This list of Mesozoic birds is a comprehensive list of all Mesozoic dinosaurs that have been assigned to the clade Avialae (birds, in the broadest sense). The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered avialan. The list currently includes 263 genera.
There is no official, canonical list of Mesozoic bird genera, but some of the more complete ones include Holtz's list of Mesozoic dinosaurs, [1] Molina-Perez & Larramendi's list of theropods, [2] and Mickey Mortimer's Theropod Database. [3] Lists of ex-birds are generally rare, but one example is the "Ex-Saurischian Taxa and Basal Saurischians" section of the Theropod Database. [4]
The authors column lists the authors of the formal description responsible for the erection of the genus listed. They are not necessarily the same as the authors of the type species, as sometimes a species from one genus is determined sufficiently distinct to warrant the erection of a new genus to house it. If this is the case, only the latter authors will be listed. The year column notes the year the genus' description was published.
Naming conventions and terminology follow the International Code of Zoological Nomenclature (ICZN). Technical terms used include:
The age column denotes the epoch of geologic time to which the fossils date. Genera that are invalid, misidentified, or otherwise do not represent a valid Mesozoic bird are listed as age N/A because there was never a time in which a Mesozoic bird by that generic name actually lived.
The location column designates the geographic region where remains of the relevant genus have been found. The regions used are countries, despite the fact that there were no political boundaries in Mesozoic times; they are only used for convenience. Genera that are invalid, misidentified, or otherwise do not represent a valid Mesozoic bird are listed as location N/A because there was never a place in which a Mesozoic bird by that generic name actually lived. The notes column is a collection of annotations on the scientific significance and taxonomic history of listed genera, as well as elaborations on the information presented in other columns.
Genus | Authors | Year | Status | Age | Location | Notes | Images |
---|---|---|---|---|---|---|---|
Abavornis [5] | Panteleyev | 1998 | Valid | Late Cretaceous | Uzbekistan | ||
Aberratiodontus [6] | Gong et al. | 2004 | Junior synonym | Early Cretaceous | China | Junior synonym of Yanornis. | |
Abitusavis [7] | Wang et al. | 2020 | Valid | Early Cretaceous | China | ||
Alamitornis [8] | Agonlin & Martinelli | 2009 | Valid | Late Cretaceous | Argentina | ||
Alcmonavis [9] | Rauhut et al. | 2019 | Valid | Late Jurassic | Germany | ||
Alethoalaornis | Li et al. | 2007 | Valid | Early Cretaceous | China | ||
Alexornis [10] | Brodkorb | 1976 | Valid | Late Cretaceous | Mexico | ||
Ambiortus | Kurochkin | 1982 | Valid | Early Cretaceous | Mongolia | ||
Ambopteryx [11] | Wang et al. | 2019 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Anatalavis | Olson & Parris | 1987 | Valid | Late Cretaceous | United States England | The type species is known from the Late Cretaceous–Paleocene border, while a referred species is from the Eocene. | |
Anchiornis [12] | Xu et al. | 2009 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Angelinornis | Kashin | 1972 | Junior synonym | Late Cretaceous | United States | Junior synonym of Ichthyornis. | |
