Shastasaurus Temporal range: Late Triassic, | |
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Partial skull of Shastasaurus pacificus (UCMP 9017) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | † Ichthyosauria |
Suborder: | † Longipinnati |
Node: | † Merriamosauria |
Family: | † Shastasauridae Merriam, 1895 |
Genus: | † Shastasaurus Merriam, 1895 |
Type species | |
†Shastasaurus pacificus Merriam, 1895 | |
Species | |
†S. pacificus Contents | |
Synonyms | |
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Shastasaurus ("Mount Shasta lizard") is an extinct genus of ichthyosaur from the Late Triassic. [2] Specimens have been found in the United States, Canada, and China. [3]
Shastasaurus lived during the late Triassic period. The type species, S. pacificus, is known from California, with the name Shastasaurus directly referencing Shasta County, Northern California, where the type specimen was found. [4] S. pacificus was a medium-sized ichthyosaur, measuring over 7 metres (23 ft) in length. [5] A second possible species of Shastasaurus, S. sikanniensis, is known from the Pardonet Formation British Columbia, dating to the middle Norian age (about 210 million years ago). [6] By comparison, S. sikanniensis was one of the largest known ichthyosaurs, similar in size to modern-day cetaceans, measuring up to 21 metres (69 ft) in length and weighing 81.5 metric tons (89.8 short tons). [7]
Shastasaurus was highly specialized, and differed considerably from other ichthyosaurs. It was very slender in profile. S. sikanniensis had a ribcage slightly less than 2 metres (6.6 ft) deep despite a distance of over 7 metres (23 ft) between its flippers. [6] With its unusually short, toothless snout (compared to the longer, toothed, dolphin-like snouts of most ichthyosaurs), it had been proposed that Shastasaurus was a suction feeder (or filled a similar ecological niche as extant beaked whales), preying primarily on soft-bodied cephalopods. [8] However, current research indicates that the jaws of shastasaurid ichthyosaurs do not fit the suction-feeding profile, since their short and narrow hyoid bones are unsuitable to withstand impact forces for such kind of feeding, [9] and since some species like Shonisaurus had robust sectorial teeth with gut contents of mollusk shells and vertebrates. [10] [11]
It is unknown whether Shastasaurus had a dorsal fin; however, the smaller, more basal ichthyosaur Mixosaurus had one. [12] The upper fluke of the tail was probably much less-developed than the shark-like tails found in later species. [13]
The type species of Shastasaurus is S. pacificus, from the late Carnian of northern California. It is known only from fragmentary remains, which have led to the assumption that it was a 'normal' ichthyosaur in terms of proportions, especially skull proportions. Several species of long-snouted ichthyosaur were referred to Shastasaurus based on this misinterpretation, but are now placed in other genera (including Callawayia and Guizhouichthyosaurus ). [8] Elizabeth Nicholls and Makoto Manabe considered this species as a nomen dubium in 2000. [14]
Shastasaurus may include a second species, Shastasaurus liangae. It is known from several good specimens, and was originally placed in the separate genus Guanlingsaurus . Complete skulls show that it had an unusual short and toothless snout. S. pacificus probably also had a short snout, although its skull is incompletely known. [8] However, a new juvenile specimen discovered in 2013 shows that the hyoid bone of Guanlingsaurus is much shorter, and considered it as a distinct genus based on phylogenetic analysis. [15]
S. sikanniensis was originally described in 2004 as a large species of Shonisaurus . However, this classification was not based on any phylogenetic analysis, and the authors also noted similarities with Shastasaurus. The first study testing its relationships, in 2011, supported the hypothesis that it was indeed more closely related to Shastasaurus than to Shonisaurus, and it was reclassified as Shastasaurus sikanniensis. [8] However, a 2013 analysis supported the original classification, finding it more closely related to Shonisaurus than to Shastasaurus. [15] In the 2019 study, S. sikanniensis was pertained within the genus Shastasaurus. [16] In the 2021 analysis, S. sikanniensis forms a clade with Shonisaurus, indicating that it is closer to Shonisaurus than to Shastasaurus. [17] Specimens belonging to S. sikanniensis have been found in the Pardonet Formation British Columbia, dating to the middle Norian age. [6]
In 2009, Shang & Li reclassified the species Guizhouichthyosaurus tangae as Shastasaurus tangae. However, later analysis showed that Guizhouichthyosaurus was in fact closer to more advanced ichthyosaurs, and so cannot be considered a species of Shastasaurus. [8]
Dubious species that were referred to this genus include S. carinthiacus (Huene, 1925) from the Austrian Alps and S. neubigi (Sander, 1997) from the German Muschelkalk. [3] S. neubigi, however, was re-described and reassigned to its own genus, Phantomosaurus . [18]
Synonyms of S. / G. liangae:
Guanlingichthyosaurus [19] liangaeWang et al., 2008 ( lapsus calami )
Synonyms of S. pacificus:
Shastasaurus alexandraeMerriam, 1902
Shastasaurus osmontiMerriam, 1902
Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides.
