Saltrio Formation

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Saltrio Formation
Stratigraphic range: Early Sinemurian
~199–196  Ma
O
S
D
C
P
T
J
K
Pg
N
Cava Salnova.jpg
“Salnova” quarry, where most of the fossils come from
Type Geological formation
Unit ofCalcari Selciferi Lombardi Unit
Sub-units
  • "Viggiù facies"
  • "Saltrio facies"
  • "Poaggia facies"
Underlies Moltrasio Formation
Overlies Tremona Formation
Thickness20 m
Lithology
PrimaryFine to compact arenitic limestones
Location
Coordinates 45°54′N8°54′E / 45.9°N 8.9°E / 45.9; 8.9
Approximate paleocoordinates 33°06′N14°48′E / 33.1°N 14.8°E / 33.1; 14.8
Region Lombardy
Country Italy, Switzerland
Type section
Named for Saltrio
Named byTaramelli [1]
Italy relief location map.jpg
Blue pog.svg
Saltrio Formation (Italy)
Italy Lombardy location map.svg
Blue pog.svg
Saltrio Formation (Lombardy)

The Saltrio Formation (Also known as the Broccatello Formation) is a geological formation in Italy and Switzerland. [2] It dates back to the Early Sinemurian, and would have represented a near-epicontinental to shallow environment, judging by the presence of marine fauna such as the nautiloid Cenoceras . [3] [4] The Fossils of the Formation were described on the late 1880s and revised on 1960s, finding first marine biota, such as Crinoids, Bivalves and other fauna related to Epicontinental basin deposits. [5] [6] It´s sandstones have been historically used in construction. [7]

Contents

Salnova Quarry

Machinery suitable for processing the stone extracted in the Salnova Quarry Cava Salnova - Macchinari.jpg
Machinery suitable for processing the stone extracted in the Salnova Quarry

The main outcrop of the formation, represents an active private extraction site. The first extraction activities of the famous Saltrio stone give back to the times of the ancient Romans, with modern reports of activity in this quarry since 1400. [8] In the Monte Oro area, on the southern slope of Monte Orsa, there were numerous trench quarries which were used to extract this precious rock, used both for structural constructions and for the production of artefacts and artistic works. In more recent times the mining activity has been transformed and we have moved from the extraction of stone for construction to the extraction for the production of stabilized and split crushed stone, useful for the production of motorway foundations and mixtures for the production of asphalt. To date it is the only active quarry where Saltrio stone is extracted. [8]

In today's quarry what is mainly known as the Saltrio Formation emerges, i.e., a group of stratified rocks dating back to the Lower Jurassic. The stratigraphy, however, is much more complex, even if so far no study has focused on this topic. Inside the quarry, Dolomia principale sediment emerges dating back to the Upper Triassic (Norian); yet the succession is dominated by the Saltrio Formation, here 15-20 meters thick. [9]

Above, the Moltrasio Formation emerges, a greyish-brown limestone composed of biocalcarenite and containing widespread nodules of spongolitic silica. This rock is rarely fossiliferous except in the contact areas between the Formations. At the roof of the Moltrasio Fm, a whitish yellow limestone emerges, again of marine-pelagic origin, where there is a lot of micro-diffused silica within the sediment. [9]

Since the early 1900s, fossil finds have been known in the Salnova Quarry and in the various quarry sites present in the surroundings of this site. The first written testimonies, and subsequent revisions, are reported starting from the sixties by Giulia Sacchi Vialli. The scholar describes the fossil faunas of Saltrio by listing and detailing various taxa belonging to ammonoids, nautiloids, gastropods, crinoids, brachiopods and bivalves. [6]

In that period, the great phase of extraction of ornamental stone using manual-mechanical methods had just ended in the quarry. Paleontologists could only recover fossils from the waste flakes near the quarry and therefore the possibility of seeing more specimens was limited to the length of manual operations. In those years, however, the quarry was acquired by Salnova SPA (1969): the purpose of the extracted material, and therefore the extraction method and processing, changed. From classic and manual extraction we move on to the use of heavy mechanical means and extraction with explosives: the moved rubble increases considerably, making it easier to observe other specimens, new lithologies and above all different faunas. [8]

