand by the local acclimatisation society in 1906, and to Canterbury a little later, and is now widespread in the eastern and northern South Island; [13] it is partially protected under Schedule 2 of New Zealand's Wildlife Act 1953, whereas most introduced birds explicitly have no protection or are game birds.
This is a sedentary species that is found in open countryside in a great range of habitats. These include agricultural land with hedgerows and trees, orchards, woodland verges, parks and gardens, as well as steppes and stony semi-deserts. It is also present in treeless areas such as dunes, and in the vicinity of ruins, quarries and rocky outcrops. It sometimes ventures into villages and suburbs. In the United Kingdom it is chiefly a bird of the lowlands, and usually occurs below 500 m (1,600 ft). [10] In continental Europe and Asia it may be found at much higher elevations; one individual was recorded from 3,600 m (12,000 ft) in Tibet. [14]
This owl usually perches in an elevated position ready to swoop down on any small creature it notices. It feeds on prey such as insects and earthworms, as well as small vertebrates including amphibians, reptiles, birds and mammals. It may pursue prey on the ground and it caches surplus food in holes or other hiding places. [12] A study of the pellets of indigestible material that the birds regurgitate found mammals formed 20 to 50% of the diet and insects 24 to 49%. Mammals taken included mice, rats, voles, shrews, moles and rabbits. The birds were mostly taken during the breeding season and were often fledglings, and including the chicks of game birds. The insects included Diptera, Dermaptera, Coleoptera, Lepidoptera and Hymenoptera. Some vegetable matter (up to 5%) was included in the diet and may have been ingested incidentally. [10]
The little owl is territorial, the male normally remaining in one territory for life. However, the boundaries may expand and contract, being largest in the courtship season in spring. The home range, in which the bird actually hunts for food, varies with the type of habitat and time of year. Little owls with home-ranges that incorporate a high diversity of habitats are much smaller (< 2 ha) than those which breed in monotonous farmland (with home-ranges over 12 ha). Larger home-ranges result in increased flight activity, longer foraging trips and fewer nest visits. [15] If a male intrudes into the territory of another, the occupier approaches and emits its territorial calls. If the intruder persists, the occupier flies at him aggressively. If this is unsuccessful, the occupier repeats the attack, this time trying to make contact with his claws. In retreat, an owl often drops to the ground and makes a low-level escape. [16] The territory is more actively defended against a strange male as compared to a known male from a neighbouring territory; it has been shown that the little owl can recognise familiar birds by voice. [17]
The little owl is partly diurnal and often perches boldly and prominently during the day. [14] If living in an area with a large amount of human activity, little owls may grow used to humans and will remain on their perch, often in full view, while people are around. The little owl has a life expectancy of about 16 years. [12] However, many birds do not reach maturity; severe winters can take their toll and some birds are killed by road vehicles at night, [12] so the average lifespan may be on the order of 3 years. [11]
This owl becomes more vocal at night as the breeding season approaches in late spring. The nesting location varies with habitat, nests being found in holes in trees, in cliffs, quarries, walls, old buildings, river banks and rabbit burrows. [14] A clutch of 3 to 5 eggs is laid (occasionally 2 to 8). The eggs are broadly elliptical, white and without gloss; they measure about 35.5 by 29.5 mm (1.40 by 1.16 in). They are incubated by the female who sometimes starts sitting after the first egg is laid. While she is incubating the eggs, the male brings food for her. The eggs hatch after 28 or 29 days. [10] At first the chicks are brooded by the female and the male brings in food which she distributes to them. Later, both parents are involved in hunting and feeding them. The young leave the nest at about 7 weeks, and can fly a week or two later. Usually there is a single brood but when food is abundant, there may be two. [12] The energy reserves that little owl chicks are able to build up when in the nest influences their post-fledgling survival, with birds in good physical condition having a much higher chance of survival than those in poor condition. [18] When the young disperse, they seldom travel more than about 20 km (12 mi). [9] Pairs of birds often remain together all year round and the bond may last until one partner dies. [9]
A. noctua has an extremely large range. It has been estimated that there are between 560 thousand and 1.3 million breeding pairs in Europe, and as Europe equates to 25 to 49% of the global range, the world population may be between 5 million and 15 million birds. The population is believed to be stable, and for these reasons, the International Union for Conservation of Nature has assessed the bird's conservation status as being of "least concern". [1]
Owls have often been depicted from the Upper Palaeolithic onwards, in forms from statuettes and drawings to pottery and wooden posts, but in the main they are generic rather than identifiable to species. The little owl is, however, closely associated with the Greek goddess Athena and the Roman goddess Minerva, and hence represents wisdom and knowledge. A little owl with an olive branch appears on a Greek tetradrachm coin from 500 BC (a copy of which appears on the modern Greek one-euro coin) and in a 5th-century B.C. bronze statue of Athena holding the bird in her hand. The call of a little owl was thought to have heralded the murder of Julius Caesar. [19] [20]