Antarcticavis [13] | Cordes-Person et al. | 2020 | Valid | Late Cretaceous | Antarctica | ||
Apatornis | Marsh | 1873 | Valid | Late Cretaceous | United States | ||
Apsaravis [14] | Norell & Clarke | 2001 | Valid | Late Cretaceous | Mongolia | ||
Archaeopteryx [15] | Meyer | 1861 | Valid | Late Jurassic | Germany | Considered a non-avialan theropod by some researchers. | |
Archaeorhynchus [16] | Zhou & Zhang | 2006 | Valid | Early Cretaceous | China | ||
Archaeornis | Petronievics | 1917 | Junior synonym | Late Jurassic | Germany | Junior synonym of Archaeopteryx. | |
Archaeornithura [17] | Wang et al. | 2015 | Valid | Early Cretaceous | China | ||
Archaeovolans | Czerkas & Xu | 2002 | Junior synonym | Early Cretaceous | China | Junior synonym of Yanornis. | |
Asiahesperornis | Nesov & Prizemlin | 1991 | Valid | Late Cretaceous | Kazakhstan | ||
Asteriornis [18] | Field et al. | 2020 | Valid | Late Cretaceous | Belgium | ||
Aurornis [19] | Godefroit et al. | 2013 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Austinornis [20] | Clarke | 2004 | Valid | Late Cretaceous | United States | ||
Avimaia [21] | Bailleul et al. | 2019 | Valid | Early Cretaceous | China | ||
Avisaurus [22] | Brett-Surman & Paul | 1985 | Valid | Late Cretaceous | United States | ||
Balaur [23] | Csiki et al. | 2010 | Valid | Late Cretaceous | Romania | Originally (and sometimes still) thought to be a non-avialan theropod. | |
Baptornis | Marsh | 1877 | Valid | Late Cretaceous | United States | ||
Bauxitornis [24] | Dyke & Ősi | 2010 | Valid | Late Cretaceous | Hungary | ||
Beiguornis [25] | Wang et al. | 2022 | Valid | Early Cretaceous | China | ||
Bellulia | Wang et al. | 2016a | Preoccupied | N/A | N/A | Name preoccupied by the moth Bellulia Fibiger 2008 and replaced by Bellulornis. | |
Bellulornis | Wang et al. | 2016b | Valid | Early Cretaceous | China | ||
Bohaiornis [26] | Hu et al. | 2011 | Valid | Early Cretaceous | China | ||
Boluochia [27] | Zhou | 1995 | Valid | Early Cretaceous | China | ||
Bradycneme [28] | Harrison & Walker | 1975 | Misidentification | N/A | N/A | Originally described as a bird; more likely a non-avialan theropod. | |
Brevidentavis [29] | O'Connor et al. | 2021 | Valid | Early Cretaceous | China | ||
Brevirostruavis [30] | Li et al. | 2021 | Valid | Early Cretaceous | China | ||
Brodavis [31] | Martin et al. | 2012 | Valid | Late Cretaceous | United States Mongolia | ||
Caihong [32] | Hu et al. | 2018 | Valid | Late Jurassic | China | Considered to be a non-avialan theropod by some researchers. | |
Camptodontornis [33] | Demirjian | 2019 | Valid | Early Cretaceous | China | ||
Camptodontus [34] | Li et al. | 2010 | Preoccupied | N/A | N/A | Name preoccupied by the beetle Camptodontus Dejean, 1826 and replaced by Camptodontornis. | |
Canadaga [35] | Hou | 1999 | Valid | Late Cretaceous | Canada | ||
Castignovolucris [36] | Buffetaut et al. | 2023 | Valid | Late Cretaceous | France | ||
Catenoleimus [5] | Panteleyev | 1998 | Valid | Late Cretaceous | Uzbekistan | ||
Cathayornis | Zhou et al. | 1992 | Valid | Early Cretaceous | China | ||
Ceramornis [37] | Brodkorb | 1963 | Valid | Late Cretaceous | United States | ||
Cerebavis [38] | Kurochkin et al. | 2006 | Valid | Late Cretaceous | Russia | ||
Changchengornis [39] | Ji et al. | 1999 | Valid | Early Cretaceous | China | ||