The fauna present at the base of the Saltrio Formation is condensed and includes ammonoids of species attributed to the entire Upper Sinemurian. The taxa attributable to the Lower Sinemurian found in the Saltrio quarries probably come from the base of the formation or have been reworked. [6] According to Sacchi-Vialli, the Formation includes taxa indicative of all the biozones between the Bucklandi Zone (Lower Sinemurian) and the Obtusum Zone, and possibly also of the Oxynotum Zone of the Upper Sinemurian, present at the base of the Formation. [6] The contact between the Main Dolomite and the Saltrio Formation also contains selachian teeth, glauconite and phosphated internal models of ammonites. [10]

Description

Salnova in Saltrio seen from the viewpoint above the quarry Cava Salnova from viewpoint 2022-09-29 02.jpg
Salnova in Saltrio seen from the viewpoint above the quarry

The Saltrio layers represent a unique sedimentary environment that is different from both the "Formazione di Saltrio" and the "Saltrio calcarenite" described by earlier researchers. These layers are characterized by being transgressive deposits, meaning they formed as the sea advanced over previously exposed land. The Saltrio deposits show signs of stratigraphic condensation, which refers to slow sediment accumulation over time, often resulting in hardgrounds, surfaces drilled by marine organisms, and the presence of minerals like Glauconite and Phosphorite. [11] Biologically, these layers are rich in fossils, especially Encrinite (crinoid-rich limestones) and bivalve lumachellas (fossilized bivalve shells). Other marine creatures like cephalopods and brachiopods appear occasionally. Faunas from the condensed Saltrio beds indicate early subsidence in the Hettangian. Additionally, the involutine limestones with a rich ammonite fauna support subsidence during the same period. The sediment composition varies in different areas, often containing reworked material from older rocks. [11] The Saltrio environment was complex, with different layers showing distinct conditions. In some areas, the Saltrio layers blend with the "Broccatello d'Arzo", a related limestone formation, but they can still be separated based on differences in their structure and fossil content. The region also experienced sedimentary discontinuities, where layers were not deposited continuously, likely due to tectonic activity or submarine erosion. [11]

The stratigraphic sequence at the Galli quarry, located at an elevation of approximately 700 meters on the southeastern flank of the ridge above Saltrio, represents one of the most detailed exposures of the Saltrio Formation. [12] This section, reaching a total thickness of about 17 meters, begins atop underlying dolomite and consists of a series of carbonate-dominated layers that reflect varying depositional conditions in a marine setting. [12]

Closeup of the Saltrio Limestones Cava Salnova - la pietra.jpg
Closeup of the Saltrio Limestones

At the base, a thin dolomitic breccia layer (up to 1 meter thick) contains angular dolomite fragments embedded in a lighter calcareous-dolomitic matrix. This is overlain by a 0.3-meter-thick marly limestone with minor detrital components, displaying an olive-gray to greenish hue and iron oxide stains. Next is a 0.8-meter saccharoidal limestone with sparse marl, glauconite, and quartz grains, followed by a thin (0.1–0.2 meter) reddish-brown clay horizon. [12]

Above this, a 1-meter oolitic limestone features intact and broken ooids in a compact calcareous cement, grading from white to grayish-yellow. This transitions into a 3-meter unit of finely to coarsely detrital marly limestone rich in organic fragments, shifting from gray-pink at the base to yellowish upward, with iron oxide patches. A 0.8-meter calcareous breccia with diverse clasts and ooids follows, exhibiting irregular surfaces. [12]

The upper part includes a 5.5-meter marly limestone with minimal detritus, progressing from gray-ashy at the base to dark smoky due to bituminous content. The sequence concludes with a 5-meter dark limestone containing chert nodules, which become more abundant and marly toward the top, before passing into overlying cherty limestones approximately 200 meters thick. [12]

Paleoenvironment

Nautiloid found in situ on the Salnova Quarry Nautiloide appena estratto dalla Cava Salnova.jpg
Nautiloid found in situ on the Salnova Quarry

