| Mollesaurus Temporal range: Middle Jurassic, | |
|---|---|
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Order: | † Ichthyosauria |
| Family: | † Ophthalmosauridae |
| Subfamily: | † Ophthalmosaurinae |
| Genus: | † Mollesaurus Fernández, 1999 |
| Species: | †M. periallus |
| Binomial name | |
| †Mollesaurus periallus Fernández, 1999 | |
| Synonyms | |
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Mollesaurus is an extinct genus of large ophthalmosaurine ophthalmosaurid ichthyosaur known from northwestern Patagonia of Argentina. [1]
Mollesaurus was named by Marta S. Fernández in 1999 and the type species is Mollesaurus periallus. The generic name is derived from the name of the Los Molles Formation, where the holotype was collected, and sauros, Greek for "lizard". The specific name is derived from periallos, Greek for "before all others", in reference to the fact that it is the oldest ophthalmosaurid and one of the oldest thunnosaurs. [1]
Mollesaurus is known from the holotype MOZ 2282 V, articulated partial skeleton which preserved partial skull and most of the vertebral column. It was collected in the Chacaico Sur locality from the Emileia giebeli ammonoid zone of the Los Molles Formation, Cuyo Group, dating to the early Bajocian stage of the Middle Jurassic, about 171.6-170 million years ago. Mollesaurus, along with Chacaicosaurus cayi which was found at the same locality, are the only diagnostic ichthyosaur specimens from the Aalenian-Bathonian interval. It was found near Zapala city of the Neuquén Basin. [1]
Maisch and Matzke (2000) regarded Mollesaurus to be a species of Ophthalmosaurus . [2] However, all recent cladistic analyses found that Mollesaurus is a valid genus of ophthalmosaurid. [3] [4] Patrick S. Druckenmiller and Erin E. Maxwell (2010) found it to be the basalmost member of the ophthalmosaurid lineage that included Brachypterygius , Caypullisaurus and Platypterygius (but not Ophthalmosaurus). [3] Valentin Fischer, Michael W. Maisch, Darren Naish, Ralf Kosma, Jeff Liston, Ulrich Joger, Fritz J. Krüger, Judith Pardo Pérez, Jessica Tainsh and Robert M. Appleby (2012) found it to be the basalmost member of Ophthalmosaurinae. [5]
The following cladogram shows a possible phylogenetic position of Mollesaurus in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020). [6]
| Ophthalmosauria |
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Ophthalmosaurus is an ichthyosaur of the Jurassic period. Possible remains from the Cretaceous, around 145 million years ago, are also known. It was a relatively medium-sized ichthyosaur, measuring 4 m (13 ft) long and weighing 930–950 kg (2,050–2,090 lb). Named for its extremely large eyes, it had a jaw containing many small but robust teeth. Major fossil finds of this genus have been recorded in Europe with a second species possibly being found in North America.
Aegirosaurus is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from the late Jurassic and early Cretaceous of Europe. It was originally named as a species of Ichthyosaurus.
Caypullisaurus is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur from the Late Jurassic to the Early Cretaceous of Argentina. Its holotype was collected from the Vaca Muerta Formation of Cerro Lotena, Neuquen, dating to the early Tithonian stage of the Late Jurassic, about 150 million years ago. Caypullisaurus was first named by Marta Fernández in 1997 and the type species is Caypullisaurus bonapartei. It was a large ichthyosaur, measuring about 7 m (23 ft) long.
Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous worldwide. Almost all ichthyosaurs from the Middle Jurassic onwards belong to the family, until the extinction of ichthyosaurs in the early Late Cretaceous. Opthalmosaurids appeared worldwide during early Bajocian, subsequent to the disappearance of most other ichthyosaur lineages after the end of the Toarcian. Currently, the oldest known ophthalmosaurids is Mollesaurus from the early Bajocian of Argentina, as well as indeterminate remains of the same age from Luxembourg and Canada. Named by George H. Baur, in 1887, the family contains the basal taxa like Ophthalmosaurus. Appleby (1956) named the taxon Ophthalmosauria which was followed by some authors, but these two names are often treated as synonyms; Ophthalmosauridae has the priority over Ophthalmosauria. However, some researchers argue that Ophthalmosauridae should be restricted to the group typically referred to as Ophthalmosaurinae, with classic Platypterygiinae instead being referred to as Undorosauridae or Brachypterygiidae and Ophthalmosauria being used to unite these two groups.
Undorosaurus is an extinct genus of ophthalmosaurid ichthyosaur known from western Russia, Svalbard, and Poland. It was a large ichthyosaur, with the type species measuring 4–6 metres (13–20 ft) long and weighing 300 kilograms (660 lb).
