Ichthyosauridae

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Ichthyosauridae
Temporal range: latest Triassic-Early Jurassic, 207–185  Ma
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Possible Early Cretaceous ghost lineage
Ichthyosaurus breviceps 2.jpg
Fossil specimen of Ichthyosaurus breviceps
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Ichthyosauria
Node: Thunnosauria
Family: Ichthyosauridae
Bonaparte, 1841
Type species
Ichthyosaurus communis
Subgroups

Ichthyosauridae is an extinct family of thunnosaur ichthyosaurs from the latest Triassic and Early Jurassic (Rhaetian to Pliensbachian stages) of Europe, and possibly also from the middle Early Cretaceous (Hauterivian or Barremian stage) of Iraq. Named by Charles Lucien Bonaparte, in 1841, it is usually thought to contain a single genus, Ichthyosaurus , which is known from several species from the Early Jurassic. In 2013, Fischer et al. named and described Malawania anachronus from the middle Early Cretaceous of Iraq. It was found to share several synapomorphies with the type species of this family, Ichthyosaurus communis , and a large phylogenetic analysis recovered these species as sister taxa. Despite its geologically younger age, M. anachronus was also assigned to Ichthyosauridae. [1]

Contents

Phylogeny

Fischer et al. (2013) applied the name Ichthyosauridae Bonaparte, 1841 for the clade that contains Malawania anachronus and Ichthyosaurus communis . This clade was recovered in the phylogenetic analysis that was based on the largest currently available data matrix of parvipelvian species. Most other phylogenetic analyses, that were based on smaller matrices, also recovered this clade or found M. anachronus to be in a more basal position. Ichthyosauridae was recovered as the sister taxon of Baracromia, a clade first defined by Fischer et al. (2013) that includes all other thunnosaurs. The cladogram below follows the large phylogenetic analysis of Fischer et al. (2013). [1]

Life reconstruction of Ichthyosaurus anningae Ichthyosaurus anningae trio NT small.jpg
Life reconstruction of Ichthyosaurus anningae

Cladogram

Mikadocephalus gracilirostris

  Parvipelvia  

Hudsonelpidia brevirostris

Macgowania janiceps

  Neoichthyosauria  

Temnodontosaurus

Leptonectes tenuirostris

Eurhinosaurus longirostris

Excalibosaurus costini

Suevoleviathan disinteger

Hauffiopteryx typicus

Thunnosauria  
 Ichthyosauridae 

Ichthyosaurus communis

Malawania anachronus

  Baracromia  

Stenopterygius quadriscissus

Chacaicosaurus cayi

  Ophthalmosauridae  

Arthropterygius chrisorum

  Ophthalmosaurinae  

Mollesaurus periallus

Ophthalmosaurus icenicus (type species)

Baptanodon natans ("O." natans)

Acamptonectes densus

  Platypterygiinae  

Brachypterygius extremus

Maiaspondylus lindoei

Aegirosaurus leptospondylus

Sveltonectes insolitus

"Platypterygius" hercynicus

Caypullisaurus bonapartei

Athabascasaurus bitumineus

"Platypterygius" australis

Related Research Articles

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Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs," although the term is also used for wider clades that the order resides in.

<i>Ichthyosaurus</i> Genus of extinct marine reptile, type genus of Ichthyosauria

Ichthyosaurus is a genus of ichthyosaurs from the Early Jurassic, with possible Late Triassic record, from Europe. It is among the best known ichthyosaur genera, as it is the type genus of the order Ichthyosauria.

<i>Ophthalmosaurus</i> Extinct genus of reptiles

Ophthalmosaurus is a genus of ichthyosaur known from the Middle-Late Jurassic. Possible remains from the earliest Cretaceous, around 145 million years ago, are also known. It was a relatively medium-sized ichthyosaur, measuring 4 m (13 ft) long and weighing 930–950 kg (2,050–2,090 lb). Named for its extremely large eyes, it had a jaw containing many small but robust teeth. Major fossil finds of this genus have been recorded in Europe with a second species possibly being found in North America.

<span class="mw-page-title-main">Megalosauroidea</span> Extinct superfamily of Dinosaurs

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<i>Stenopterygius</i> Extinct genus of reptiles

Stenopterygius is an extinct genus of thunnosaur ichthyosaur known from Europe.