From the Hettangian to earliest Sinemurian, the western Lombardy Basin featured a notable continental area, wider than previously estimated, characterized by a warm, humid paleoclimate. This region transitioned gradually from Upper Rhaetian shallow-water carbonates to Lombardian siliceous limestones and thick Lowermost Jurassic. Dinosaur fossils from the Saltrio Formation may have been transported from this continental zone or the Arbostora swell in Switzerland, an emerged structural high that divided the subsiding Monte Nudo (east) and Monte Generoso (west) basins. The swell overlay a carbonate platform connected to broader shallow-water areas extending west to southeast, forming a large northern gulf influenced by horst and graben tectonics. Outcrops of "terra rossa" paleosols, including those at Castello Cabiaglio-Orino, west of Saltrio, indicate emerged lands during the Hettangian-Sinemurian, with forests covering the modern Lake Maggiore area, evidenced by large plant fragments in the coeval/younger Moltrasio Formation. [13] [14] Recovered flora from Cellina and Arolo on Lake Maggiore's eastern side includes Bennettitales ( Ptilophyllum ) and Conifers ( Pagiophyllum , Brachyphyllum ), suggesting inland dry-warm conditions. [15]

Calcareous Sponge-dominated bioherms represent the main facies of the Saltrio Formation Sponges at SAS Transvaal DSC09129.JPG
Calcareous Sponge-dominated bioherms represent the main facies of the Saltrio Formation

In the early Sinemurian, the Arbostora swell submerged into a shallow open sea (ramp-slope), still bordered south and southwest by emerged land. Dinosaur bones were likely washed into the Monte Nudo gulf during this phase, supported by terrigenous sands from eroded igneous/metamorphic rocks and terrestrial plants in the limestones. While plants (mainly Araucariaceae and Bennettitales leaves/branches) and sands could have been transported by wind or currents, the nearest major continental land was the Sardinia-Corsica mountains, tens of kilometers WNW. [16] Coeval dinosaur footprints in Trento province (160 km east of Saltrio) challenge the traditional atoll model, indicating larger emerged lands for sustaining herbivorous prey and fresh water. [17] [18] The Peri-Adriatic platforms likely featured temporary continental bridges connecting Laurasia and Gondwana across central Tethys, enabling migration and potential isolation leading to endemism and dwarfism in terrestrial faunas. [19]

The Saltrio Formation records a shallow open marine depositional environment on a subsiding carbonate platform along the Tethyan passive margin. The 17-meter-thick sequence at the Galli quarry transitions from basal dolomitic breccia (transgressive reworking) through marly and oolitic limestones (higher-energy subtidal shoals) to bituminous, cherty limestones (deeper open-shelf with reduced oxygenation). Faunal assemblages, dominated by cephalopods (ammonites like Asteroceras , nautiloids Cenoceras ), bivalves (Cardinia), gastropods (Pleurotomaria), and brachiopods (Lobothyris punctata), indicate normal-marine, oxygenated waters with deepening trends. [12] Fragmentary ichthyosaur remains and bioeroded dinosaur bones (e.g., Saltriovenator zanellai) suggest transport from nearby terrestrial sources into a proximal slope or ramp, that is, an open subtidal zone some dozen of meters depth reached by the effects of storm waves and with constant bottom currents. [4] The Broccatello sector represents a deeper-water carbonate mound environment formed during episodes of platform drowning, made up of stromatactis limestones originated through early diagenetic processes involving organic matter decay, with abundant sponges, along brachiopods, crinoids, and mollusks, lacking photic-dependent organisms. The facies reflect a benthic, sponge reef-dominated carbonate factory in a deeper setting. [20]

Regional studies link this to platform drowning amid rifting, with carbon-isotope excursions implying volcanic influences and ocean perturbations. A modern analogue is the Bahama Banks, featuring oolitic shoals and lagoons in a subtropical passive-margin setting. [12]

Invertebrate fauna

Porifera

GenusSpeciesMaterialLocationNotesImages
Corynella [20] [21]
  • C. spp.
Spicules & Imprints
  • Broccatello of Arzo
A Calcareous sponge of the family Endostomatidae.

