Ichthyosauridae | |
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Fossil specimen of Ichthyosaurus breviceps | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | † Ichthyosauria |
Node: | † Thunnosauria |
Family: | † Ichthyosauridae Bonaparte, 1841 |
Type species | |
† Ichthyosaurus communis | |
Subgroups | |
Ichthyosauridae is an extinct family of thunnosaur ichthyosaurs from the latest Triassic and Early Jurassic (Rhaetian to Pliensbachian stages) of Europe, and possibly also from the middle Early Cretaceous (Hauterivian or Barremian stage) of Iraq. Named by Charles Lucien Bonaparte, in 1841, it is usually thought to contain a single genus, Ichthyosaurus , which is known from several species from the Early Jurassic. In 2013, Fischer et al. named and described Malawania anachronus from the middle Early Cretaceous of Iraq. It was found to share several synapomorphies with the type species of this family, Ichthyosaurus communis , and a large phylogenetic analysis recovered these species as sister taxa. Despite its geologically younger age, M. anachronus was also assigned to Ichthyosauridae. [1]
Fischer et al. (2013) applied the name Ichthyosauridae Bonaparte, 1841 for the clade that contains Malawania anachronus and Ichthyosaurus communis . This clade was recovered in the phylogenetic analysis that was based on the largest currently available data matrix of parvipelvian species. Most other phylogenetic analyses, that were based on smaller matrices, also recovered this clade or found M. anachronus to be in a more basal position. Ichthyosauridae was recovered as the sister taxon of Baracromia, a clade first defined by Fischer et al. (2013) that includes all other thunnosaurs. The cladogram below follows the large phylogenetic analysis of Fischer et al. (2013). [1]
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Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs," although the term is also used for wider clades that the order resides in.
Ichthyosaurus is a genus of ichthyosaurs from the Early Jurassic, with possible Late Triassic record, from Europe. It is among the best known ichthyosaur genera, as it is the type genus of the order Ichthyosauria.
Ophthalmosaurus is a genus of ichthyosaur known from the Middle-Late Jurassic. Possible remains from the earliest Cretaceous, around 145 million years ago, are also known. It was a relatively medium-sized ichthyosaur, measuring 4 m (13 ft) long and weighing 930–950 kg (2,050–2,090 lb). Named for its extremely large eyes, it had a jaw containing many small but robust teeth. Major fossil finds of this genus have been recorded in Europe with a second species possibly being found in North America.
Megalosauroidea is a superfamily of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as Megalosaurus bucklandii and all taxa sharing a more recent common ancestor with it than with Allosaurus fragilis or Passer domesticus. Members of the group include Spinosaurus, Megalosaurus, and Torvosaurus. They are possibly paraphyletic in nature with respect to Allosauroidea.
Stenopterygius is an extinct genus of thunnosaur ichthyosaur known from Europe.
Aegirosaurus is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from the late Jurassic and early Cretaceous of Europe. It was originally named as a species of Ichthyosaurus.
Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous worldwide. Almost all ichthyosaurs from the Middle Jurassic onwards belong to the family, until the extinction of ichthyosaurs in the early Late Cretaceous. Opthalmosaurids appeared worldwide during early Bajocian, subsequent to the disappearance of most other ichthyosaur lineages after the end of the Toarcian. Currently, the oldest known ophthalmosaurids is Mollesaurus from the early Bajocian of Argentina, as well as indeterminate remains of the same age from Luxembourg and Canada. Named by George H. Baur, in 1887, the family contains the basal taxa like Ophthalmosaurus. Appleby (1956) named the taxon Ophthalmosauria which was followed by some authors, but these two names are often treated as synonyms; Ophthalmosauridae has the priority over Ophthalmosauria. However, some researchers argue that Ophthalmosauridae should be restricted to the group typically referred to as Ophthalmosaurinae, with classic Platypterygiinae instead being referred to as Undorosauridae or Brachypterygiidae and Ophthalmosauria being used to unite these two groups.
Brachypterygius is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the Late Jurassic of England. The type species was originally described and named as Ichthyosaurus extremus by Boulenger in 1904. Brachypterygius was named by Huene in 1922 for the width and shortness of the forepaddle, and the type species is therefore Brachypterygius extremus. The holotype of B. extremus was originally thought to be from the Lias Group of Bath, United Kingdom, but other specimens suggest it more likely came from the Kimmeridgian Kimmeridge Clay of Kimmeridge Bay, Dorset, UK.
Nannopterygius is an extinct genus of ophthalmosaurid ichthyosaur that lived during the Middle Jurassic to the Early Cretaceous. Fossils are known from England, Kazakhstan, Russia, and Norway and six species are currently assigned to the genus.