In 1843 several little owls that had been brought from Italy were released by the English naturalist Charles Waterton on his estate at Walton Hall in Yorkshire but these failed to establish themselves. Later successful introductions were made by Lord Lilford on his Lilford Hall estate near Oundle in Northamptonshire and by Edmund Meade-Waldo at Stonewall Park near Edenbridge, Kent. From these areas the birds spread and had become abundant by 1900. [21] The owls acquired a bad reputation and were believed to prey on game bird chicks. They therefore became a concern to game breeders who tried to eliminate them. In 1935 the British Trust for Ornithology initiated a study into the little owl's diet led by the naturalist Alice Hibbert-Ware. The report showed that the owls feed almost entirely on insects, other invertebrates and small mammals and thus posed little threat to game birds. [22] [23]
There is evidence that from the 19th century little owls were occasionally kept as ornamental birds. In Italy, tamed and docked little owls were kept to hunt rodents and insects in the house and garden. [24]
More common was keeping little owls to use them in so-called cottage hunting. This took advantage of the fact that many bird species react to owls with aggressive behaviour when they discover them during the day (mobbing). Such huntings, particularly with tawny owls, were practiced in Italy from 350 B.C. until the 20th century and in Germany from the 17th to the 20th century. [25] In Italy, mainly skylarks were caught in this way. The main place of trade was Crespina, a small town near Pisa. Here, little owls were traditionally sold on 29 September, after being taken from their nests and raised in human care. [25] Only since the 1990s has this trade been officially banned; however, because of the long cultural tradition for hunting with little owls, exemptions are still granted. Thus, there is still a breeding center for little owls near Crespina, which is maintained by hunters. [26] [27]
In 1992, the little owl appeared as a watermark on Jaap Drupsteen’s 100 guilder banknote for the Netherlands. [28]
The whinchat is a small migratory passerine bird breeding in Europe and western Asia and wintering in central Africa. At one time considered to be in the thrush family, Turdidae, it is now placed in the Old World flycatcher family, Muscicapidae. Both sexes have a strong supercilium, brownish upper parts mottled darker, a pale throat and breast, a pale buff to whitish belly, and a blackish tail with white bases to the outer tail feathers, but in the breeding season, the male has an orange-buff throat and breast.
The European nightjar, common goatsucker, Eurasian nightjar or just nightjar, is a crepuscular and nocturnal bird in the nightjar family that breeds across most of Europe and the Palearctic to Mongolia and Northwestern China. The Latin generic name refers to the old myth that the nocturnal nightjar suckled from goats, causing them to cease to give milk. The six subspecies differ clinally, the birds becoming smaller and paler towards the east of the range. All populations are migratory, wintering in sub-Saharan Africa. Their densely patterned grey and brown plumage makes individuals difficult to see in the daytime when they rest on the ground or perch motionless along a branch, although the male shows white patches in the wings and tail as he flies at night.
The Eurasian tree sparrow is a passerine bird in the sparrow family with a rich chestnut crown and nape and a black patch on each pure white cheek. The sexes are similarly plumaged, and young birds are a duller version of the adult. This sparrow breeds over most of temperate Eurasia and Southeast Asia, where it is known as the tree sparrow, and it has been introduced elsewhere including the United States, where it is known as the Eurasian tree sparrow or German sparrow to differentiate it from the native unrelated American tree sparrow. Although several subspecies are recognised, the appearance of this bird varies little across its extensive range.
The hen harrier is a bird of prey. It breeds in Eurasia. The term "hen harrier" refers to its former habit of preying on free-ranging fowl.
The purple heron is a wide-ranging heron species. It breeds in Africa, central and southern Europe, and southern and eastern Palearctic. The Western Palearctic populations migrate between breeding and wintering habitats whereas the African and tropical-Asian populations are primarily sedentary, except for occasional dispersive movements. It is similar in appearance to the more common grey heron but is slightly smaller, more slender and has darker plumage. It is also a more evasive bird, favouring densely vegetated habitats near water, particularly reed beds. It hunts for a range of prey including fish, rodents, frogs and insects, either stalking them or standing waiting in ambush.
The barn owl is the most widely distributed species of owl in the world and one of the most widespread of all species of birds, living almost everywhere except for polar and desert regions, Asia north of the Himalayas, some Indonesian islands and some Pacific Islands. It is also known as the common barn owl, to distinguish it from the other species in its family, Tytonidae, which forms one of the two main lineages of living owls, the other being the typical owls (Strigidae).
