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Pseudotsuga macrocarpa | |
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Bigcone Douglas-fir at Mount San Antonio | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Gymnospermae |
Division: | Pinophyta |
Class: | Pinopsida |
Order: | Pinales |
Family: | Pinaceae |
Genus: | Pseudotsuga |
Species: | P. macrocarpa |
Binomial name | |
Pseudotsuga macrocarpa | |
Natural range | |
Synonyms | |
Tsuga macrocarpa(Vasey) Lemmon Contents |
Pseudotsuga macrocarpa, commonly called the bigcone spruce or bigcone Douglas-fir, is an evergreen conifer native to the mountains of southern California. It is notable for having the largest (by far) cones in the genus Pseudotsuga , hence the name.
The tree occurs from the San Rafael Mountains in central Santa Barbara County and the Tehachapi Mountains of southwestern Kern County, south through the Transverse Ranges, to the Cuyamaca Mountains in San Diego County. The tree is shade-tolerant and prefers to grow on slopes.
Pseudotsuga macrocarpa is a Douglas-fir. The name "bigcone spruce", though confusing as it is not a spruce species, is often still used, [3] [4] and occurs in place names. [5]
Pseudotsuga macrocarpa typically grows from 15–30 m (49–98 ft) in height and 56–155 cm (1 ft 10 in – 5 ft 1 in) in trunk diameter. [6] The growth form is straight, with a conical crown from 12–30 m (39–98 ft) broad, and a strong and spreading root system. The bark is deeply ridged, composed of thin, woodlike plates separating heavy layers of cork; bark of trees over 1 m (3.3 ft) in diameter is from 15–20 cm (5.9–7.9 in) thick. The main branches are long and spreading with pendulous side shoots.
The leaves are needle-like, 2.5–4.5 cm (0.98–1.77 in) long, [7] and are shed when about five years old. The female cones are from 11–17 cm (4.3–6.7 in) long, [4] larger and with thicker scales than those of other douglas-firs, and with exserted tridentine bracts. The seeds are large and heavy, 10 mm long and 8 mm broad, with a short rounded wing 13 mm long; [4] they may be bird or mammal dispersed as the wing is too small to be effective for wind dispersal. Trees start producing seeds at about 20 years of age.
The largest known individual of this species is 53 m (174 ft) tall, 231 cm (91 in) in diameter, and is estimated to be from 600 to 700 years of age.
Pseudotsuga macrocarpa is restricted to the California montane chaparral and woodlands and California coastal sage and chaparral ecoregions of California. It prefers a Mediterranean climate, characterized by hot dry summers and wet, mild winters. Annual rainfall during a 30-year period on a bigcone Douglas-fir site in the San Gabriel Mountains averaged 75 cm (30 in) and ranged from 25–125 cm (9.8–49.2 in).
Bigcone Douglas-fir occurs between 300–2,700 m (980–8,860 ft). At low elevation, it occurs near streams in moist, shaded canyons and draws where aspects are mostly north and east. At elevations from 1,350–1,700 m (4,430–5,580 ft), aspects include south- and east-facing slopes. At these elevations, it also grows on sloping hillsides, ridges, and benches. At higher elevations, it occurs on south and west aspects on all types of terrain. The average angle of slope on which it grows is 35 degrees, ranging from level to 90 degrees, although these extremes are uncommon.
Ranges the tree is found in, south to north, include: [8] [9]
Pseudotsuga macrocarpa, has several features to tolerate and survive wildfire, notably the very thick bark, and the presence of numerous adventitious buds on the upper side of the branches; this enables the trees to survive even crown fires which burn off all the branchlets, the apparently dead trees becoming green again the following spring. Wildfire frequencies in the chaparral habitats in which it often grows typically range from 15 to 50 year intervals. Bigcone Douglas-fir is closely associated with canyon live oak (Quercus chrysolepis) and often establishes itself in its shade; after about 50 years, it emerges above the oak canopy.
The number of plant associates in bigcone Douglas-fir communities is usually small.
Pseudotsuga macrocarpa populations are suspected to be declining due to possibly larger and more extreme wildland fires with greater frequencies. Major wildfires within its range, since 2003, have clearly proven a reduced extent when compared to early 1930s extents derived from historical aerial photos. Although historical information has provided the opportunity to detect stand level patch changes, post-fire resprouting of older more mature trees and natural regeneration and recruitment of the species into higher canopy has yet to be adequately quantified.
