Ichthyosaurus Temporal range: Early Jurassic, Possible Late Triassic (Rhaetian) record, but yet to be confirmed [1] | |
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Fossil specimen of I. somersetensis at Natural History Museum, London | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | † Ichthyosauria |
Family: | † Ichthyosauridae |
Genus: | † Ichthyosaurus De la Beche & Conybeare, 1821 |
Type species | |
†Ichthyosaurus communis De la Beche & Conybeare, 1822 | |
Other species | |
Ichthyosaurus (derived from Greek ἰχθύς (ichthys) meaning 'fish' and σαῦρος (sauros) meaning 'lizard') is a genus of ichthyosaurs from the Early Jurassic (Hettangian - Pliensbachian [3] ), with possible Late Triassic record, [1] from Europe (Belgium, England, Germany, Switzerland, and Portugal [4] ). It is among the best known ichthyosaur genera, as it is the type genus of the order Ichthyosauria. [5] [6] [7]
Ichthyosaurus was the first complete fossil to be discovered in the early 19th century by Mary Anning in England; [8] the holotype of I. communis, no coll. number given, [9] was a fairly complete specimen discovered by Mary and Joseph Anning around 1814 in Lyme Regis [10] but was reported as lost by McGowan (1974) in his review of the latipinnate ichthyosaurs of England. [11] The name Ichthyosaurus was first used by Charles König in 1818, but it was not used in a formal scientific description, with the earliest described ichthyosaur being Proteosaurus by James Everard Home in 1819 for a skeleton which is now attributed to Temnodontosaurus platyodon. Henry De la Beche and William Conybeare in 1821 considered Ichthyosaurus to have taxonomic priority over Proteosaurus and named the species I. communis based on BMNH 2149 (now NHMUK PV R1158), a now partially lost specimen now assigned to Temnodontosaurus that was discovered and collected between 1811 and 1812. [12] [13] One specimen that Home had assigned to Proteosaurus was the first complete ichthyosaur skeleton known, but it was destroyed in WW2. Two casts were rediscovered in 2022, showing that the specimen belonged to Ichthyosaurus, but of uncertain species. [14] During the 19th century, almost all fossil ichthyosaurs were attributed to Ichthyosaurus, resulting in the genus having over 50 species by 1900. These species were subsequently moved to separate genera or synonymised with other species. [15]
I. anningae, described in 2015 from a fossil found in the early 1980s in Dorset, England, was named after Anning. [16] [17] [15] The fossil was acquired by Doncaster Museum and Art Gallery, where it was misidentified as a plaster cast. In 2008, Dean Lomax, from the University of Manchester, recognised it as genuine and worked with Judy Massare, of the State University of New York, to establish it as a new species. [16]
Ichthyosaurus was smaller than most of its relatives, with the largest specimen of I. somersetensis measuring up to 3–3.3 m (9.8–10.8 ft) in length. [18] In comparison, other species were much smaller, with the I. communis reaching up to 2 metres (6.6 ft) in length, I. larkini probably up to 2.5 metres (8.2 ft), I. anningae up to 1.8 metres (5.9 ft), I. breviceps up to 1.9 metres (6.2 ft), and I. conybeari up to 1.5 metres (4.9 ft). [1] [9] Many Ichthyosaurus fossils are well-preserved and fully articulated. Some fossils still had baby specimens inside them, indicating that Ichthyosaurus was viviparous. Similar finds in the related Stenopterygius also show this. [19] [20] Jurassic ichthyosaurs had a fleshy dorsal fin on their back as well as a large caudal fin. Icthyosaurus is distinguished from other ichthyosaurs by having a wide forefin with 5 or more digits with an anterior digital bifurcation, but the morphology of the humerus and coracoids are also distinct from that of other Lower Jurassic ichthyosaurs, as is the arrangement of the dermal bones, though the suture lines used to diagnose these are not always visible. [15]
This cladogram below follows the topology from a 2010 analysis by Patrick S. Druckenmiller and Erin E. Maxwell. [21]
Thunnosauria |
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Ichthyosaurus is suggested to have been a ram feeder, with the morphology of its hyobranchial apparatus suggesting that it was incapable of suction feeding, [23] using the jaws and teeth alone to capture prey. Ichthyosaurus is thought to have been a pursuit predator that was capable of sustained swift swimming via thunniform locomotion. [24] Stomach contents of Ichthyosaurus anningae indicate that it fed on cephalopods (likely belemnites) and fish. [25] Like other ichthyosaurs, it likely relied on its sense of sight, possibly in combination with olfaction. [26]
It was initially believed that Ichthyosaurus laid eggs on land, but fossil evidence shows that in fact the females gave birth to live young. As such, they were well-adapted to life as fully pelagic organisms (i.e. they never came onto land). The babies were born tail first to prevent them from drowning in the water. [19]
Joseph Victor von Scheffels poem Der Ichthyosaurus describes its extinction in humouristic verses. A monument on Hohentwiel cites it as well. [27] The poem has been translated among others by Charles Godfrey Leland [28] Some of the stanzas:
The rushes are strangely rustling,
The ocean uncannily gleams,
As with tears in his eyes down gushing,
An Ichthyosaurus swims.