The piping plover is a small sand-colored, sparrow-sized shorebird that nests and feeds along coastal sand and gravel beaches in North America. The adult has yellow-orange-red legs, a black band across the forehead from eye to eye, and a black stripe running along the breast line. This chest band is usually thicker in males during the breeding season, and it is the only reliable way to tell the sexes apart. The bird is difficult to see when it is standing still, as it blends well with open, sandy beach habitats. It typically runs in short, quick spurts and then stops.
The little ringed plover is a small plover. The genus name Charadrius is a Late Latin word for a yellowish bird mentioned in the fourth-century Vulgate. It derives from Ancient Greek kharadrios a bird found in river valleys. The specific dubius is Latin for doubtful, since Sonnerat, writing in 1776, thought this bird might be just a variant of common ringed plover.
The long-eared owl, also known as the northern long-eared owl or, more informally, as the lesser horned owl or cat owl, is a medium-sized species of owl with an extensive breeding range. The genus name, Asio, is Latin for "horned owl", and the specific epithet, otus, is derived from Greek and refers to a small eared owl. The species breeds in many areas through Europe and the Palearctic, as well as in North America. This species is a part of the larger grouping of owls known as typical owls, of the family Strigidae, which contains most extant species of owl.
Savi's warbler is a species of Old World warbler in the grass warbler genus Locustella. It breeds in Europe and the western Palearctic. It is migratory, wintering in northern and sub-Saharan Africa.
The eastern bluebird is a small North American migratory thrush found in open woodlands, farmlands, and orchards.
The burrowing owl, also called the shoco, is a small, long-legged owl found throughout open landscapes of North and South America. Burrowing owls can be found in grasslands, rangelands, agricultural areas, deserts, or any other open, dry area with low vegetation. They nest and roost in burrows, such as those excavated by prairie dogs. Unlike most owls, burrowing owls are often active during the day, although they tend to avoid the midday heat. Like many other kinds of owls, though, burrowing owls do most of their hunting during dusk and dawn, when they can use their night vision and hearing to their advantage. Living in open grasslands as opposed to forests, the burrowing owl has developed longer legs that enable it to sprint, as well as fly, when hunting.
The spotted owlet is a small owl which breeds in tropical Asia from North Pakistan to Southeast Asia. A common resident of open habitats including farmland and human habitation, it has adapted to living in cities. They roost in small groups in the hollows of trees or in cavities in rocks and buildings. It nests in a hole in a tree or building, laying 3–5 eggs. The species shows great variation including clinal variation in size and forms a superspecies with the very similar little owl.
The mountain plover is a medium-sized ground bird in the plover family (Charadriidae). It is misnamed, as it lives on level land. Unlike most plovers, it is usually not found near bodies of water or even on wet soil; it prefers dry habitat with short grass and bare ground. Its height is in the range of 5-9 inches, and length in the range of 8-10 inches ; and it weighs around 102 grams.
Verreaux's eagle-owl, also commonly known as the milky eagle owl or giant eagle owl, is a member of the family Strigidae. This species is widespread in sub-Saharan Africa. A member of the genus Ketupa, it is the largest African owl, measuring up to 66 cm (26 in) in total length. This eagle-owl is a resident primarily of dry, wooded savanna. Verreaux's eagle-owl is mainly grey in color and is distinguishable from other large owls by its bright pink eyelids, a feature shared with no other owl species in the world.
The Amazonian pygmy owl, also known as Hardy's pygmy owl, is an owl found in northern South America and the Amazon Basin. A member of the Strigidae family, it is a true owl.
The Asian Barred Owlet also known as the Cuckoo owlet is a species of pygmy owl native to the forests and shrublands of mainland Southeast Asia to the foothills of the Himalayas of northern Pakistan. It is a smaller owl, measuring 22–25 cm (8.7–9.8 in) making it one of the larger pygmy owls. They are primarily insectivorous but will eat lizards, small rodents and birds as well.
The eastern barn owl is usually considered a subspecies group and together with the American barn owl group, the western barn owl group, and sometimes the Andaman masked owl make up the barn owl. The cosmopolitan barn owl is recognized by most taxonomic authorities. A few separate them into distinct species, as is done here. The eastern barn owl is native to southeastern Asia and Australasia.
The western barn owl is usually considered a subspecies group and together with the American barn owl group, the eastern barn owl group, and sometimes the Andaman masked owl make up the barn owl. The cosmopolitan barn owl is recognized by most taxonomic authorities. A few separate them into distinct species, as is done here. The western barn owl is native to Eurasia and Africa.
The American barn owl is usually considered a subspecies group and together with the western barn owl group, the eastern barn owl group, and sometimes the Andaman masked owl, make up the barn owl, cosmopolitan in range. The barn owl is recognized by most taxonomic authorities. A few separate them into distinct species, as is done here. The American barn owl is native to North and South America, and has been introduced to Hawaii.