<i>Aegirosaurus</i> Extinct genus of reptiles

Aegirosaurus is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from the late Jurassic and early Cretaceous of Europe. It was originally named as a species of Ichthyosaurus.

<span class="mw-page-title-main">Ophthalmosauridae</span> Extinct family of reptiles

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<i>Brachypterygius</i> Extinct genus of reptiles

Brachypterygius is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the Late Jurassic of England. The type species was originally described and named as Ichthyosaurus extremus by Boulenger in 1904. Brachypterygius was named by Huene in 1922 for the width and shortness of the forepaddle, and the type species is therefore Brachypterygius extremus. The holotype of B. extremus was originally thought to be from the Lias Group of Bath, United Kingdom, but other specimens suggest it more likely came from the Kimmeridgian Kimmeridge Clay of Kimmeridge Bay, Dorset, UK.

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Protoichthyosaurus is a genus of ichthyosaur from the early Jurassic of southern England. Two species are known, P. prostaxalis—the type species, named by Appleby in 1979—and P. applebyi. A third species, P. prosostealis, was named by Appleby, but it was removed from the genus in 2017 due to its similarity to Ichthyosaurus. The genus Protoichthyosaurus was synonymized with Ichthyosaurus by Maisch and Hungerbuhler in 1997, and again by Maisch and Matzke in 2000. However, it was found to be distinct in 2017 by Dean Lomax and colleagues, who separated it from Ichthyosaurus on account of differences in the arrangement and shape of the carpal ossifications, as well as the absence of the fifth digit. The species most likely lived during the Hettangian stage, but may have lived as early as the Rhaetian and as late as the Sinemurian.

Sveltonectes is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from Ul’yanovsk region, western Russia.

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Thunnosauria is an extinct clade of parvipelvian ichthyosaurs from the Early Jurassic to the early Late Cretaceous (Hettangian–Cenomanian) of Asia, Australia, Europe, North America, and South America. Named by Ryosuke Motani in 1999, it contains the basal taxa Ichthyosaurus and Stenopterygius and the family Ophthalmosauridae. In thunnosaurs, the fore fin is at least twice as long as the hind fin.

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<span class="mw-page-title-main">Ophthalmosaurinae</span> Extinct subfamily of reptiles

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<i>Malawania</i> Extinct genus of reptiles

Malawania is an extinct genus of basal thunnosaur ichthyosaur known from the middle Early Cretaceous of Iraq. Malawania was named by Valentin Fischer, Robert M. Appleby, Darren Naish, Jeff Liston, Riding, J. B., Brindley, S. and Pascal Godefroit in 2013 and the type species is Malawania anachronus. It is unusual as it is much more primitive than other Cretaceous ichthyosaurs, being most closely related to Ichthyosaurus from the Late Triassic and Early Jurassic, over 70 million years earlier than Malawania, with all other known ichthyosaurs from the Late Jurassic onwards belonging to the family Ophthalmosauridae.

<i>Sisteronia</i> Extinct genus of reptiles

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<span class="mw-page-title-main">Timeline of ichthyosaur research</span>

This timeline of ichthyosaur research is a chronological listing of events in the history of paleontology focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, sharks, or swordfish. Scientists have documented ichthyosaur fossils at least as far back as the late 17th century. At that time, a scholar named Edward Lhuyd published a book on British fossils that misattributed some ichthyosaur vertebrae to actual fishes; their true nature was not recognized until the 19th century. In 1811, a boy named Joseph Anning discovered the first ichthyosaur fossils that would come to be scientifically recognized as such. His sister Mary would later find the rest of its skeleton and would go on to become a respected fossil collector and paleontologist in her own right.

References

  1. 1 2 Fischer, V.; Appleby, R. M.; Naish, D.; Liston, J.; Riding, J. B.; Brindley, S.; Godefroit, P. (2013). "A basal thunnosaurian from Iraq reveals disparate phylogenetic origins for Cretaceous ichthyosaurs". Biology Letters. 9 (4): 20130021. doi:10.1098/rsbl.2013.0021. PMC   3730615 . PMID   23676653.