Endostoma [21]

  • E. spp.
Spicules & Imprints
  • Broccatello of Arzo

A Calcareous sponge of the family Endostomatidae.

Stellispongia [21]
  • S. spp.
Spicules & Imprints
  • Broccatello of Arzo
A Calcareous sponge of the family Stellispongiidae.
Neuropora [22]
  • N. cf. undulata
  • N. spp.
Spicules & Imprints
  • Cava Salnova quarry
  • Galli Quarry
A Demosponge of the family Neuroporidae. A notorious reef developing genus

Brachiopoda

GenusSpeciesMaterialLocationNotesImages
Arzonellina [23]
  • A. exotica
Shells
  • Broccatello of Arzo
A Terebratulidan of the family Arzonellinidae.
Aulacothyris [21]
  • A. waterhousi
Shells
  • Broccatello of Arzo
A Terebratulidan of the family Zeilleriidae.

Cirpa [6] [21] [24] [22]

  • C. fronto
  • C. briseis
  • C. variabilis
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

A Rhynchonellidan of the family Wellerellidae. Identified originally as "Rhynchonella variabilis".

Furcirhynchia [21]
  • F. cf. melvillei
Shells
  • Broccatello of Arzo
A Rhynchonellidan of the family Rhynchonellidae.

Liospiriferina [21] [24] [22]

  • L. rostrata
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

A Rhynchonellidan of the family Spiriferinidae. Was identified originally as "Spiriferina haasi".

Lobothyris [6] [21] [24] [22]

  • L. punctata
  • L. andleri
  • L. subpunctata
  • L. ovatissinia
  • L. sp.
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry
  • Galli Quarry

A Terebratulidan of the family Lobothyrididae. Was identified originally as "Terebratula punctata".

Prionorhynchia [21] [23] [25]
  • P. belemnitica
  • P. greppini
  • P. flabellum
  • P. ssp.
Shells
  • Broccatello of Arzo
A Rhynchonellidan of the family Prionorhynchiidae.
Rimirhynchia [21]
  • R. aff. anglica
Shells
  • Broccatello of Arzo
A Rhynchonellidan of the family Rhynchonellidae.

Rhynchonella [6] [21] [24] [22]

  • R. acanthica
  • R. belemnitica
  • R. latifrons
  • R. rimata
  • R. cf.polyptycha
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry
  • Galli Quarry

A Rhynchonellidan of the family Wellerellidae.

La terre avant le deluge, 1863 "Rhynchonella sulcata". (4513809761).jpg

Rhynchonellina [21]
  • R. alpina
  • R. ssp.
Shells
  • Broccatello of Arzo
A Rhynchonellidan of the family Dimerellidae.
Tetrarhynchia [21]
  • T. spp.
Shells
  • Broccatello of Arzo
A Rhynchonellidan of the family Tetrarhynchiidae.

Spiriferina [6] [21] [24] [22]

  • S. expansa
  • S. hartmanni
  • S. oxigona
  • S. walcotti
  • S. rupestris
  • S. rostrata
  • S. haasi
  • S. tumida
  • S. cf. spirigeroides
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

A Rhynchonellidan of the family Spiriferinidae.

Sulcirostra [23]
  • S. alpina
Shells
  • Broccatello of Arzo
A Rhynchonellidan of the family Dimerellidae.
Viallithyris [24]
  • V. gozzanensis
  • V. adnetensis
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry
A Rhynchonellidan of the family Rhynchonellidae.
Zeilleria [6] [21]
  • Z. sarthacensis
  • Z. choffati
  • Z. cor
  • Z. ewaldi
  • Z. stapia
  • Z. cornuta
  • Z. subnumismalis
  • Z. (Zeilleria) indentata
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry
  • Galli Quarry
A Terebratulidan of the family Zeilleriidae.

Bryozoa

GenusSpeciesMaterialLocationNotesImages

Ceriopora [6]

  • C.? cf. orbignyi
Imprints
  • Cava Salnova quarry
  • Galli Quarry

A Cyclostomatidan of the family Cerioporidae.

Bivalves

GenusSpeciesMaterialLocationNotesImages

Astarte [22] [26]

  • A. praeobliqua
Shells
  • Cava Salnova quarry

A Clam of the family Astartidae. Some shells identified as Cardium probably belong to this genus.