He bewails the frightful corruption
Of his age, for an awful tone
Has lately been noticed by many
In the Lias formation shown.
Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides.
Temnodontosaurus is an extinct genus of ichthyosaur from the Early Jurassic period. They lived between 200 and 175 million years ago (Hettangian-Toarcian) in what is now Western Europe and possibly Chile. It lived in the deeper areas of the open ocean. University of Bristol paleontologist Jeremy Martin described the genus Temnodontosaurus as "one of the most ecologically disparate genera of ichthyosaurs," although the number of valid Temnodontosaurus species has varied over the years.
Stenopterygius is an extinct genus of thunnosaur ichthyosaur known from Europe.
Eurhinosaurus is an extinct genus of ichthyosaur from the Early Jurassic (Toarcian), ranging between 183 and 175 million years. Fossils of the aquatic reptile have been found in Western Europe. They used to live in the deep, open sea area. Eurhinosaurus was a large genus of ichthyosaurs. An adult individual could reach up to 7 metres (23 ft) in length.
Shastasauridae is an extinct family of Triassic ichthyosaurs. The family contains the largest known species of ichthyosaurs, which include some of and possibly the largest known marine reptiles.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1857.
Brachypterygius is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the Late Jurassic of England. The type species was originally described and named as Ichthyosaurus extremus by Boulenger in 1904. Brachypterygius was named by Huene in 1922 for the width and shortness of the forepaddle, and the type species is therefore Brachypterygius extremus. The holotype of B. extremus was originally thought to be from the Lias Group of Bath, United Kingdom, but other specimens suggest it more likely came from the Kimmeridgian Kimmeridge Clay of Kimmeridge Bay, Dorset, UK.
Nannopterygius is an extinct genus of ophthalmosaurid ichthyosaur that lived during the Middle Jurassic to the Early Cretaceous. Fossils are known from England, Kazakhstan, Russia, and Norway and six species are currently assigned to the genus.
Chacaicosaurus is a genus of neoichthyosaurian ichthyosaur known from the Middle Jurassic of Argentina. The single known specimen of this genus was excavated from the Los Molles Formation in Neuquén Province, and is housed at the Museo Olsacher under the specimen number MOZ 5803. This specimen consists of a skull, forelimb, some vertebrae, and some additional postcranial elements. The genus was named by Marta Fernández in 1994, and contains a single species, Chacaicosaurus cayi, making it the first named distinctive ichthyosaur from the Bajocian stage. It is a medium-sized ichthyosaur with a very long snout, which bears a ridge running along each side. The forelimbs of Chacaicosaurus are small and contain four main digits.
