Nerve tract

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White matter tracts within a human brain, as visualized by MRI tractography White Matter Connections Obtained with MRI Tractography.png
White matter tracts within a human brain, as visualized by MRI tractography

A nerve tract is a bundle of nerve fibers (axons) connecting nuclei of the central nervous system. [1] [2] [3] In the peripheral nervous system, this is known as a nerve fascicle, and has associated connective tissue. The main nerve tracts in the central nervous system are of three types: association fibers, commissural fibers, and projection fibers. A nerve tract may also be referred to as a commissure, decussation, or neural pathway. [4] A commissure connects the two cerebral hemispheres at the same levels, while a decussation connects at different levels (crosses obliquely).

Contents

Types

Tracts of human brain.png

The nerve fibers in the central nervous system can be categorized into three groups on the basis of their course and connections. [5] Different tracts may also be referred to as projections or radiations such as thalamocortical radiations.

Association fibers

The cingulum shown in red in the cingulate gyrus. Sobo 1909 671 - Cingulum.png
The cingulum shown in red in the cingulate gyrus.

The tracts that connect cortical areas within the same hemisphere are called association tracts. [5] Long association fibers connect different lobes of a hemisphere to each other whereas short association fibers connect different gyri within a single lobe. Among their roles, association tracts link perceptual and memory centers of the brain. [6]

The cingulum is a major association tract. The cingulum forms the white matter core of the cingulate gyrus and links from this to the entorhinal cortex. Another major association tract is the superior longitudinal fasciculus (SLF) that has three parts.

Commissural fibers

Commissural tracts connect corresponding cortical areas in the two hemispheres. [5] They cross from one cerebral hemisphere to the other through bridges called commissures. The great majority of commissural tracts pass through the largest commissure the corpus callosum. A few tracts pass through the much smaller anterior and posterior commissures. Commissural tracts enable the left and right sides of the cerebrum to communicate with each other. Other commissures are the hippocampal commissure, and the habenular commissure.

Projection fibers

Projection tracts connect the cerebral cortex with the corpus striatum, diencephalon, brainstem and the spinal cord. [5] The corticospinal tract for example, carries motor signals from the cerebrum to the spinal cord. Other projection tracts carry signals upward to the cerebral cortex. Superior to the brainstem, such tracts form a broad, dense sheet called the internal capsule between the thalamus and basal nuclei, then radiate in a diverging, fanlike array to specific areas of the cortex.

See also

Related Research Articles

<span class="mw-page-title-main">Central nervous system</span> Brain and spinal cord

The central nervous system (CNS) is the part of the nervous system consisting of the brain and spinal cord, the retina and optic nerve, and the olfactory nerve and epithelia. The CNS is so named because the brain integrates the received information and coordinates and influences the activity of all parts of the bodies of bilaterally symmetric and triploblastic animals—that is, all multicellular animals except sponges and diploblasts. It is a structure composed of nervous tissue positioned along the rostral to caudal axis of the body and may have an enlarged section at the rostral end which is a brain. Only arthropods, cephalopods and vertebrates have a true brain, though precursor structures exist in onychophorans, gastropods and lancelets.

Articles related to anatomy include:

<span class="mw-page-title-main">Brainstem</span> Posterior part of the brain, adjoining and structurally continuous

The brainstem is the stalk-like part of the brain that interconnects the cerebrum and diencephalon with the spinal cord. In the human brain, the brainstem is composed of the midbrain, the pons, and the medulla oblongata. The midbrain is continuous with the thalamus of the diencephalon through the tentorial notch.

<span class="mw-page-title-main">Trigeminal nerve</span> Cranial nerve responsible for the faces senses and motor functions

In neuroanatomy, the trigeminal nerve (lit. triplet nerve), also known as the fifth cranial nerve, cranial nerve V, or simply CN V, is a cranial nerve responsible for sensation in the face and motor functions such as biting and chewing; it is the most complex of the cranial nerves. Its name (trigeminal, from Latin tri- 'three', and -geminus 'twin') derives from each of the two nerves (one on each side of the pons) having three major branches: the ophthalmic nerve (V1), the maxillary nerve (V2), and the mandibular nerve (V3). The ophthalmic and maxillary nerves are purely sensory, whereas the mandibular nerve supplies motor as well as sensory (or "cutaneous") functions. Adding to the complexity of this nerve is that autonomic nerve fibers as well as special sensory fibers (taste) are contained within it.

