Shonisaurus Temporal range: Late Triassic (Carnian), | |
---|---|
Restored skull in a Japanese museum | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | † Ichthyosauria |
Family: | † Shastasauridae |
Genus: | † Shonisaurus Camp, 1976 |
Type species | |
†Shonisaurus popularis Camp, 1976 | |
Species [2] [1] | |
|
Shonisaurus is a genus of very large ichthyosaurs. At least 37 incomplete fossil specimens of the marine reptile have been found in the Luning Formation of Nevada, USA. This formation dates to the late Carnian age of the late Triassic period, about 237–227 million years ago. [1]
Shonisaurus lived during the Carnian stage of the late Triassic period. With a large skull about 2.75 m (9.0 ft) long, S. popularis and measured around 13.5–15 metres (44–49 ft) in length and 21.6–29.7 metric tons (23.8–32.7 short tons) in body mass. [3] [4] S. sikanniensis was one of the largest marine reptiles of all time, measuring 21 metres (69 ft) and weighing 81.5 metric tons (89.8 short tons). [3]
Shonisaurus had a long snout, and its flippers were much longer and narrower than in other ichthyosaurs. While Shonisaurus was initially reported to have had socketed teeth (rather than teeth set in a groove as in more advanced forms), these were present only at the jaw tips, and only in the very smallest, juvenile specimens. All of these features suggest that Shonisaurus may be a relatively specialised offshoot of the main ichthyosaur evolutionary line. [5] More recent finds however indicate that Shonisaurus possessed teeth in all ontogenetic stages. [6] Robust sectorial teeth and gut contents indicate that Shonisaurus was a macrophagous raptorial predator which fed on vertebrates and shelled mollusks like cephalopods, possibly even large-bodied prey. [7] [8] Additionally, Shonisaurus was historically depicted with a rather rotund body, but studies of its body shape since the early 1990s have shown that the body was much more slender than traditionally thought, [9] and had a relatively deep body compared with related marine reptiles. [1]
Fossils of Shonisaurus were first found in a large deposit in Nevada in 1920. Thirty years later, they were excavated, uncovering the remains of 37 very large ichthyosaurs. These were named Shonisaurus, which means "lizard from the Shoshone Mountains", after the formation where the fossils were found.
S. popularis, was adopted as the state fossil of Nevada in 1984. Excavations, begun in 1954 under the direction of Charles Camp and Samuel Welles of the University of California, Berkeley, were continued by Camp throughout the 1960s. It was named by Charles Camp in 1976. [10]
The Nevada fossil sites can currently be viewed at the Berlin-Ichthyosaur State Park.
A second species from British Columbia was named Shonisaurus sikanniensis in 2004. [3] However, a phylogenetic study by Sander and colleagues in 2011 later showed S. sikanniensis to be a species of Shastasaurus rather than Shonisaurus. [11] A subsequent study by Ji and colleagues published in 2013 reasserted the original classification, finding it more closely related to Shonisaurus than to Shastasaurus. [12] Support for both hypotheses has been found in later studies, with some authors classifying the species in Shonisaurus and others in Shastasaurus. [13] [14] Specimens belonging to S. sikanniensis have been found in the Pardonet Formation British Columbia, dating to the middle Norian age (about 210 million years ago). [1]
Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides.
Cymbospondylus is an extinct genus of large ichthyosaurs, of which it is among the oldest representatives, that lived during the Anisian stage of the Middle Triassic in what are now North America and Europe. The first known fossils of this taxon are a set of more or less complete vertebrae which were discovered in the 19th century in various mountain ranges of Nevada, in the United States, before being named and described by Joseph Leidy in 1868. It is in the beginning of the 20th century that more complete fossils were discovered through several expeditions launched by the University of California, and described in more detail by John Campbell Merriam in 1908, thus visualizing the overall anatomy of the animal. While many species have been assigned to the genus, only five are recognized as valid, the others being considered synonymous, doubtful or belonging to other genera. Cymbospondylus was formerly classified as a representative of the Shastasauridae, but more recent studies consider it to be more basal, view as the type genus of the Cymbospondylidae.
Mixosaurus is an extinct genus of Middle Triassic ichthyosaur. Its fossils have been found near the Italy–Switzerland border and in South China.
Utatsusaurus hataii is the earliest-known ichthyopterygian which lived in the Early Triassic period. It was nearly 2.5–3 metres (8.2–9.8 ft) long with a slender body. The first specimen was found in Utatsu-cho, Miyagi Prefecture, Japan. It is the only described species in the genus Utatsusaurus and the only member of the family Utatsusauridae. The name Utatsusaurus was given after the city. The fossils have been found from the Early Triassic Osawa Formation of Miyagi Prefecture, Japan and British Columbia, Canada.
Shastasauridae is an extinct family of Triassic ichthyosaurs. The family contains the largest known species of ichthyosaurs, which include some of and possibly the largest known marine reptiles.