Changmaornis [40] | Wang et al. | 2013 | Valid | Early Cretaceous | China | ||
Changzuiornis [41] | Huang et al. | 2016 | Disputed | Early Cretaceous | China | Possibly synonymous with Juehuaornis. | |
Chaoyangia | Hou & Zhang | 1993 | Valid | Early Cretaceous | China | ||
Chiappeavis [42] | O'Connor et al. | 2016 | Valid | Early Cretaceous | China | ||
Chongmingia [43] | Wang et al. | 2016 | Valid | Early Cretaceous | China | ||
Chupkaornis [44] | Tanaka et al. | 2017 | Valid | Late Cretaceous | Japan | ||
Cimoliornis | Owen | 1846 | Misidentification | N/A | N/A | Originally described as a bird; actually a pterosaur. | |
Cimolopteryx | Marsh | 1892 | Valid | Late Cretaceous | United States | May be Paleocene. | |
Colonosaurus [45] | Marsh | 1872 | Junior synonym | Late Cretaceous | United States | Junior synonym of Ichthyornis. | |
Concornis [46] | Sanz & Buscalioni | 1992 | Valid | Early Cretaceous | Spain | ||
Confuciusornis [47] | Hou et al. | 1995 | Valid | Early Cretaceous | China | ||
Coniornis | Marsh | 1893 | Junior synonym | Late Cretaceous | United States | Junior synonym of Hesperornis. | |
Cosesaurus [48] | Ellenberger & de Villalta | 1974 | Misidentification | N/A | N/A | Sometimes proposed to be a bird ancestor; actually a tanystropheid. | |
Cratoavis [49] | Carvalho et al. | 2015 | Valid | Early Cretaceous | Brazil | ||
Cratonavis [50] | Li et al. | 2023 | Valid | Early Cretaceous | China | ||
Cretaaviculus | Bazhanov | 1969 | Valid | Late Cretaceous | Kazakhstan | ||
Cretornis [51] | Fritsch | 1880 | Misidentification | N/A | N/A | Originally described as a bird; actually a pterosaur. | |
Cruralispennia [52] | Wang et al. | 2017 | Valid | Early Cretaceous | China | ||
Cuspirostrisornis [53] | Hou | 1997 | Valid | Early Cretaceous | China | ||
Dalianraptor | Gao & Liu | 2005 | Disputed | N/A | N/A | Likely a chimaera consisting of a Jeholornis and another theropod. | |
Dalingheornis [54] | Zhang et al. | 2006 | Valid | Early Cretaceous | China | ||
Dapingfangornis [55] | Li et al. | 2006 | Valid | Early Cretaceous | China | ||
Didactylornis [56] | Yuan | 2008 | Junior synonym | Early Cretaceous | China | Junior synonym of Sapeornis. | |
Dingavis [57] | O'Connor et al. | 2016 | Disputed | Early Cretaceous | China | Possibly synonymous with Changzuiornis and/or Juehuaornis. | |
Dunhuangia [58] | Wang et al. | 2015 | Valid | Early Cretaceous | China | ||
Elbretornis [59] | Walker & Dyke | 2009 | Valid | Late Cretaceous | Argentina | Possibly synonymous with Lectavis, Soroavisaurus and/or Yungavolucris. | |
Elektorornis [60] | Xing et al. | 2019 | Valid | Early Cretaceous | Myanmar | ||
Elopteryx [61] | Andrews | 1913 | Misidentification | N/A | N/A | Originally described as a bird; more likely to be a non-avialan theropod. | |
Elsornis [62] | Chiappe et al. | 2007 | Valid | Late Cretaceous | Mongolia | ||
Enaliornis | Seeley | 1876 | Valid | Late Cretaceous | England | ||
Enantiophoenix [63] | Cau & Arduini | 2008 | Valid | Late Cretaceous | Lebanon | ||
Enantiornis [64] | Walker | 1981 | Valid | Late Cretaceous | Argentina | ||
Eoalulavis [65] | Sanz et al. | 1996 | Valid | Early Cretaceous | Spain | ||
Eocathayornis [66] | Zhou | 2002 | Valid | Early Cretaceous | China | ||
Eoconfuciusornis [67] | Zhang et al. | 2008 | Valid | Early Cretaceous | China | ||
Eoenantiornis [68] | Hou et al. | 1999 | Valid | Early Cretaceous | China | ||