One tree species in direct competition with bigcone is Calocedrus decurrens , with preliminary post-fire regeneration of this species exponentially greater than bigcone. One or two years after the Station fire in 2009 on the Angeles National Forest, there was an estimated 20:1 cedar:bigcone seedling density in fixed radius plots on Mount Wilson. It may be more appropriate to perform population stability estimates up to five years or much later (i.e. 20 yrs) after a large conflagration due to the potential for immediate and delayed post-fire sprouting and regeneration and interplant competition, as well as the well-noted strategy of seed germination in shrub understories, which is likely to escape detection by surveyors until much later in its life.
Research related to the role of mycorrhizae and its relationship to seed establishment needs evaluated in these vegetation communities due to the suspected role it has with the relationship with water, especially in water-limited systems such as those in the wildlands of southern California. In addition, an aggressive seed cone collection strategy should be drafted for this species which includes extensive collection during large cone production years such as 2013, and should include a tracking system to determine correlations to climatic conditions in order to develop a foundation from which to perform species viability assessments w/ varying future climate scenarios.
This tree is being considered for more extensive plantings in semiarid locales. Its favorable qualities include resistance to drought, fire, insects, decay, and damage from ozone, and its aggressive rooting system and tolerance to variable growing medium. The needles of older trees sometimes fade to yellow, drop, and trees appear dead only to sprout with renewed vigor within two years. The reason is unknown, although drought or insects may be possible causes.
Bigcone Douglas-fir stands provide habitat for black-tailed deer, black bear, and various small animals. These trees provide preferred spring habitat for black bear in the San Bernardino Mountains.
The seeds are eaten by various rodents and birds.
Bigcone Douglas-fir is used for watershed and habitat restoration. The Los Angeles County Department of Forestry has extensively planted the tree over a 50-year period for that purpose.
Some hybrids of Pseudotsuga macrocarpa (Bigcone Douglas-fir) × Pseudotsuga menziesii (Coast Douglas-fir) show promise for planting on drier restoration sites within the P. menziesii−Douglas-fir natural range. These hybrids produce wood of comparable quality to that of Douglas-fir and have the drought tolerance of bigcone Douglas-fir.
There is no current commercial market for bigcone Douglas-fir wood due to limited distribution and access. It is heavy, hard, and fine grained but not durable. There is less sapwood than heartwood, and the latter contains pockets of resin.
In the past, the wood was used locally for fuel and lumber.
Pseudotsuga is a genus of evergreen coniferous trees in the family Pinaceae. Common names for species in the genus include Douglas fir, Douglas-fir, Douglas tree, Oregon pine and Bigcone spruce. Pseudotsuga menziesii is widespread in western North America and is an important source of timber. The number of species has long been debated, but two in western North America and two to four in eastern Asia are commonly acknowledged.
The Douglas fir is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three varieties: coast Douglas-fir, Rocky Mountain Douglas-fir and Mexican Douglas-fir.
Pseudotsuga menziesii var. glauca, or Rocky Mountain Douglas-fir, is an evergreen conifer native to the interior mountainous regions of western North America, from central British Columbia and southwest Alberta in Canada southward through the United States to the far north of Mexico. The range is continuous in the northern Rocky Mountains south to eastern Washington, eastern Oregon, Idaho, western and south-central Montana and western Wyoming, but becomes discontinuous further south, confined to "sky islands" on the higher mountains in Utah, Colorado, Arizona and New Mexico, with only very isolated small populations in eastern Nevada, westernmost Texas, and northern Mexico. It occurs from 600 m altitude in the north of the range, up to 3,000 m, rarely 3,200 m, in the south. Further west towards the Pacific coast, it is replaced by the related coast Douglas-fir, and to the south, it is replaced by Mexican Douglas-fir in high mountains as far south as Oaxaca. Some botanists have grouped Mexican Douglas-fir with P. menziesii var. glauca, but genetic and morphological evidence suggest that Mexican populations should be considered a different variety.
Quercus kelloggii, the California black oak or Kellogg oak, is an oak in the red oak section native to western North America. Although genetically separated from them for more than 20 million years, its leaves are remarkably similar in appearance to several other members of the red oak section including the red oak and the black oak found in eastern and central North America.
Pinus sabiniana, with vernacular names including towani pine, foothill pine, gray pine, bull pine, and digger pine, is a pine endemic to California in the United States. Some sources discourage using the name "digger pine," considering it pejorative.