Avicula [5] [22] [26]

  • A. sinemuriensis
  • A. (Oxytoma) dumortieri
Shells
  • Cava Salnova quarry
  • Galli Quarry

A Pearl Oyster of the family Pteriidae.

Calvaentolium [22] [26]

  • C. hehlii
Shells
  • Cava Salnova quarry

A Scallop of the family Pectinoidae. It was identified originally as "Pecten (Pseudamusium) hehlii".

Cardinia [5] [22] [26]

  • C. hybrida
  • C. similis
  • C. rugosa
  • C. hybrida
  • C. concinna
  • C. depressa
  • C. spp.
Shells
  • Cava Salnova quarry
  • Galli Quarry

A Clam of the family Cardiniidae.

Cardium [5] [22] [26]

  • C.? italicum
  • C. cf. multicostatum
  • C. spp.
Shells
  • Cava Salnova quarry

A Clam of the family Cardiniidae.

Entolium [5]
  • E. hehli
  • E. calvum
  • E. disciforme
Shells
  • Cava Salnova quarry
  • Galli Quarry
A Scallop of the family Pectinoidae.

Chlamys [5] [22] [26]

  • C. textoria
Shells
  • Cava Salnova quarry
  • Galli Quarry

A Scallop of the family Pectinoidae It was identified as "Pecten (Chlamys) textorius".

Fimbria [22] [26]

  • F. semireticulata
  • F. (Sphaeriola) sp.
Shells
  • Cava Salnova quarry

A Clam of the family Lucinidae.

Gresslya [5]
  • G. lunulata
Shells
  • Cava Salnova quarry
  • Galli Quarry
A Clam of the family Pleuromyidae.

Goniomya [22] [26]

  • G. verbana
Shells
  • Cava Salnova quarry

A Clam of the family Lucinidae.

Gryphaea [22] [26]

  • G. arcuata
Shells
  • Cava Salnova quarry

An Oyster of the family Gryphaeidae.

Gryphaea arcuata fossil oyster (Blue Lias, Lower Jurassic; coastal cliffs near Lyme Regis, far-western Dorset County, southwestern England) 1 (15206675956).jpg

Lima [5] [22] [26]

  • L. (Radula) succincta
  • L. (Radula) valmariannae
  • L. (Radula) hettangiensis
  • L. spp.
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

A File Clam of the family Limidae.

Lucina [22] [26]

  • L. ? liasina
Shells
  • Cava Salnova quarry

A Clam of the family Lucinidae.

Mactromya [5]
  • M. arenacea
  • M. cingulata
  • M. problematica
Shells
  • Cava Salnova quarry
  • Galli Quarry
An Adapedont of the family Edmondiidae.

Modiolus [5] [22] [26]

  • M. vomer
  • M. cf.scalpra
Shells
  • Cava Salnova quarry
  • Galli Quarry

A Mussel of the family Mytilidae. Identified as the genus "Modiola", now junior synonym of Modiolus.

Myoconcha [5] [22] [26]

  • M. scabra
  • M. ssp.
Shells
  • Cava Salnova quarry
  • Galli Quarry

A Clam of the family Pleuromyidae.

Ostrea [22] [26]

  • O. cf. chillyensis
  • O. electra
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

An Oyster of the family Ostreidae.

Parallelodon [5]
  • P. (Grammatodon) cypriniforme
Shells
  • Cava Salnova quarry
  • Galli Quarry
A Clam of the family Parallelodontidae.

Pecten [22] [26]

  • P. (Pseudamusium) diblasii
  • P. (Pseudamusium) hehlii
  • P. (Chlamys) subalpinus
  • P. (Chlamys) textorius
  • P. spp.
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

A Scallop of the family Pectinoidae.

Pecten fossil.jpg

Pholadomya [22] [26]

  • P. sp.
Shells
  • Cava Salnova quarry

A Clam of the family Pholadomyidae.

Plagiostoma [5] [22] [26]

  • P. giganteum
  • P. stabilei
  • P. spp.
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

A File Clam of the family Limidae. Identified originally as "Lima (Plagiostoma) gigantea".