Eogranivora [69] | Zheng et al. | 2018 | Valid | Early Cretaceous | China | ||
Eopengornis | Wang et al. | 2014 | Valid | Early Cretaceous | China | ||
Eosinopteryx [70] | Godefroit et al. | 2013 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Epidendrosaurus [71] | Zhang et al. | 2002 | Junior synonym | Late Jurassic | China | Possible synonym of Scansoriopteryx. | |
Epidexipteryx [72] | Zhang et al. | 2008 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Eurolimnornis [73] | Kessler & Jurcsák | 1986 | Misidentification | N/A | N/A | Originally described as a bird; reinterpreted as a pterodactyloid pterosaur. | |
Evgenavis [74] | O'Connor et al. | 2014 | Valid | Early Cretaceous | Russia | ||
Explorornis [5] | Panteleyev | 1998 | Valid | Late Cretaceous | Uzbekistan | ||
Falcatakely [75] | O'Connor et al. | 2020 | Valid | Late Cretaceous | Madagascar | ||
Feitianius [76] | O'Connor et al. | 2015 | Valid | Early Cretaceous | China | ||
Flexomornis [77] | Tykoski & Fiorillo | 2010 | Valid | Late Cretaceous | United States | ||
Fortipesavis [78] | Clark & O'Connor | 2021 | Valid | Early Cretaceous | Myanmar | ||
Fortunguavis [79] | Wang et al. | 2014 | Valid | Early Cretaceous | China | ||
Fujianvenator [80] | Xu et al. | 2023 | Valid | Late Jurassic | China | Found within the family Anchiornithidae, which may represent non-avialan dinosaurs | |
Fukuipteryx [81] | Imai et al. | 2019 | Valid | Early Cretaceous | Japan | ||
Fumicollis [82] | Bell & Chiappe | 2015 | Valid | Late Cretaceous | United States | ||
Gallornis | Lambrecht | 1931 | Valid | Early Cretaceous | France | ||
Gansus [83] | Hou & Liu | 1984 | Valid | Early Cretaceous | China | ||
Gargantuavis [84] | Buffetaut & Le Loeuff | 1998 | Valid | Late Cretaceous | France Spain Romania | ||
Gettyia [85] | Atterholt et al. | 2018 | Valid | Late Cretaceous | United States | ||
Gobipipus [86] | Kurochkin & Chatterjee | 2013 | Valid | Early Cretaceous | Mongolia | Very similar to embryonic Gobipteryx. | |
Gobipteryx [87] | Elżanowski | 1994 | Valid | Late Cretaceous | Mongolia | ||
Grabauornis [88] | Dalsätt et al. | 2014 | Valid | Early Cretaceous | China | ||
Gracilornis [89] | Li & Hou | 2011 | Valid | Early Cretaceous | China | ||
Graculavus | Marsh | 1872 | Valid | Late Cretaceous | United States | May be Paleocene. | |
Gretcheniao | Chiappe et al. | 2019 | Valid | Early Cretaceous | China | ||
Griphornis | Woodward | 1862 | Junior synonym | Late Jurassic | Germany | Name officially rejected in favor of Archaeopteryx. | |
Griphosaurus | Wagner | 1862 | Junior synonym | Late Jurassic | Germany | Name officially rejected in favor of Archaeopteryx. | |
Guildavis [20] | Clarke | 2004 | Valid | Late Cretaceous | United States | ||
Gurilynia [90] | Kurochkin | 1999 | Valid | Late Cretaceous | Mongolia | ||
Halimornis [91] | Chiappe | 2002 | Valid | Late Cretaceous | United States | ||
Hargeria | Lucas | 1903 | Junior synonym | Late Cretaceous | United States | Junior synonym of Hesperornis. | |
"Hebeiornis" [92] | Xu et al. | 1999 | Nomen nudum | N/A | N/A | Synonym of Vescornis due to being based on the same holotype. However, despite predating the description of Vescornis by five years, its description is so poor that it is regarded as a nomen nudum and a synonym. | |
Heptasteornis [28] | Harrison & Walker | 1975 | Misidentification | N/A | N/A | Originally described as a bird; more likely a non-avialan theropod. | |
