Shastasaurus

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Shastasaurus
Temporal range: Late Triassic, 235.0–205.6  Ma [1]
Shastasaurus pacificus.jpg
Partial skull of Shastasaurus pacificus (UCMP 9017)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Ichthyosauria
Suborder: Longipinnati
Node: Merriamosauria
Family: Shastasauridae
Merriam, 1895
Genus: Shastasaurus
Merriam, 1895
Type species
Shastasaurus pacificus
Merriam, 1895
Species

S. pacificus
Merriam, 1895
S.? liangae
(Yin et al., 2000 [originally Guanlingsaurus ])
S.? sikanniensis
(Nicholls & Manabe, 2004 [originally Shonisaurus ])

Synonyms
  •  ?GuanlingsaurusYin et al., 2000

Shastasaurus ("Mount Shasta lizard") is an extinct genus of ichthyosaur from the Late Triassic. [2] Specimens have been found in the United States, Canada, and China. [3]

Description

Size of Shonisaurus popularis (green) and S. sikanniensis (red), a possible species of Shastasaurus, compared with a human Shonisaurus scale mmartyniuk.png
Size of Shonisaurus popularis (green) and S. sikanniensis (red), a possible species of Shastasaurus, compared with a human

Shastasaurus lived during the late Triassic period. The type species, S. pacificus, is known from California, with the name Shastasaurus directly referencing Mount Shasta in Siskiyou County, Northern California. S. pacificus was a medium-sized ichthyosaur, measuring over 7 metres (23 ft) in length. [4] A second possible species of Shastasaurus, S. sikanniensis, is known from the Pardonet Formation British Columbia, dating to the middle Norian age (about 210 million years ago). [5] By comparison, S. sikanniensis was one of the largest known ichthyosaurs, similar in size to modern-day cetaceans, measuring up to 21 metres (69 ft) in length and weighing 81.5 metric tons (89.8 short tons). [6]

Shastasaurus was highly specialized, and differed considerably from other ichthyosaurs. It was very slender in profile. S. sikanniensis had a ribcage slightly less than 2 metres (6.6 ft) deep despite a distance of over 7 metres (23 ft) between its flippers. [5] With its unusually short, toothless snout (compared to the longer, toothed, dolphin-like snouts of most ichthyosaurs), it had been proposed that Shastasaurus was a suction feeder (or filled a similar ecological niche as extant beaked whales), preying primarily on soft-bodied cephalopods. [7] However, current research indicates that the jaws of shastasaurid ichthyosaurs do not fit the suction-feeding profile, since their short and narrow hyoid bones are unsuitable to withstand impact forces for such kind of feeding, [8] and since some species like Shonisaurus had robust sectorial teeth with gut contents of mollusk shells and vertebrates. [9] [10]

It is unknown whether Shastasaurus had a dorsal fin; however, the smaller, more basal ichthyosaur Mixosaurus had one. [11] The upper fluke of the tail was probably much less-developed than the shark-like tails found in later species. [12]

Species and synonyms

Restoration of S. pacificus Shastasaurus pacificus restoration.jpg
Restoration of S. pacificus

The type species of Shastasaurus is S. pacificus, from the late Carnian of northern California. It is known only from fragmentary remains, which have led to the assumption that it was a 'normal' ichthyosaur in terms of proportions, especially skull proportions. Several species of long-snouted ichthyosaur were referred to Shastasaurus based on this misinterpretation, but are now placed in other genera (including Callawayia and Guizhouichthyosaurus ). [7]

S. sikanniensis specimen, Royal Tyrrell Museum Shonisaurus sikanniensis Royal Tyrrell.jpg
S. sikanniensis specimen, Royal Tyrrell Museum

Shastasaurus may include a second species, Shastasaurus liangae. It is known from several good specimens, and was originally placed in the separate genus Guanlingsaurus . Complete skulls show that it had an unusual short and toothless snout. S. pacificus probably also had a short snout, although its skull is incompletely known. [7] However, a new juvenile specimen discovered in 2013 shows that the hyoid bone of Guanlingsaurus is much shorter, and considered it as a distinct genus based on phylogenetic analysis. [13]

S. sikanniensis was originally described in 2004 as a large species of Shonisaurus . However, this classification was not based on any phylogenetic analysis, and the authors also noted similarities with Shastasaurus. The first study testing its relationships, in 2011, supported the hypothesis that it was indeed more closely related to Shastasaurus than to Shonisaurus, and it was reclassified as Shastasaurus sikanniensis. [7] However, a 2013 analysis supported the original classification, finding it more closely related to Shonisaurus than to Shastasaurus. [13] In the 2019 study, S. sikanniensis was pertained within the genus Shastasaurus. [14] In the 2021 analysis, S. sikanniensis forms a clade with Shonisaurus, indicating that it is closer to Shonisaurus than to Shastasaurus. [15] Specimens belonging to S. sikanniensis have been found in the Pardonet Formation British Columbia, dating to the middle Norian age (about 210 million years ago). [5]