Protoichthyosaurus is a genus of ichthyosaur from the early Jurassic of southern England. Two species are known, P. prostaxalis—the type species, named by Appleby in 1979—and P. applebyi. A third species, P. prosostealis, was named by Appleby, but it was removed from the genus in 2017 due to its similarity to Ichthyosaurus. The genus Protoichthyosaurus was synonymized with Ichthyosaurus by Maisch and Hungerbuhler in 1997, and again by Maisch and Matzke in 2000. However, it was found to be distinct in 2017 by Dean Lomax and colleagues, who separated it from Ichthyosaurus on account of differences in the arrangement and shape of the carpal ossifications, as well as the absence of the fifth digit. The species most likely lived during the Hettangian stage, but may have lived as early as the Rhaetian and as late as the Sinemurian.
Baptanodon is an ichthyosaur of the Late Jurassic period, named for its supposed lack of teeth. It had a graceful 3.5 m (11 ft) long dolphin-shaped body, and its jaws were well adapted for catching squid. Major fossil finds of this genus have been recorded in North America. The type species, Sauranodon natans, was originally included under Sauranodon in 1879, but this name was preoccupied.
The Blue Lias is a geological formation in southern, eastern and western England and parts of South Wales, part of the Lias Group. The Blue Lias consists of a sequence of limestone and shale layers, laid down in latest Triassic and early Jurassic times, between 195 and 200 million years ago. The Blue Lias is famous for its fossils, especially ammonites.
Suevoleviathan is an extinct genus of primitive ichthyosaur found in the Early Jurassic (Toarcian) of Holzmaden, Germany.
Acamptonectes is a genus of ophthalmosaurid ichthyosaurs, a type of dolphin-like marine reptiles, that lived during the Early Cretaceous around 130 million years ago. The first specimen, a partial adult skeleton, was discovered in Speeton, England, in 1958, but was not formally described until 2012 by Valentin Fischer and colleagues. They also recognised a partial subadult skeleton belonging to the genus from Cremlingen, Germany, and specimens from other localities in England. The genus contains the single species Acamptonectes densus; the generic name means "rigid swimmer" and the specific name means "compact" or "tightly packed".
The Charmouth Mudstone Formation is a geological formation in England, dating to the Early Jurassic (Sinemurian–Pliensbachian). It forms part of the lower Lias Group. It is most prominently exposed at its type locality in cliff section between Lyme Regis and Charmouth but onshore it extends northwards to Market Weighton, Yorkshire, and in the subsurface of the East Midlands Shelf and Wessex Basin. The formation is notable for its fossils, including those of ammonites and marine reptiles and rare dinosaur remains. The formation played a prominent role in the history of early paleontology, with its Lyme Regis-Charmouth exposure being frequented by fossil collectors including Mary Anning.
This timeline of ichthyosaur research is a chronological listing of events in the history of paleontology focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, sharks, or swordfish. Scientists have documented ichthyosaur fossils at least as far back as the late 17th century. At that time, a scholar named Edward Lhuyd published a book on British fossils that misattributed some ichthyosaur vertebrae to actual fishes; their true nature was not recognized until the 19th century. In 1811, a boy named Joseph Anning discovered the first ichthyosaur fossils that would come to be scientifically recognized as such. His sister Mary would later find the rest of its skeleton and would go on to become a respected fossil collector and paleontologist in her own right.
Temnodontosaurus eurycephalus is an extinct species of marine reptile. It is thought to have been a nektonic carnivore. Its name comes from the Greek ευρύς, for "wide" and κεφαλή for "head". It possibly belongs to a new genus.
Wahlisaurus is an extinct genus of leptonectid ichthyosaur. The holotype was found in the Scunthorpe Mudstone in Nottinghamshire in 1951 and described in 2016 by Dean Lomax. The type species is Wahlisaurus massare, and two specimens have been found: the first consisting of a skull and an incomplete skeleton, and the second a single coracoid.
Ichthyotitan is an extinct genus of giant ichthyosaur from the Late Triassic (Rhaetian), known from the Westbury Mudstone Formation in Somerset, England. It is believed to be a shastasaurid, extending the family's range by 13 million years up to the latest Triassic. The discovery of Ichthyotitan has been considered evidence that shastasaurids were still thriving until their disappearance in the Triassic–Jurassic extinction event.