Hesperornis | Marsh | 1872 | Valid | Late Cretaceous | United States Canada Russia | ||
Holbotia [93] | Zelenkov & Averianov | 2015 | Valid | Early Cretaceous | Mongolia | ||
Hollanda [94] | Bell et al. | 2010 | Valid | Late Cretaceous | Mongolia | ||
Hongshanornis [95] | Zhou & Zhang | 2005 | Valid | Early Cretaceous | China | ||
Horezmavis | Nessov & Borkin | 1983 | Valid | Late Cretaceous | Uzbekistan | ||
Houornis [96] | Wang & Liu | 2015 | Valid | Early Cretaceous | China | ||
Huoshanornis [97] | Wang et al. | 2010 | Valid | Early Cretaceous | China | ||
Iaceornis [20] | Clarke | 2004 | Valid | Late Cretaceous | United States | ||
Iberomesornis [98] | Sanz & Bonaparte | 1992 | Valid | Early Cretaceous | Spain | ||
Ichthyornis | Marsh | 1873 | Valid | Late Cretaceous | United States | ||
Ilerdopteryx | Lacasa-Ruiz | 1985 | Valid | Early Cretaceous | Spain | ||
Imparavis [99] | Wang et al. | 2024 | Valid | Early Cretaceous | China | ||
Incolornis [5] | Panteleyev | 1998 | Valid | Late Cretaceous | Uzbekistan | ||
Intiornis [100] | Novas et al. | 2010 | Valid | Late Cretaceous | Argentina | ||
Iteravis | Zhou et al. | 2014 | Valid | Early Cretaceous | China | ||
Janavis [101] | Benito et al. | 2022 | Valid | Late Cretaceous | Belgium | ||
Jeholornis [102] | Zhou et al. | 2002 | Valid | Early Cretaceous | China | ||
Jianchangornis [103] | Zhou et al. | 2009 | Valid | Early Cretaceous | China | ||
Jibeinia [53] | Hou | 1997 | Valid | Early Cretaceous | China | ||
Jinfengopteryx [104] | Ji et al. | 2005 | Misidentification | N/A | N/A | Originally described as a bird; actually a non-avian dinosaur. | |
Jinguofortis [105] | Wang et al. | 2018 | Valid | Early Cretaceous | China | ||
Jinzhouornis | Hou et al. | 2002 | Junior synonym | Early Cretaceous | China | Junior synonym of Confuciusornis. | |
Jiuquanornis [40] | Wang et al. | 2013 | Valid | Early Cretaceous | China | ||
Jixiangornis [106] | Ji et al. | 2002 | Valid | Early Cretaceous | China | ||
Judinornis | Nessov & Borkin | 1983 | Valid | Late Cretaceous | Mongolia | ||
Juehuaornis [107] | Wang et al. | 2015 | Valid | Early Cretaceous | China | ||
Junornis [108] | Liu et al. | 2017 | Valid | Early Cretaceous | China | ||
Jurapteryx | Howgate | 1984 | Junior synonym | Late Jurassic | Germany | Junior synonym of Archaeopteryx. | |
Kaririavis [109] | Carvalho et al. | 2021 | Valid | Early Cretaceous | Brazil | ||
Khinganornis [110] | Wang et al. | 2020 | Valid | Early Cretaceous | China | ||
Kizylkumavis [111] | Nessov | 1984 | Valid | Late Cretaceous | Uzbekistan | ||
Kompsornis [112] | Wang et al. | 2020 | Valid | Early Cretaceous | China | ||
Kookne | Novas et al. | 2019 | Valid | Late Cretaceous | Argentina | ||
Kuszholia [113] | Nesov | 1992 | Valid | Late Cretaceous | Uzbekistan | ||
Lamarqueavis [114] | Agnolin | 2010 | Valid | Late Cretaceous | Argentina United States Canada | ||
Laopteryx [115] | Marsh | 1881 | Misidentification | N/A | N/A | Originally described as a bird; actually a pterosaur. | |
Laornis [116] | Marsh | 1870 | Valid | Late Cretaceous | United States | May be Paleocene. | |
Largirostrornis [53] | Hou | 1997 | Disputed | Early Cretaceous | China | Possible synonym of Cathayornis. | |
Lectavis [117] | Chiappe | 1993 | Valid | Late Cretaceous | Argentina | ||
Lenesornis [118] | Kurochkin | 1996 | Valid | Late Cretaceous | Uzbekistan | ||
