Panderodus Temporal range: | |
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A body fossil of Panderodus from the Waukesha biota | |
Life restoration of the species Panderodus unicostatus, based on the Waukesha specimen | |
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Genus: | †Panderodus Ethington, 1959 |
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†Panderodus denticulatus | |
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Panderodus Is an extinct genus of jawless fish belonging to the order Conodonta. This genus had a long temporal range, surviving from the middle Ordovician to late Devonian. [1] In 2021, extremely rare body fossils of Panderodus from the Waukesha Biota were described, and it revealed that Panderodus had a more thick body compared to the more slender bodies of more advanced conodonts. [2] It also revealed that this conodont was a macrophagous predator, meaning it went after large prey. [2]
Panderodus was first described in 1959 on the basis of its tooth structures known as conodont elements. [2] In 1985 a poorly preserved body fossil of the species Panderodus unicostatus was discovered in the Waukesha biota, a fossil site located in southeastern Wisconsin. [2] [3] At the time of its discovery this specimen was the one of only two known conodont body fossils, the other one being a body fossil possibly belonging to Clydagnathus which was found in 1983 from Carboniferous rocks of Scotland. [4] More body fossils have since been found, like a large fossil of the species Promissum pulchrum found in the upper Ordovician soom shale in South Africa in the 1990s, but only 11 total are so far known. [2] [5] [6] [7] The Waukesha fossil was known to have belonged to Panderodus, but wasn't fully analyzed until 2021. [2] This fossil revealed that at least P. unicostatus had a thicker body then most other conodonts, and had more primitive shark-like teeth. These features suggest P. unicostatus was a macrophagous hunter, something rare in vertebrates from the Ordovician-Silurian (due to the fact that many of them had no jaws). [2] [5]
Panderodus is a conodont of the family Panderodontidae, with Its closest relative being Besselodus. This conodont was relatively primitive; unlike later conodonts whose teeth formed into a complex sieve, the teeth of Panderodus were large and shark-like. The teeth probably had their own roles, like locking prey into place. Conodonts themselves are an unusual grouping, as for more than a hundred years we only knew of them by their teeth structures. [8] Conodonts are classed in the grouping Agnatha alongside jawless fish like lampreys and hagfish. [9]
Conodonts such as Panderodus were typically small to medium sized, elongate, marine vertebrates that look similar to eels today. [10] These creatures have been classified under a large amount of animals, however now they are recognized as agnathan vertebrates. [11] [2]
The phylogenetic position of Panderodus relative to other conodonts with complete apparatus reconstructions: [2]
Chordata |
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Another study found that some species of Panderodus exhibited body parts that would mean they would be venomous. This makes these conodonts some of the earliest known venomous organisms in the fossil record. [11]
Conodonts are an extinct group of eel-looking agnathan (jawless) vertebrates, classified in the class Conodonta. For many years, they were known only from fossils of their spiky oral elements, which are usually found in isolation and are now called conodont elements, while knowledge about soft tissues remains limited. A resilient group of prehistoric fish, conodonts existed in the world's oceans for over 300 million years, from the Cambrian to the beginning of the Jurassic. Due to their cosmopolitan distribution, conodont elements are widely used as index fossils, fossils used to define and identify geological periods.
The Maotianshan Shales (帽天山页岩) are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation, famous for their Konservat Lagerstätten, deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale of British Columbia, Canada. They take their name from Maotianshan Hill in Chengjiang County, Yunnan Province, China.
Agnatha is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both living (cyclostomes) and extinct species. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes.
Promissum is an extinct genus of conodonts, primitive chordates, that lived during the Upper Ordovician period.
Jamoytius kerwoodi is an extinct species of primitive, eel-like jawless fish known from the Patrick Burn Formation in Scotland, dating to the Llandovery epoch of the Early Silurian period.
Bromalites are the fossilized remains of material sourced from the digestive system of organisms. As such, they can be broadly considered to be trace fossils. The most well-known types of bromalites are fossilized faeces or coprolites. However, other types are recognised, including: regurgitalites ; cololites ; and gastrolites. Regurtitalites and coprolites are thus essentially known only after they have left the body of the producing organisms, whereas gastrolites and cololites are found in situ in their respective organs, but there are rare exceptions.
Onychopterella is a genus of predatory eurypterid, an extinct group of aquatic arthropods. Fossils of Onychopterella have been discovered in deposits from the Late Ordovician to the Late Silurian. The genus contains three species: O. kokomoensis, the type species, from the Early Pridoli epoch of Indiana; O. pumilus, from the Early Llandovery epoch of Illinois, both from the United States; and O. augusti, from the Late Hirnantian to Early Rhuddanian stages of South Africa.