Plagiostoma gigantea.JPG

Pleuromya [5] [22] [26]

  • P. cf. angusta
  • P. saltriensis
  • P. galathea
Shells
  • Cava Salnova quarry
  • Galli Quarry

A Clam of the family Pleuromyidae.

Sphaeriola [5] [22] [26]

  • S. cf. kurremolinae
  • S. sp.
Shells
  • Cava Salnova quarry

A Clam of the family Lucinidae. Identified originally as "Fimbria (Sphaeriola) sp.".

Terquemia [22] [26]

  • T. heberti
Shells
  • Cava Salnova quarry

A Clam of the family Prospondylidea.

Gastropods

GenusSpeciesMaterialLocationNotesImages
Anticonulus [27]
  • A. profunde-umbilicatus
Shells
  • Cava Salnova quarry
  • Galli Quarry
A Snail of the family Trochidae.
Bathrotomaria [6]
  • B. cf. trocheata
Shells
  • Cava Salnova quarry
  • Galli Quarry
A Snail of the family Pleurotomariidae.
Coelostylina [27]
  • C. lepontina
Shells
  • Cava Salnova quarry
  • Galli Quarry
A Snail of the family Coelostylinidae.
Discohelix [20]
  • D. sp.
Shells
  • Broccatello of Arzo
A Snail of the family Discohelicidae.
Discotoma [6]
  • D.? aff. ornata
Shells
  • Cava Salnova quarry
A Snail of the family Pleurotomariidae.

Pleurotomaria [6] [21] [27] [28]

  • P. oblita
  • P. anglica
  • P. escheri
  • P. granulatocincta
  • P. hettangiensis
  • P. cf. gigas
  • P. sp.
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

A Snail of the family Pleurotomariidae.

Pleurotomaria expansa 01.JPG

Ptychomphalus [6]
  • P. expansus
Shells
  • Cava Salnova quarry
A Snail of the family Eotomariidae.
Pyrgotrochus [6]
  • P. basilicus
  • P. cf.basilicus
Shells
  • Cava Salnova quarry
  • Galli Quarry
A Snail of the family Pleurotomariidae.

Trochus [27]

  • T. sp
Shells
  • Broccatello of Arzo
  • Cava Salnova quarry

A Snail of the family Trochidae.

Trochotoma [6]
  • T. subturrita
Shells
  • Cava Salnova quarry
A Snail of the family Pleurotomariidae.

Cephalopoda

GenusSpeciesMaterialLocationNotesImages

Agassiceras [2] [29]

  • A. nodosaries
  • A. aff. scipionianum
Shells
  • Cava Salnova quarry

An Ammonitidan of the family Arietitidae.

Arietites [2] [29]

  • A. bucklandi
  • A. raricostatus
  • A. ceratitoides
  • A. kridioides
  • A. dimorphus
  • A. arnoui
  • A. conybeari
  • A. sauzeanus
Shells
  • Cava Salnova quarry

An Ammonitidan of the family Arietitidae.

Arietites bucklandi Ammonite (Arietites bucklandi), Lee Kong Chian Natural History Museum (103314).jpg
Arietites bucklandi
Arnioceras [2]
  • A. cf. bodleyi
Shells
  • Cava Salnova quarry
  • Galli Quarry
An Ammonitidan of the family Arietitidae.
Asteroceras [2]
  • A. obtusum
  • A. stellare
  • A. retusum
  • A. confusum
Shells
  • Cava Salnova quarry
  • Burgioli quarry
  • Galli quarry
An Ammonitidan of the family Arietitidae.
Asteroceras Asteroceras BW.jpg
Asteroceras
Aulacoceras [6]
  • A. ("Ausseites") italicus
Phagmocones
  • Cava Salnova quarry
  • Galli Quarry
A Belemnite of the family Aulacoceratidae.