Lestornis [119] | Marsh | 1876 | Junior synonym | Late Cretaceous | United States | Junior synonym of Hesperornis. | |
Liaoningornis [120] | Hou | 1996 | Valid | Early Cretaceous | China | ||
Liaoxiornis [121] | Hou & Chen | 1999 | Valid | Early Cretaceous | China | ||
Limenavis [122] | Clarke & Chiappe | 2001 | Valid | Late Cretaceous | Argentina | ||
Limnornis [123] | Kessler & Jurcsak | 1984 | Misidentification | N/A | N/A | Name preoccupied by the ovenbird Limnornis Gould 1839 and replaced by Palaeocursornis. Originally described as a bird; actually a pterosaur. | |
Lingyuanornis [124] | Ji & Ji | 1999 | Junior synonym | N/A | N/A | Junior synonym of Liaoxiornis as it is based on the same holotype. | |
Linyiornis [125] | Wang et al. | 2016 | Valid | Early Cretaceous | China | ||
Lisboasaurus [126] | Seiffert | 1970 | Misidentification | N/A | N/A | Sometimes proposed to be a bird; actually a crocodilian. | |
Lonchodytes [37] | Brodkorb | 1963 | Valid | Late Cretaceous | United States | May be Paleocene. | |
Longchengornis [53] | Hou | 1997 | Disputed | Early Cretaceous | China | Possible synonym of Cathayornis. | |
Longicrusavis [127] | O'Connor et al. | 2010 | Valid | Early Cretaceous | China | ||
Longipteryx [128] | Zhang et al. | 2001 | Valid | Early Cretaceous | China | ||
Longirostravis [129] | Hou et al. | 2004 | Valid | Early Cretaceous | China | ||
Longisquama [130] | Sharov | 1970 | Misidentification | N/A | N/A | Sometimes proposed to be a bird ancestor; actually an indeterminate reptile. | |
Longusunguis [131] | Wang et al. | 2014 | Valid | Early Cretaceous | China | ||
Maaqwi [132] | McLachlan et al. | 2017 | Valid | Late Cretaceous | Canada | ||
Magnusavis [133] | Clark et al. | 2024 | Valid | Late Cretaceous | United States | ||
Martinavis [134] | Walker et al. | 2007 | Valid | Late Cretaceous | France Argentina | ||
Meemannavis [29] | O'Connor et al. | 2021 | Valid | Early Cretaceous | China | ||
Megalancosaurus [135] | Calzavara et al. | 1980 | Misidentification | N/A | N/A | Sometimes proposed to be a bird ancestor, but actually a drepanosaurid. | |
Mengciusornis [136] | Wang et al. | 2019 | Valid | Early Cretaceous | China | ||
Microenantiornis | Wei & Li | 2017 | Valid | Early Cretaceous | China | ||
Mirarce [85] | Atterholt et al. | 2018 | Valid | Late Cretaceous | United States | ||
Mirusavis [137] | Wang et al. | 2020 | Valid | Early Cretaceous | China | ||
Monoenantiornis [138] | Hu & O'Connor | 2016 | Valid | Early Cretaceous | China | ||
Musivavis [139] | Wang et al. | 2022 | Valid | Early Cretaceous | China | ||
Mystiornis [140] | Kurochkin et al. | 2011 | Valid | Early Cretaceous | Russia | ||
Nanantius [141] | Molnar | 1986 | Valid | Early Cretaceous | Australia | ||
Neimengornis [142] | Wang et al. | 2021 | Valid | Early Cretaceous | China | ||
Neogaeornis [143] | Lambrecht | 1929 | Valid | Late Cretaceous | Chile | ||
Neuquenornis [144] | Chiappe & Calvo | 1994 | Valid | Late Cretaceous | Argentina | ||
Noguerornis [145] | Lacasa-Ruiz | 1989 | Valid | Early Cretaceous | Spain | ||
Novacaesareala | Parris & Hope | 2002 | Valid | Late Cretaceous | United States | May be Paleocene. | |
Oculudentavis [146] | Xing et al. | 2020 | Misidentification | N/A | N/A | Originally described as a bird; reinterpreted as an indeterminate lizard. | |
Omnivoropteryx [147] | Czerkas & Ji | 2002 | Valid | Early Cretaceous | China | ||