Prioniodontida, also known as the "complex conodonts", is a large clade of conodonts that includes two major evolutionary grades; the Prioniodinina and the Ozarkodinina. It includes many of the more famous conodonts, such as the giant ordovician Promissum (Prioniodinina) from the Soom Shale and the Carboniferous specimens from the Granton Shrimp bed (Ozarkodinina). They are euconodonts, in that their elements are composed of two layers; the crown and the basal body, and are assumed to be a clade.
The Soom Shale is a member of the Late Ordovician (Hirnantian) Cederberg Formation in South Africa, renowned for its remarkable preservation of soft-tissue in fossil material. Deposited in still waters, the unit lacks bioturbation, perhaps indicating anoxic conditions.
Paleontology in Wisconsin refers to paleontological research occurring within or conducted by people from the U.S. state of Wisconsin. The state has fossils from the Precambrian, much of the Paleozoic, some a parts of the Mesozoic and the later part of the Cenozoic. Most of the Paleozoic rocks are marine in origin. Because of the thick blanket of Pleistocene glacial sediment that covers the rock strata in most of the state, Wisconsin’s fossil record is relatively sparse. In spite of this, certain Wisconsin paleontological occurrences provide exceptional insights concerning the history and diversity of life on Earth.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
Proconodontida is an order of conodonts which originated in the late Cambrian (Furongian) and persisted partly through the Ordovician. The ancestral proconodont, Proconodontus, was one of the earliest euconodonts to appear. Proconodonts are often equated with the broader group Cavidonti, which occupies one side of a basal division in the evolution of early euconodonts in the Cambrian. All other euconodonts occupy Conodonti, the other side of the Cambrian split.
Balognathidae is an extinct conodont family.
Notiodella is an extinct conodont genus in the family Balognathidae. It has been described from a 17-element apparatus from the Soom Shale Lagerstätte in South Africa.
The conodont feeding apparatus is a series of phosphatic-mineralized elements, resembling a set of “teeth”, which are found lining the oral surface of the conodont animal.
Parioscorpio is an extinct genus of arthropod containing the species P. venator known from the Silurian-aged Waukesha Biota of the Brandon Bridge Formation near Waukesha, Wisconsin. This animal has gone through a confusing taxonomic history, being called an arachnid, crustacean, and an artiopodan arthropod at various points. This animal is one of the more famous fossil finds from Wisconsin, due to the media coverage it received based on its original description in 2020 as a basal scorpion.
The Waukesha Biota is an important fossil site located in Waukesha County and Franklin, Milwaukee County within the state of Wisconsin. This biota is preserved in certain strata within the Brandon Bridge Formation, which dates to the early Silurian period. It is known for the exceptional preservation of soft-bodied organisms, including many species found nowhere else in rocks of similar age. The site's discovery was announced in 1985, leading to a plethora of discoveries. This biota is one of the few well studied Lagerstätten from the Silurian, making it important in our understanding of the period's biodiversity. Some of the species are not easily classified into known animal groups, showing that much research remains to be done on this site. Other taxa that are normally common in Silurian deposits are rare here, but trilobites are quite common.
Venustulus is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Venustulus was regarded as part of the clade Prosomapoda. Fossils of the single and type species, V. waukeshaensis, have been discovered in deposits of the Silurian period in Wisconsin, in the United States. Venustulus is one of the few synziphosurine genera with fossil showing evidence of appendages, the other ones being Weinbergina, Anderella and Camanchia. Despite often being aligned close to horseshoe crabs, it has been found that Venustulus and its relatives form a group made up of various basal euchelicerate arthropods more distant to the xiphosurans.
The Winneshiek Shale is a Middle Ordovician (Darriwilian-age) geological formation in Iowa. The formation is restricted to the Decorah crater, an impact crater near Decorah, Iowa. Despite only being discovered in 2005, the Winneshiek Shale is already renowned for the exceptional preservation of its fossils. The shale preserves a unique ecosystem, the Winneshiek biota, which is among the most remarkable Ordovician lagerstätten in the United States. Fossils include the oldest known eurypterid, Pentecopterus, as well as giant conodonts such as Iowagnathus and Archeognathus.
Acheronauta is a genus of extinct worm-shaped arthropod that lived in the early Silurian Waukesha biota fossil site in southeast Wisconsin. This arthropod was first discovered alongside the biota in 1985, but was not fully described until October 2022. This creature was recognized and described as a possible early mandibulate. This description is very important as much of the fauna of the biota remain undescribed, and its discovery has allowed for paleontologists to get a better grasp of the diversity of the arthropod fauna at the site. Multiple phylogenetic analyses were performed, and it was found that this arthropod forms a previously undiscovered clade with the Devonian stem-arthropod Captopodus, and the somewhat enigmatic group Thylacocephala.