Alnus rubra, the red alder, is a deciduous broadleaf tree native to western North America.
Abies bracteata, the Santa Lucia fir or bristlecone fir, is the rarest fir in North America, and according to some, the world. It is confined to steep-sided slopes and the bottoms of rocky canyons in the Santa Lucia Mountains, in the Big Sur region on the central coast of California, United States.
Abies grandis is a fir native to the Pacific Northwest and Northern California of North America, occurring at altitudes of sea level to 1,700 metres (5,600 ft). It is a major constituent of the Grand Fir/Douglas Fir Ecoregion of the Cascade Range.
Abies concolor, the white fir, concolor fir, or Colorado fir, is a coniferous tree in the pine family Pinaceae. This tree is native to the mountains of western North America, including the Cascade Range and southern Rocky Mountains, and into the isolated mountain ranges of southern Arizona, New Mexico, and Northern Mexico. It naturally occurs at elevations between 900 and 3,400 metres.
Cupressus arizonica, the Arizona cypress, is a North American species of tree in the cypress family Cupressaceae, native to the southwestern United States and Mexico. Populations may be scattered rather than in large, dense stands.
The Northern California coastal forests are a temperate coniferous forests ecoregion of coastal Northern California and southwestern Oregon.
Abies lasiocarpa, the subalpine fir or Rocky Mountain fir, is a western North American fir tree.
The Sierra Madre Occidental pine–oak forests are a Temperate broadleaf and mixed forests ecoregion of the Sierra Madre Occidental range from the southwest USA region to the western part of Mexico. They are home to a large number of endemic plants and important habitat for wildlife.
California mixed evergreen forest is a plant community found in the mountain ranges of California and southwestern Oregon.
Abies vejarii is a species of fir native to northeastern Mexico, in the states of Coahuila, Nuevo León, and Tamaulipas, where it grows at high altitudes in the Sierra Madre Oriental.
The Klamath Mountains ecoregion of Oregon and California lies inland and north of the Coast Range ecoregion, extending from the Umpqua River in the north to the Sacramento Valley in the south. It encompasses the highly dissected ridges, foothills, and valleys of the Klamath and Siskiyou Mountains. It corresponds to the Level III ecoregion designated by the Environmental Protection Agency and to the Klamath-Siskiyou forests ecoregion designated by the World Wide Fund for Nature.
The California montane chaparral and woodlands is an ecoregion defined by the World Wildlife Fund, spanning 7,900 square miles (20,000 km2) of mountains in the Transverse Ranges, Peninsular Ranges, and Coast Ranges of southern and central California. The ecoregion is part of the larger California chaparral and woodlands ecoregion, and belongs to the Mediterranean forests, woodlands, and scrub biome.
The California interior chaparral and woodlands ecoregion covers 24,900 square miles (64,000 km2) in an elliptical ring around the California Central Valley. It occurs on hills and mountains ranging from 300 feet (91 m) to 3,000 feet (910 m). It is part of the Mediterranean forests, woodlands, and scrub biome, with cool, wet winters and hot, dry summers. Temperatures within the coast can range from 53° to 65 °F and 32° to 60 °F within the mountains. Many plant and animal species in this ecoregion are adapted to periodic fire.
Cupressus stephensonii is a species of conifer known as the Cuyamaca cypress, and is endemic to Southern California. It has been classified as Hesperocyparis stephensonii. It was previously listed as Cupressus arizonica subsp. stephensonii and Cupressus arizonica var. glabra.
Pseudotsuga menziesii var. menziesii, commonly known as Coast Douglas-fir, Pacific Douglas-fir, Oregon pine, or Douglas spruce, is an evergreen conifer native to western North America from west-central British Columbia, Canada southward to central California, United States. In Oregon and Washington its range is continuous from the Cascades crest west to the Pacific Coast Ranges and Pacific Ocean. In California, it is found in the Klamath and California Coast Ranges as far south as the Santa Lucia Mountains with a small stand as far south as the Purisima Hills, Santa Barbara County. In the Sierra Nevada it ranges as far south as the Yosemite region. It occurs from near sea level along the coast to 1,800 metres (5,900 ft) in the California Mountains. Further inland, coast Douglas-fir is replaced by Rocky Mountain or interior Douglas-fir. Interior Douglas-fir intergrades with coast Douglas-fir in the Cascades of northern Washington and southern British Columbia.