In 2009, Shang & Li reclassified the species Guizhouichthyosaurus tangae as Shastasaurus tangae. However, later analysis showed that Guizhouichthyosaurus was in fact closer to more advanced ichthyosaurs, and so cannot be considered a species of Shastasaurus. [7]

Dubious species that were referred to this genus include S. carinthiacus (Huene, 1925) from the Austrian Alps and S. neubigi (Sander, 1997) from the German Muschelkalk. [3] S. neubigi, however, was re-described and reassigned to its own genus, Phantomosaurus . [16]

Synonyms of S. / G. liangae:
Guanlingichthyosaurus [17] liangaeWang et al., 2008 ( lapsus calami )

Synonyms of S. pacificus:
Shastasaurus alexandraeMerriam, 1902
Shastasaurus osmontiMerriam, 1902

See also

Related Research Articles

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Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides.

<i>Shonisaurus</i> Genus of reptiles

Shonisaurus is a genus of very large ichthyosaurs. At least 37 incomplete fossil specimens of the marine reptile have been found in the Luning Formation of Nevada, USA. This formation dates to the late Carnian age of the late Triassic period, about 237–227 million years ago.

<i>Cymbospondylus</i> Extinct genus of reptiles

Cymbospondylus is an extinct genus of large ichthyosaurs, of which it is among the oldest representatives, that lived during the Anisian stage of the Middle Triassic in what are now North America and Europe. The first known fossils of this taxon are a set of more or less complete vertebrae which were discovered in the 19th century in various mountain ranges of Nevada, in the United States, before being named and described by Joseph Leidy in 1868. It is in the beginning of the 20th century that more complete fossils were discovered through several expeditions launched by the University of California, and described in more detail by John Campbell Merriam in 1908, thus visualizing the overall anatomy of the animal. While many species have been assigned to the genus, only five are recognized as valid, the others being considered synonymous, doubtful or belonging to other genera. Cymbospondylus was formerly classified as a representative of the Shastasauridae, but more recent studies consider it to be more basal, view as the type genus of the Cymbospondylidae.

<i>Mixosaurus</i> Extinct genus of reptiles

Mixosaurus is an extinct genus of Middle Triassic ichthyosaur. Its fossils have been found near the Italy–Switzerland border and in South China.

<span class="mw-page-title-main">Shastasauridae</span> Extinct family of reptiles

Shastasauridae is an extinct family of Triassic ichthyosaurs. The family contains the largest known species of ichthyosaurs, which include some of and possibly the largest known marine reptiles.

<i>Contectopalatus</i> Extinct species of reptile

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Guizhouichthyosaurus is an extinct genus of ichthyosaur which is known primarily from the Xiaowa Formation of the lower Carnian stage of the Late Triassic in southwest China. The type species of this genus is Guizhouichthyosaurus tangae, of which multiple skeletons are known. It has been reassigned as a species of the genus Shastasaurus in the past, though it has since been considered distinct. The ichthyosaurs Cymbospondylus asiaticus, named in 2002, and Panjiangsaurus epicharis, named in 2003, are junior synonyms of G. tangae. The genus is also known from the Ladinian-aged Middle Triassic Zhuganpo Formation; additionally, the species "Callawayia" wollongangense may belong to Guizhouichthyosaurus.

<i>Phantomosaurus</i> Extinct genus of reptiles

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Mixosauridae was an early group of ichthyosaurs, living between 247.2 and 235 million years ago, during the Triassic period. Fossils of mixosaurs have been found all over the world: China, Timor, Indonesia, Italy, Germany, Spitsbergen, Switzerland, Svalbard, Canada, Alaska, and Nevada.

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<span class="mw-page-title-main">Merriamosauria</span> Extinct clade of reptiles

Merriamosauria is an extinct clade of ichthyosaurs. It was named by Ryosuke Motani in his 1999 analysis of the relationships of ichthyopterygian marine reptiles and was defined in phylogenetic terms as a stem-based taxon including "the last common ancestor of Shastasaurus pacificus and Ichthyosaurus communis, and all of its descendants." The name honours John Campbell Merriam. Based on this definition, Merriamosauria includes most ichthyosaurs except for several Triassic groups such as the clade Mixosauria, the family Cymbospondylidae, and perhaps the family Toretocnemidae. Merriamosaurs are characterized by features in their pectoral girdles and limb bones, including an extensive connection between the scapula and the coracoid bone, the absence of the first metacarpal and the absence of a pisiform bone.

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<span class="mw-page-title-main">Timeline of ichthyosaur research</span>

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<i>Ichthyotitan</i> Genus of giant ichthyosaurs

Ichthyotitan is an extinct genus of giant ichthyosaur from the Late Triassic (Rhaetian), known from the Westbury Mudstone Formation in Somerset, United Kingdom. It is believed to be a shastasaurid, extending the family's range by 13 million years up to the latest Triassic. The discovery of Ichthyotitan has been considered evidence that shastasaurids were still thriving until their disappearance in the Triassic–Jurassic extinction event.

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