Cenoceras [30] [26] [22]

  • C. amorettii
  • C. stoppanii
  • C. intermedium
  • C. arare
  • C. spreaficoi
  • C. breislacki
  • C. striatum
  • C. sturi
  • C. balsamocrivellii
  • C. affinis
  • C. brancoi
  • C. ornatus
Shells
  • Cava Salnova quarry
  • Galli Quarry

A Nautilidan of the family Cenoceratidae. Cenoceras was identified as member of the genus Nautilus originally.

Cenoceras reconstruction Cenoceras NT.jpg
Cenoceras reconstruction

Coroniceras [2] [29]

  • C. cf.gmuendense
  • C. rotiformis
  • C. bucklandi
  • C. orbiculatus
  • C. bisulcatus
Shells
  • Cava Salnova quarry

An Ammonitidan of the family Arietitidae.

Coroniceras specimen Coroniceras rotiforme.JPG
Coroniceras specimen
Crucilobiceras [12]
  • C. cf. tardarmatum
Shells
  • Galli Quarry
An Ammonitidan of the family Eoderoceratidae.
Eparietites [21]
  • E. impendens
Shells
  • Cava Salnova quarry
  • Burgioli quarry
  • Galli Quarry
An Ammonitidan of the family Arietitidae
Euasteroceras [2]
  • E. cf.turneri
Shells
  • Cava Salnova quarry
  • Galli Quarry
An Ammonitidan of the family Arietitidae
Lytoceras [12]
  • L. sp.
Shells
  • Galli Quarry
An Ammonitidan of the family Lytoceratidae
Microderoceras [2]
  • M. birchii
Shells
  • Cava Salnova quarry
An Ammonitidan of the family Eoderoceratidae.
Microderoceras specimen Microderoceras birchi 348.jpg
Microderoceras specimen
Nannobelus [6]
  • N. cf. acutus
Phagmocones
  • Cava Salnova quarry
  • Galli Quarry
A Belemnite of the family Belemnitidae.

Oxynoticeras [29]

  • O. oxynotum
  • O. actinotum
Shells
  • Cava Salnova quarry
  • Burgioli quarry
  • Galli Quarry

An Ammonitidan of the family Oxynoticeratidae.

Oxynoticeras specimen Oxynoticeras greenoughi.jpg
Oxynoticeras specimen
Paltechioceras [21]
  • P. (Plesechioceras) demissum
  • P.(Plesechioceras) delicatum
Shells
  • Burgioli quarry
An Ammonitidan of the family Echioceratidae.
Paracoroniceras [2]
  • P. cf. gmuendense
Shells
  • Cava Salnova quarry
An Ammonitidan of the family Arietitidae.
Paradasyceras [2]
  • P. stella
Shells
  • Cava Salnova quarry
  • Galli Quarry
An Ammonitidan of the family Juraphyllitidae.
Promicroceras [21]
  • P. planicosta
Shells
  • Burgioli quarry
An Ammonitidan of the family Eoderoceratidae.
Schlotheimia [12]
  • S.(Charmasseiceras?) sp.
Shells
  • Galli Quarry
An Ammonitidan of the family Schlotheimiidae
Vermiceras [2]
  • V. spiratissimum
  • V. cf.spiratissimum
Shells
  • Cava Salnova quarry
An Ammonitidan of the family Arietitidae.
Vermiceras specimen Vermiceras bristol museum.JPG
Vermiceras specimen

Annelida

GenusSpeciesStratigraphic positionMaterialNotesImages
Terebella [20]
  • T. lapilloides
  • T. spp.
  • Broccatello of Arzo
  • Cava Salnova quarry
Tubiform structuresA terebellid, member of the family Terebellidae
Terebella specimen Terebellidae (YPM IZ 080460) 002.jpeg
Terebella specimen

Echinoderms

GenusSpeciesMaterialLocationNotesImages

Isocrinus [31]

  • I. tuberculatus

Multiple ossicles

  • M. Campo dei Fiori

A Sea lily of the family Isocrininae.

Miocidaris [26]

  • M. amalthei

Multiple ossicles

  • Cava Salnova quarry

An Echinoidean of the family Miocidaridae.

Millericrinus [26]

  • M. cf. adneticus

Multiple ossicles

  • Cava Salnova quarry

A Sea lily of the family Millericrinida.