Shonisaurus is a genus of very large ichthyosaurs. At least 37 incomplete fossil specimens of the type species, Shonisaurus popularis, have been found in the Luning Formation of Nevada, USA. This formation dates to the late Carnian-early Norian age of the Late Triassic, around 227 million years ago. Other possible species of Shonisaurus have been discovered from the middle Norian deposits of Canada and Alaska.
Cymbospondylus is an extinct genus of large ichthyosaurs, of which it is among the oldest representatives, that lived from the Lower to Middle Triassic in what are now North America and Europe. The first known fossils of this taxon are a set of more or less complete vertebrae which were discovered in the 19th century in various mountain ranges of Nevada, in the United States, before being named and described by Joseph Leidy in 1868. It is in the beginning of the 20th century that more complete fossils were discovered through several expeditions launched by the University of California, and described in more detail by John Campbell Merriam in 1908, thus visualizing the overall anatomy of the animal. While many species have been assigned to the genus, only five are recognized as valid, the others being considered synonymous, doubtful or belonging to other genera. Cymbospondylus was formerly classified as a representative of the Shastasauridae, but more recent studies consider it to be more basal, view as the type genus of the Cymbospondylidae.
Mixosaurus is an extinct genus of Middle Triassic ichthyosaur. Its fossils have been found near the Italy–Switzerland border and in South China.
Shastasauridae is an extinct family of ichthyosaurs from the Late Triassic with a possible Early Jurassic record. The family contains the largest known species of ichthyosaurs, which include some of and possibly the largest known marine reptiles.
Thalattosauria is an extinct order of marine reptiles that lived in the Middle to Late Triassic. Thalattosaurs were diverse in size and shape, and are divided into two superfamilies: Askeptosauroidea and Thalattosauroidea. Askeptosauroids were endemic to the Tethys Ocean, their fossils have been found in Europe and China, and they were likely semiaquatic fish eaters with straight snouts and decent terrestrial abilities. Thalattosauroids were more specialized for aquatic life and most had unusual downturned snouts and crushing dentition. Thalattosauroids lived along the coasts of both Panthalassa and the Tethys Ocean, and were most diverse in China and western North America. The largest species of thalattosaurs grew to over 4 meters (13 feet) in length, including a long, flattened tail utilized in underwater propulsion. Although thalattosaurs bore a superficial resemblance to lizards, their exact relationships are unresolved. They are widely accepted as diapsids, but experts have variously placed them on the reptile family tree among Lepidosauromorpha, Archosauromorpha, ichthyosaurs, and/or other marine reptiles.
Omphalosaurus is an extinct genus of marine reptile from the Early Triassic to Middle Triassic, thought to be in the order of Ichthyosauria. Most of what is known about Omphalosaurus is based on multiple jaw fragments, ribs, and vertebrae. Specimens of Omphalosaurus have been described from the western United States, Poland, Austria and the island of Spitsbergen off the northern coast of Norway.
Callawayia is an extinct genus of ichthyosaur. It contains the species Callawayia neoscapularis.
Guizhouichthyosaurus is an extinct genus of ichthyosaur which is known primarily from the Xiaowa Formation of the lower Carnian stage of the Late Triassic in southwest China. The type species of this genus is Guizhouichthyosaurus tangae, of which multiple skeletons are known. It has been reassigned as a species of the genus Shastasaurus in the past, though it has since been considered distinct. The ichthyosaurs Cymbospondylus asiaticus, named in 2002, and Panjiangsaurus epicharis, named in 2003, are junior synonyms of G. tangae. The genus is also known from the Ladinian-aged Middle Triassic Zhuganpo Formation; additionally, the species "Callawayia" wollongangense may belong to Guizhouichthyosaurus.