<span class="mw-page-title-main">Cerebral peduncle</span> Stalks that attach the cerebrum to the brainstem

The cerebral peduncles are the two stalks that attach the cerebrum to the brainstem. They are structures at the front of the midbrain which arise from the ventral pons and contain the large ascending (sensory) and descending (motor) nerve tracts that run to and from the cerebrum from the pons. Mainly, the three common areas that give rise to the cerebral peduncles are the cerebral cortex, the spinal cord and the cerebellum. The region includes the tegmentum, crus cerebri and pretectum. By this definition, the cerebral peduncles are also known as the basis pedunculi, while the large ventral bundle of efferent fibers is referred to as the cerebral crus or the pes pedunculi.

<span class="mw-page-title-main">Cerebrum</span> Large part of the brain containing the cerebral cortex

The cerebrum, telencephalon or endbrain is the largest part of the brain containing the cerebral cortex, as well as several subcortical structures, including the hippocampus, basal ganglia, and olfactory bulb. In the human brain, the cerebrum is the uppermost region of the central nervous system. The cerebrum develops prenatally from the forebrain (prosencephalon). In mammals, the dorsal telencephalon, or pallium, develops into the cerebral cortex, and the ventral telencephalon, or subpallium, becomes the basal ganglia. The cerebrum is also divided into approximately symmetric left and right cerebral hemispheres.

<span class="mw-page-title-main">Internal capsule</span> White matter structure situated in the inferomedial part of each cerebral hemisphere of the brain

The internal capsule is a white matter structure situated in the inferomedial part of each cerebral hemisphere of the brain. It carries information past the basal ganglia, separating the caudate nucleus and the thalamus from the putamen and the globus pallidus. The internal capsule contains both ascending and descending axons, going to and coming from the cerebral cortex. It also separates the caudate nucleus and the putamen in the dorsal striatum, a brain region involved in motor and reward pathways.

<span class="mw-page-title-main">Neural pathway</span> Connection formed between neurons that allows neurotransmission

In neuroanatomy, a neural pathway is the connection formed by axons that project from neurons to make synapses onto neurons in another location, to enable neurotransmission. Neurons are connected by a single axon, or by a bundle of axons known as a nerve tract, or fasciculus. Shorter neural pathways are found within grey matter in the brain, whereas longer projections, made up of myelinated axons, constitute white matter.

<span class="mw-page-title-main">Pyramidal tracts</span> Include both the corticobulbar tract and the corticospinal tract

The pyramidal tracts include both the corticobulbar tract and the corticospinal tract. These are aggregations of efferent nerve fibers from the upper motor neurons that travel from the cerebral cortex and terminate either in the brainstem (corticobulbar) or spinal cord (corticospinal) and are involved in the control of motor functions of the body.

<span class="mw-page-title-main">Spinothalamic tract</span> Sensory pathway from the skin to the thalamus

The spinothalamic tract is a part of the anterolateral system or the ventrolateral system, a sensory pathway to the thalamus. From the ventral posterolateral nucleus in the thalamus, sensory information is relayed upward to the somatosensory cortex of the postcentral gyrus.

<span class="mw-page-title-main">Dorsal column–medial lemniscus pathway</span> Sensory spinal pathway

The dorsal column–medial lemniscus pathway (DCML) is a sensory pathway of the central nervous system that conveys sensations of fine touch, vibration, two-point discrimination, and proprioception from the skin and joints. It transmits information from the body to the primary somatosensory cortex in the postcentral gyrus of the parietal lobe of the brain. The pathway receives information from sensory receptors throughout the body, and carries this in nerve tracts in the white matter of the dorsal column of the spinal cord to the medulla, where it is continued in the medial lemniscus, on to the thalamus and relayed from there through the internal capsule and transmitted to the somatosensory cortex. The name dorsal-column medial lemniscus comes from the two structures that carry the sensory information: the dorsal columns of the spinal cord, and the medial lemniscus in the brainstem.