Millericrinus specimen Millericrinus sp 567.jpg
Millericrinus specimen

Palaeocoma [32]

  • P. milleri
MSNVI 044/017, dorsally-ventrally oriented ophiuroid
  • Mount Campo dei Fiori

An Ophiuridan of the family Ophiodermatidae. Extant tropical species like Ophioderma are benthic predators and scavengers that show the same short spines seen in Palaeocoma. [32]

Palaeocoma milleri Palaeocoma milleri 34.jpg
Palaeocoma milleri

Pentacrinites [21] [26]

  • P. tuberculatus
  • P. spp.
  • Fragments
  • Broccatello of Arzo
  • Cava Salnova quarry

A Sea lily of the family Pentacrinitidae.

Pentacrinites Pentacrinites NT.jpg
Pentacrinites
Plegiocidaris [6]
  • P. falsani
Multiple ossicles
  • Cava Salnova quarry
  • Galli Quarry
An Echinoidean of the family Cidaridae.
Plegiocidaris Plegiocidaris cervicalis FOS 2191 - 4.jpg
Plegiocidaris

Vertebrate fauna

In 2016 new vertebrate remains were discovered in the Salnova quarry, the remains are being studied to understand if it is a new dinosaur or some other creature. [33] [34] Latter has been confirmed to be Marine Diapsid material. [35]

Fish

GenusSpeciesMaterialLocationNotesImages
Notidanoides [10]
  • N. arzoensis
  • Isolated Teeth
  • Broccatello of Arzo
  • Cava Salnova quarry
A Crassodontidanidae Hexanchiform

Osteichthyes [4]

Indeterminate

  • Tooth and one jaw fragment
  • Cava Salnova quarry

Non-determined afinitties

Sphenodus [10]
  • S. helveticus
  • S. cf. alpinus
  • S. stschurowskii
  • Isolated Teeth
  • Broccatello of Arzo
  • Cava Salnova quarry
An Orthacodontidae Synechodontiform

Icthyosaurs

GenusSpeciesMaterialLocationNotesImages

Ichthyosaurus [6] [35]

  • I. spp.
  • 3 Vertebrae Imprints
  • Galli Quarry

A Neoichthyosaurian of the family Ichthyosauridae.

Ichthyosaurus 202006 Ichthyosaurus communis.svg
Ichthyosaurus

Temnodontosaurus [1] [35]

  • T. cf.platyodon
  • Isolated Tooth
  • Isolated Vertebrae
  • Cava Salnova quarry

A Neoichthyosaurian of the family Temnodontosauridae. Quoted on the 1880s, specimen that apparently has never been described or figured and whose present repository is unknown

Temnodontosaurus platyodon Temnodontosaurus DB.jpg
Temnodontosaurus platyodon

Pterosaurs

GenusSpeciesMaterialLocationNotesImages

Pterosauria [1] [35]

"Pterodactylus" longirostris

  • Isolated Tooth
  • Cava Salnova quarry

A Pterosaur. Quoted on the 1880s, specimen that apparently has never been fully described or figured and whose present repository is unknown

Dimorphodon, a coeval Pterosaur from UK Dimorphodon2DB.jpg
Dimorphodon, a coeval Pterosaur from UK

Dinosaurs

GenusSpeciesMaterialLocationNotesImages

Saltriovenator [16] [4] [3]

S. zanellai [4] [3]

  • MSNM V3664, very fragmentary and disarticulated skeleton
  • MSNM V3659, one maxillary or dentary tooth (Referred material)
  • Cava Salnova quarry

The oldest known Ceratosaur, falling outside Neoceratosauria, being sister taxa to the maroccan genus Berberosaurus . Saltriovenator is also the largest known Theropod of the Lower Jurassic. It probably was washed into the Sea. [4] Traces on the bones show that the dinosaur carcass remained exposed to the water-sediment interface for months or years, long enough to first be defleshed by mobile scavengers, then colonized by a microbial community that spanned the bone–water interface, which in turn attracted slow-moving grazers and epibionts. [4]

Saltriovenator on an ancient beach SaltriovantorRes 01.jpg
Saltriovenator on an ancient beach


See also

References

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Further reading