Qianichthyosaurus is an extinct genus of ichthyosaur from the Ladinian and Carnian stages of the Late Triassic epoch. Its fossils have been found in southeastern China, in Carnian rocks of the Falang Formation near Huangtutang, Guizhou. The type species is Qianichthyosaurus zhoui, named by Chun Li in 1999. A second species, Qianichthyosaurus xingyiensis, was named from older (Ladinian) deposits in the Falang Formation in 2013 by Pengfei Yang and colleagues. Complete Qianichthyosaurus fossils are common in the Xiaowa Formation, with both juveniles and pregnant specimens being known; its larger contemporaries, Guizhouichthyosaurus and Guanlingsaurus, are rarer.
Suevoleviathan is an extinct genus of primitive ichthyosaur found in the Early Jurassic (Toarcian) of Holzmaden, Germany.
Toretocnemus is an extinct genus of ichthyosaur. Its remains have been found in California, United States, in Triassic layers of the Carnian Hosselkus Limestone.
Sikannisuchus is an extinct genus of large archosaur from upper Triassic deposits of northeastern British Columbia, Canada. It is known from the holotype, TMP 94.382.3, a posterior portion of skull roof and from other fragmentary remains. It was found from four localities of the Pardonet Formation, near the community of Sikanni Chief. It was first named by Elizabeth L. Nicholls, Donald B. Brinkman, and Xiao-Chun Wu in 1998 and the type species is Sikannisuchus huskyi. It would have reached about 4 metres (13 ft) in length. Ichthyosaurs such as Macgowania, Callawayia and possibly the giant shastasaurid Shonisaurus, coelacanths Whiteia banffensis and possibly Garnbergia, and various genera of molluscs including ammonites and bivalves have also been found in the Pardonet Formation.
Guanlingsaurus is an extinct genus of shastasaurid ichthyosaur from the Late Triassic of China. It grew up to 8.3 metres (27 ft) in length and has a wide, triangular skull with a short and toothless snout.
Merriamosauria is an extinct clade of ichthyosaurs. It was named by Ryosuke Motani in his 1999 analysis of the relationships of ichthyopterygian marine reptiles and was defined in phylogenetic terms as a stem-based taxon including "the last common ancestor of Shastasaurus pacificus and Ichthyosaurus communis, and all of its descendants." The name honours John Campbell Merriam. Based on this definition, Merriamosauria includes most ichthyosaurs except for several Triassic groups such as the clade Mixosauria, the family Cymbospondylidae, and perhaps the family Toretocnemidae. Merriamosaurs are characterized by features in their pectoral girdles and limb bones, including an extensive connection between the scapula and the coracoid bone, the absence of the first metacarpal and the absence of a pisiform bone.
Elizabeth (Betsy) Laura Nicholls was an American-Canadian paleontologist who specialized in Triassic marine reptiles. She was a paleontologist at the Royal Tyrrell Museum in Alberta, Canada.
This timeline of ichthyosaur research is a chronological listing of events in the history of paleontology focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, sharks, or swordfish. Scientists have documented ichthyosaur fossils at least as far back as the late 17th century. At that time, a scholar named Edward Lhuyd published a book on British fossils that misattributed some ichthyosaur vertebrae to actual fishes; their true nature was not recognized until the 19th century. In 1811, a boy named Joseph Anning discovered the first ichthyosaur fossils that would come to be scientifically recognized as such. His sister, Mary Anning, would later find the rest of its skeleton and would go on to become a respected fossil collector and paleontologist in her own right.
Ichthyotitan is an extinct genus of giant ichthyosaur from the Late Triassic (Rhaetian), known from the Westbury Mudstone Formation in Somerset, England. It is believed to be a shastasaurid, extending the family's range by 13 million years up to the latest Triassic. The discovery of Ichthyotitan has been considered evidence that shastasaurids were still thriving until their disappearance in the Triassic–Jurassic extinction event.