Chacaicosaurus is a genus of neoichthyosaurian ichthyosaur known from the Middle Jurassic of Argentina. The single known specimen of this genus was excavated from the Los Molles Formation in Neuquén Province, and is housed at the Museo Olsacher under the specimen number MOZ 5803. This specimen consists of a skull, forelimb, some vertebrae, and some additional postcranial elements. The genus was named by Marta Fernández in 1994, and contains a single species, Chacaicosaurus cayi, making it the first named distinctive ichthyosaur from the Bajocian stage. It is a medium-sized ichthyosaur with a very long snout, which bears a ridge running along each side. The forelimbs of Chacaicosaurus are small and contain four main digits.
Protoichthyosaurus is a genus of ichthyosaur from the early Jurassic of southern England. Two species are known, P. prostaxalis—the type species, named by Appleby in 1979—and P. applebyi. A third species, P. prosostealis, was named by Appleby, but it was removed from the genus in 2017 due to its similarity to Ichthyosaurus. The genus Protoichthyosaurus was synonymized with Ichthyosaurus by Maisch and Hungerbuhler in 1997, and again by Maisch and Matzke in 2000. However, it was found to be distinct in 2017 by Dean Lomax and colleagues, who separated it from Ichthyosaurus on account of differences in the arrangement and shape of the carpal ossifications, as well as the absence of the fifth digit. The species most likely lived during the Hettangian stage, but may have lived as early as the Rhaetian and as late as the Sinemurian.
Sveltonectes is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from Ul’yanovsk region, western Russia.
Thunnosauria is an extinct clade of parvipelvian ichthyosaurs from the Early Jurassic to the early Late Cretaceous (Hettangian–Cenomanian) of Asia, Australia, Europe, North America, and South America. Named by Ryosuke Motani in 1999, it contains the basal taxa Ichthyosaurus and Stenopterygius and the family Ophthalmosauridae. In thunnosaurs, the fore fin is at least twice as long as the hind fin.
Acamptonectes is a genus of ophthalmosaurid ichthyosaurs, a type of dolphin-like marine reptiles, that lived during the Early Cretaceous around 130 million years ago. The first specimen, a partial adult skeleton, was discovered in Speeton, England, in 1958, but was not formally described until 2012 by Valentin Fischer and colleagues. They also recognised a partial subadult skeleton belonging to the genus from Cremlingen, Germany, and specimens from other localities in England. The genus contains the single species Acamptonectes densus; the generic name means "rigid swimmer" and the specific name means "compact" or "tightly packed".
Platypterygiinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the early Late Jurassic to the early Late Cretaceous of Asia, Australia, Europe, North America and South America. Currently, the oldest known platypterygiine is Brachypterygius. Platypterygiines were characterized by square tooth roots in cross-section, an extremely reduced extracondylar area of the basioccipital, prominent dorsal and ventral trochanters on humerus and ischiopubis lacking an obturator foramen.
Ophthalmosaurinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the Middle Jurassic to the late Early Cretaceous of Europe, North America and South America. Currently, the oldest and the basalmost, known as ophthalmosaurine is Mollesaurus from the early Bajocian of Argentina. Ophthalmosaurines were characterized by a large extracondylar area of the basioccipital in the form of a thick and concave peripheral band, posterodistally deflected ulnar facet of the humerus, large ulna with concave and edgy posterior surface and ischiopubis with obturator foramen.
Malawania is an extinct genus of basal thunnosaur ichthyosaur known from the middle Early Cretaceous of Iraq. Malawania was named by Valentin Fischer, Robert M. Appleby, Darren Naish, Jeff Liston, Riding, J. B., Brindley, S. and Pascal Godefroit in 2013 and the type species is Malawania anachronus. It is unusual as it is much more primitive than other Cretaceous ichthyosaurs, being most closely related to Ichthyosaurus from the Late Triassic and Early Jurassic, over 70 million years earlier than Malawania, with all other known ichthyosaurs from the Late Jurassic onwards belonging to the family Ophthalmosauridae.
Sisteronia is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the 'middle' Cretaceous of southeastern England and southeastern France. It contains a single species, Sisteronia seeleyi.
The Ichthyosauromorpha are an extinct clade of marine reptiles consisting of the Ichthyosauriformes and the Hupehsuchia, living during the Mesozoic.
This timeline of ichthyosaur research is a chronological listing of events in the history of paleontology focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, sharks, or swordfish. Scientists have documented ichthyosaur fossils at least as far back as the late 17th century. At that time, a scholar named Edward Lhuyd published a book on British fossils that misattributed some ichthyosaur vertebrae to actual fishes; their true nature was not recognized until the 19th century. In 1811, a boy named Joseph Anning discovered the first ichthyosaur fossils that would come to be scientifically recognized as such. His sister Mary would later find the rest of its skeleton and would go on to become a respected fossil collector and paleontologist in her own right.