<span class="mw-page-title-main">Corticobulbar tract</span> Motor pathway in the brain connecting the motor cortex to the medullary pyramids

In neuroanatomy, the corticobulbartract is a two-neuron white matter motor pathway connecting the motor cortex in the cerebral cortex to the medullary pyramids, which are part of the brainstem's medulla oblongata region, and are primarily involved in carrying the motor function of the non-oculomotor cranial nerves. The corticobulbar tract is one of the pyramidal tracts, the other being the corticospinal tract.

<span class="mw-page-title-main">Precentral gyrus</span> Motor gyrus of the posterior frontal lobe of the brain

The precentral gyrus is a prominent gyrus on the surface of the posterior frontal lobe of the brain. It is the site of the primary motor cortex that in humans is cytoarchitecturally defined as Brodmann area 4.

The projection fibers consist of efferent and afferent fibers uniting the cortex with the lower parts of the brain and with the spinal cord. In human neuroanatomy, bundles of axons called tracts, within the brain, can be categorized by their function into association fibers, projection fibers, and commissural fibers.

<span class="mw-page-title-main">Commissural fiber</span> Axons that connect the two hemispheres of the brain

The commissural fibers or transverse fibers are axons that connect the two hemispheres of the brain. In contrast to commissural fibers, association fibers connect regions within the same hemisphere of the brain, and projection fibers connect each region to other parts of the brain or to the spinal cord.

A commissure is the location at which two objects abut or are joined. The term is used especially in the fields of anatomy and biology.

<span class="mw-page-title-main">Association fiber</span> Axons that connect cortical areas within the same cerebral hemisphere

Association fibers are axons that connect cortical areas within the same cerebral hemisphere.

<span class="mw-page-title-main">Spinal cord</span> Long, tubular central nervous system structure in the vertebral column

The spinal cord is a long, thin, tubular structure made up of nervous tissue that extends from the medulla oblongata in the brainstem to the lumbar region of the vertebral column (backbone) of vertebrate animals. The center of the spinal cord is hollow and contains a structure called central canal, which contains cerebrospinal fluid. The spinal cord is also covered by meninges and enclosed by the neural arches. Together, the brain and spinal cord make up the central nervous system (CNS).

<span class="mw-page-title-main">Anatomical terms of neuroanatomy</span> Terminology used to describe the central and peripheral nervous systems

This article describes anatomical terminology that is used to describe the central and peripheral nervous systems - including the brain, brainstem, spinal cord, and nerves.

References

  1. Moore, Keith; Dalley, Arthur (2005). Clinically Oriented Anatomy (5th ed.). LWW. pp.  47. ISBN   0-7817-3639-0. A bundle of nerve fibers (axons) connecting neighboring or distant nuclei of the CNS is a tract.
  2. Blumenfeld, Hal (2010). Neuroanatomy through clinical cases (2nd ed.). Sunderland, Mass.: Sinauer Associates. p. 22. ISBN   9780878936137.
  3. "nerve tract". TheFreeDictionary.com.
  4. Solomon. Biology. Cengage. p. 863.
  5. 1 2 3 4 Standring, Susan (2005). Gray's Anatomy: The Anatomical Basis of Clinical Practice (39th ed.). Churchill Livingstone. pp.  411. ISBN   9780443071683. The nerve fibres which make up the white matter of the cerebral hemispheres are categorized on the basis of their course and connections. They are association fibres, which link different cortical areas in the same hemisphere; commissural fibres, which link corresponding cortical areas in the two hemispheres; or projection fibres, which connect the cerebral cortex with the corpus striatum, diencephalon, brain stem and the spinal cord
  6. Saladin, Kenneth (2012). Anatomy & Physiology: The Unity of Form and Function. New York: McGraw Hill. p. 531. ISBN   978-0-07-337825-1.