Chaohusaurus is an extinct genus of basal ichthyopterygian, depending on definition possibly ichthyosaur, from the Early Triassic of Chaohu and Yuanan, China.
Shastasaurus is an extinct genus of ichthyosaur from the Late Triassic. Specimens have been found in the United States, Canada, and China.
Thalattosauria is an extinct order of prehistoric marine reptiles that lived in the Middle to Late Triassic. Thalattosaurs were diverse in size and shape, and are divided into two superfamilies: Askeptosauroidea and Thalattosauroidea. Askeptosauroids were endemic to the Tethys Ocean, their fossils have been found in Europe and China, and they were likely semiaquatic fish eaters with straight snouts and decent terrestrial abilities. Thalattosauroids were more specialized for aquatic life and most had unusual downturned snouts and crushing dentition. Thalattosauroids lived along the coasts of both Panthalassa and the Tethys Ocean, and were most diverse in China and western North America. The largest species of thalattosaurs grew to over 4 meters (13 feet) in length, including a long, flattened tail utilized in underwater propulsion. Although thalattosaurs bore a superficial resemblance to lizards, their exact relationships are unresolved. They are widely accepted as diapsids, but experts have variously placed them on the reptile family tree among Lepidosauromorpha, Archosauromorpha, ichthyosaurs, and/or other marine reptiles.
Callawayia is an extinct genus of ichthyosaur. It contains the species Callawayia neoscapularis.
Guizhouichthyosaurus is an extinct genus of ichthyosaur which is known primarily from the Xiaowa Formation of the lower Carnian stage of the Late Triassic in southwest China. The type species of this genus is Guizhouichthyosaurus tangae, of which multiple skeletons are known. It has been reassigned as a species of the genus Shastasaurus in the past, though it has since been considered distinct. The ichthyosaurs Cymbospondylus asiaticus, named in 2002, and Panjiangsaurus epicharis, named in 2003, are junior synonyms of G. tangae. The genus is also known from the Ladinian-aged Middle Triassic Zhuganpo Formation; additionally, the species "Callawayia" wollongangense may belong to Guizhouichthyosaurus.
Qianichthyosaurus is an extinct genus of ichthyosaur from the Ladinian and Carnian stages of the Late Triassic epoch. Its fossils have been found in southeastern China, in Carnian rocks of the Falang Formation near Huangtutang, Guizhou. The type species is Qianichthyosaurus zhoui, named by Chun Li in 1999. A second species, Qianichthyosaurus xingyiensis, was named from older (Ladinian) deposits in the Falang Formation in 2013 by Pengfei Yang and colleagues. Complete Qianichthyosaurus fossils are common in the Xiaowa Formation, with both juveniles and pregnant specimens being known; its larger contemporaries, Guizhouichthyosaurus and Guanlingsaurus, are rarer.
The Guanling Formation is a Middle Triassic geologic formation in southwestern China.
Mixosauridae was an early group of ichthyosaurs, living between 247.2 and 235 million years ago, during the Triassic period. Fossils of mixosaurs have been found all over the world: China, Timor, Indonesia, Italy, Germany, Spitsbergen, Switzerland, Svalbard, Canada, Alaska, and Nevada.
Guanlingsaurus is an extinct genus of shastasaurid ichthyosaur from the Late Triassic of China. It grew up to 8.3 metres (27 ft) in length and has a wide, triangular skull with a short and toothless snout.
Cymbospondylidae is an extinct family of Hueneosaurian Ichthyosaurs known from the Middle Triassic of Europe, North America, and Asia.
Merriamosauria is an extinct clade of ichthyosaurs. It was named by Ryosuke Motani in his 1999 analysis of the relationships of ichthyopterygian marine reptiles and was defined in phylogenetic terms as a stem-based taxon including "the last common ancestor of Shastasaurus pacificus and Ichthyosaurus communis, and all of its descendants." The name honours John Campbell Merriam. Based on this definition, Merriamosauria includes most ichthyosaurs except for several Triassic groups such as the clade Mixosauria, the family Cymbospondylidae, and perhaps the family Toretocnemidae. Merriamosaurs are characterized by features in their pectoral girdles and limb bones, including an extensive connection between the scapula and the coracoid bone, the absence of the first metacarpal and the absence of a pisiform bone.
Elizabeth (Betsy) Laura Nicholls was an American-Canadian paleontologist who specialized in Triassic marine reptiles. She was a paleontologist at the Royal Tyrrell Museum in Alberta, Canada.
This timeline of ichthyosaur research is a chronological listing of events in the history of paleontology focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, sharks, or swordfish. Scientists have documented ichthyosaur fossils at least as far back as the late 17th century. At that time, a scholar named Edward Lhuyd published a book on British fossils that misattributed some ichthyosaur vertebrae to actual fishes; their true nature was not recognized until the 19th century. In 1811, a boy named Joseph Anning discovered the first ichthyosaur fossils that would come to be scientifically recognized as such. His sister Mary would later find the rest of its skeleton and would go on to become a respected fossil collector and paleontologist in her own right.