Chacaicosaurus is a genus of neoichthyosaurian ichthyosaur known from the Middle Jurassic of Argentina. The single known specimen of this genus was excavated from the Los Molles Formation in Neuquén Province, and is housed at the Museo Olsacher under the specimen number MOZ 5803. This specimen consists of a skull, forelimb, some vertebrae, and some additional postcranial elements. The genus was named by Marta Fernández in 1994, and contains a single species, Chacaicosaurus cayi, making it the first named distinctive ichthyosaur from the Bajocian stage. It is a medium-sized ichthyosaur with a very long snout, which bears a ridge running along each side. The forelimbs of Chacaicosaurus are small and contain four main digits.
Baptanodon is an ichthyosaur of the Late Jurassic period, named for its supposed lack of teeth. It had a graceful 3.5 m (11 ft) long dolphin-shaped body, and its jaws were well adapted for catching squid. Major fossil finds of this genus have been recorded in North America. The type species, Sauranodon natans, was originally included under Sauranodon in 1879, but this name was preoccupied.
Maiaspondylus is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from Northwest Territories of Canada, the Cambridge Greensand of England and the Voronezh Region of Russia.
Simbirskiasaurus is an extinct genus of ichthyosaur from the Early Cretaceous of Ulyanovsk Oblast, Russia. Its type specimen is YKM 65119, a fragmentary skull and vertebral column.
The Speeton Clay Formation (SpC) is a Lower Cretaceous geological formation in Yorkshire, northern England. Unlike the contemporaneous terrestrial Wealden Group to the south, the Speeton Clay was deposited in marine conditions. The most common fossils in the unit are belemnites, followed by ammonites and the lobster Meyeria ornata. Dinosaur remains are among the fossils that have been recovered from the formation, although none have yet been referred to a specific genus.
Arthropterygius is a widespread genus of ophthalmosaurid ichthyosaur which existed in Canada, Norway, Russia, and Argentina from the late Jurassic period and possibly to the earliest Cretaceous.
Athabascasaurus is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from Alberta, Canada.
Sveltonectes is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from Ul’yanovsk region, western Russia.
Acamptonectes is a genus of ophthalmosaurid ichthyosaurs, a type of dolphin-like marine reptiles, that lived during the Early Cretaceous around 130 million years ago. The first specimen, a partial adult skeleton, was discovered in Speeton, England, in 1958, but was not formally described until 2012 by Valentin Fischer and colleagues. They also recognised a partial subadult skeleton belonging to the genus from Cremlingen, Germany, and specimens from other localities in England. The genus contains the single species Acamptonectes densus; the generic name means "rigid swimmer" and the specific name means "compact" or "tightly packed".
Platypterygiinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the early Late Jurassic to the early Late Cretaceous of Asia, Australia, Europe, North America and South America. Currently, the oldest known platypterygiine is Brachypterygius. Platypterygiines were characterized by square tooth roots in cross-section, an extremely reduced extracondylar area of the basioccipital, prominent dorsal and ventral trochanters on humerus and ischiopubis lacking an obturator foramen.
Ophthalmosaurinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the Middle Jurassic to the late Early Cretaceous of Europe, North America and South America. Currently, the oldest and the basalmost known ophthalmosaurine is Mollesaurus from the early Bajocian of Argentina. Ophthalmosaurines were characterized by a large extracondylar area of the basioccipital in the form of a thick and concave peripheral band, posterodistally deflected ulnar facet of the humerus, large ulna with concave and edgy posterior surface and ischiopubis with obturator foramen.
Sisteronia is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the 'middle' Cretaceous of southeastern England and southeastern France. It contains a single species, Sisteronia seeleyi.
This timeline of ichthyosaur research is a chronological listing of events in the history of paleontology focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, sharks, or swordfish. Scientists have documented ichthyosaur fossils at least as far back as the late 17th century. At that time, a scholar named Edward Lhwyd published a book on British fossils that misattributed some ichthyosaur vertebrae to actual fishes; their true nature was not recognized until the 19th century. In 1811, a boy named Joseph Anning discovered the first ichthyosaur fossils that would come to be scientifically recognized as such. His sister Mary would later find the rest of its skeleton and would go on to become a respected fossil collector and paleontologist in her own right.
Keilhauia is a genus of ophthalmosaurid ichthyosaur, a type of dolphin-like, large-eyed marine reptile, from the Early Cretaceous shallow marine Slottsmøya Member of the Agardhfjellet Formation in Svalbard, Norway. The genus contains a single species, K. nui, known from a single specimen discovered in 2010 and described by Delsett et al. in 2017. In life, Keilhauia probably measured approximately 4 metres (13 ft) in length; it can be distinguished by other ophthalmosaurids by the wide top end of its ilium and the relatively short ischiopubis compared to the femur. Although it was placed in a basal position within the Ophthalmosauridae by phylogenetic analysis, this placement is probably incorrect.
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