Californosaurus Temporal range: Middle-Late Triassic | |
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Holotype skeleton | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | † Ichthyosauria |
Clade: | † Euichthyosauria |
Genus: | † Californosaurus Kuhn, 1934 |
Type species | |
†Californosaurus perrini Kuhn, 1934 | |
Synonyms | |
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Californosaurus ('California lizard') is an extinct genus of ichthyosaur, an extinct marine reptile, from the Lower Hosselkus Limestone (Carnian, Late Triassic) of California, and also the Muschelkalk (Ladinian, Middle Triassic) of Germany. [1]
John Campbell Merriam (1902) described it as a new species of Shastasaurus , S. perrini, honouring Professor James Perrin Smith as discoverer, based on holotype UCMP 9119. [2] He later recognized the species as generically distinct from the Shastasaurus type species, in 1905 erecting Delphinosaurus for S. perrini. However, Delphinosaurus had in 1853 by Eichwald been previously used for an ophthalmosaurid from Albian-Cenomanian deposits in European Russia, Delphinosaurus kiprijanoffii, and Oscar Kuhn (1934) provided the generic replacement name Californosaurus. Merriam (1938) independently erected Perrinosaurus to replace Delphinosaurus, but this is a junior objective synonym. [3] [4]
The long-snouted head is small in comparison with the rest of the body, as in basal ichthyosaurs such as Mixosaurus and Cymbospondylus . [5] The tail is sharply turned downwards, in common with more advanced ichthyosaurs, with a small vertical fluke. It may have had a small dorsal fin. There is a small number of pre-sacral vertebrae (45 or 50). The phalanges (digit bones) are circular and widely spaced, giving the flipper a round appearance. It was a medium-sized ichthyosaur, measuring up to 3 m (9.8 ft) long. [6]
It fed on fish and other small marine creatures. Like other ichthyosaurs it probably never ventured onto dry land, and gave birth in the water.
Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides.
Cymbospondylus is an extinct genus of large ichthyosaurs, of which it is among the oldest representatives, that lived from the Lower to Middle Triassic in what are now North America and Europe. The first known fossils of this taxon are a set of more or less complete vertebrae which were discovered in the 19th century in various mountain ranges of Nevada, in the United States, before being named and described by Joseph Leidy in 1868. It is in the beginning of the 20th century that more complete fossils were discovered through several expeditions launched by the University of California, and described in more detail by John Campbell Merriam in 1908, thus visualizing the overall anatomy of the animal. While many species have been assigned to the genus, only five are recognized as valid, the others being considered synonymous, doubtful or belonging to other genera. Cymbospondylus was formerly classified as a representative of the Shastasauridae, but more recent studies consider it to be more basal, view as the type genus of the Cymbospondylidae.
Stenopterygius is an extinct genus of thunnosaur ichthyosaur known from Europe.
Eurhinosaurus is an extinct genus of ichthyosaur from the Early Jurassic (Toarcian), ranging between 183 and 175 million years. Fossils of the aquatic reptile have been found in Western Europe. They used to live in the deep, open sea area. Eurhinosaurus was a large genus of ichthyosaurs. An adult individual could reach up to 7 metres (23 ft) in length.
Shastasauridae is an extinct family of ichthyosaurs from the Late Triassic with a possible Early Jurassic record. The family contains the largest known species of ichthyosaurs, which include some of and possibly the largest known marine reptiles.
Delphinosaurus is a dubious genus of ophthalmosaurid ichthyosaur from Albian-Cenomanian deposits in the Kursk region of European Russia.
Shastasaurus is an extinct genus of ichthyosaur from the Late Triassic. Specimens have been found in the United States, Canada, and China.
Omphalosaurus is an extinct genus of marine reptile from the Early Triassic to Middle Triassic, thought to be in the order of Ichthyosauria. Most of what is known about Omphalosaurus is based on multiple jaw fragments, ribs, and vertebrae. Specimens of Omphalosaurus have been described from the western United States, Poland, Austria and the island of Spitsbergen off the northern coast of Norway.
Leptonectes is a genus of ichthyosaur that lived in the Late Triassic to Early Jurassic. Fossils have been found in Belgium, Germany, Spain and the United Kingdom. A possible specimen from Switzerland first reported in 2006, has been reclassified as that of Hauffiopteryx. The type species, L. tenuirostris, reached 4 metres (13 ft) long, while L. moorei reached 3 metres (9.8 ft) long; the largest species, L. solei, was approximately 5–6 metres (16–20 ft) long.
Callawayia is an extinct genus of ichthyosaur. It contains the species Callawayia neoscapularis.
Guizhouichthyosaurus is an extinct genus of ichthyosaur which is known primarily from the Xiaowa Formation of the lower Carnian stage of the Late Triassic in southwest China. The type species of this genus is Guizhouichthyosaurus tangae, of which multiple skeletons are known. It has been reassigned as a species of the genus Shastasaurus in the past, though it has since been considered distinct. The ichthyosaurs Cymbospondylus asiaticus, named in 2002, and Panjiangsaurus epicharis, named in 2003, are junior synonyms of G. tangae. The genus is also known from the Ladinian-aged Middle Triassic Zhuganpo Formation; additionally, the species "Callawayia" wollongangense may belong to Guizhouichthyosaurus.
Hudsonelpidia is an extinct genus of small parvipelvian ichthyosaur known from British Columbia of Canada.
Parvinatator, from Latin, “parvus” little and “natator” swimmer, is an extinct genus of small ichthyopterygian marine reptile that lived during the Early to Middle Triassic. Its fossils have been found in British Columbia, Canada.
Phalarodon is an extinct genus of mixosaurid ichthyosaur known from the Middle Triassic. Its name is derived from the Greek φάλαρα (phálara) and odon ("tooth"). The genus has had a tumultuous history since its classification in 1910, with different workers describing species under different genera or declaring the genus to be a nomen dubium. Currently three species are recognized, but more have been identified in the past.
Qianichthyosaurus is an extinct genus of ichthyosaur from the Ladinian and Carnian stages of the Late Triassic epoch. Its fossils have been found in southeastern China, in Carnian rocks of the Falang Formation near Huangtutang, Guizhou. The type species is Qianichthyosaurus zhoui, named by Chun Li in 1999. A second species, Qianichthyosaurus xingyiensis, was named from older (Ladinian) deposits in the Falang Formation in 2013 by Pengfei Yang and colleagues. Complete Qianichthyosaurus fossils are common in the Xiaowa Formation, with both juveniles and pregnant specimens being known; its larger contemporaries, Guizhouichthyosaurus and Guanlingsaurus, are rarer.
Toretocnemus is an extinct genus of ichthyosaur. Its remains have been found in California, United States, in Triassic layers of the Carnian Hosselkus Limestone.
Guanlingsaurus is an extinct genus of shastasaurid ichthyosaur from the Late Triassic of China. It grew up to 8.3 metres (27 ft) in length and has a wide, triangular skull with a short and toothless snout.
The Hueneosauria are a group of Ichthyosauria, living during the Mesozoic.
This timeline of ichthyosaur research is a chronological listing of events in the history of paleontology focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, sharks, or swordfish. Scientists have documented ichthyosaur fossils at least as far back as the late 17th century. At that time, a scholar named Edward Lhuyd published a book on British fossils that misattributed some ichthyosaur vertebrae to actual fishes; their true nature was not recognized until the 19th century. In 1811, a boy named Joseph Anning discovered the first ichthyosaur fossils that would come to be scientifically recognized as such. His sister, Mary Anning, would later find the rest of its skeleton and would go on to become a respected fossil collector and paleontologist in her own right.