Orienantius | Liu et al. | 2019 | Valid | Early Cretaceous | China | ||
Ostromia [148] | Foth & Rauhut | 2017 | Valid | Late Jurassic | Germany | Considered a non-avialan theropod by some researchers. | |
Otogornis | Hou | 1994 | Valid | Early Cretaceous | China | ||
Palaeocursornis [73] | Kessler & Jurcsak | 1986 | Misidentification | N/A | N/A | Originally described as a bird; actually a pterosaur. | |
Palaeonornis | Emmons | 1857 | Misidentification | N/A | N/A | Originally described as a bird; actually a junior synonym of the phytosaur Rutiodon . [4] | |
Palaeopteryx | Jensen | 1981 | Misidentification | N/A | N/A | Originally described as a bird; more likely a non-avian theropod. | |
Palaeornis | Mantell | 1844 | Misidentification | N/A | N/A | Name preoccupied by the parrot Palaeornis Vigors 1825. Originally described as a bird; actually a pterosaur. | |
Palaeotringa | Marsh | 1870 | Valid | Late Cretaceous | United States | May be Paleocene. | |
Palintropus [149] | Brodkorb | 1970 | Valid | Late Cretaceous | United States Canada | ||
Parabohaiornis [131] | Wang et al. | 2014 | Valid | Early Cretaceous | China | ||
Parahesperornis [150] | Martin | 1984 | Valid | Late Cretaceous | United States | ||
Parahongshanornis [151] | Li et al. | 2011 | Valid | Early Cretaceous | China | ||
Parapengornis [152] | Hu et al. | 2015 | Valid | Early Cretaceous | China | ||
Paraprotopteryx [153] | Zheng et al. | 2007 | Valid | Early Cretaceous | China | ||
Parascaniornis | Lambrecht | 1933 | Junior synonym | Late Cretaceous | Sweden | Junior synonym of Baptornis. | |
Parvavis [154] | Wang et al. | 2014 | Valid | Late Cretaceous | China | ||
Pasquiaornis [155] | Tokaryk et al. | 1997 | Valid | Late Cretaceous | Canada | ||
Patagopteryx [156] | Alvarenga & Bonaparte | 1992 | Valid | Late Cretaceous | Argentina | ||
Pedopenna [157] | Xu & Zhang | 2005 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Pelagornis | Seeley | 1866 | Preoccupied | N/A | N/A | Named preoccupied by the Miocene bird Pelagornis Lartet 1857 and replaced by Enaliornis. | |
Pengornis [158] | Zhou et al. | 2008 | Valid | Early Cretaceous | China | ||
Piksi [159] | Varricchio | 2002 | Misidentification | N/A | N/A | Originally described as a bird; reinterpreted as a pterodactyloid pterosaur. | |
Piscivoravis [160] | Zhou et al. | 2013 | Valid | Early Cretaceous | China | ||
Piscivorenantiornis [161] | Wang & Zhou | 2017 | Valid | Early Cretaceous | China | ||
Platanavis [113] | Nesov | 1992 | Valid | Late Cretaceous | Uzbekistan | ||
Plegadornis | Wetmore | 1962 | Preoccupied | N/A | N/A | Name preoccupied by the modern bird Plegadornis Brehm 1855; junior synonym of Ichthyornis. | |
Polarornis [162] | Chatterjee | 2002 | Valid | Late Cretaceous | Antarctica | ||
Potamornis [163] | Elzanowski et al. | 2001 | Valid | Late Cretaceous | United States | ||
Praeornis [164] | Rautian | 1978 | Valid | Late Jurassic | Kazakhstan | ||
Priscavolucris [165] | Gomez Pallerola | 1979 | Misidentification | N/A | N/A | Originally described as a bird; reinterpreted as a hybodontiform fish. [166] | |
"Proornis" | Lim | 1993 | Nomen nudum | N/A | N/A | ||
Protarchaeopteryx [167] | Ji & Ji | 1997 | Misidentification | N/A | N/A | Originally described as a bird; actually a non-avian dinosaur. | |
Protoavis [168] | Chatterjee | 1991 | Misidentification | N/A | N/A | Originally described as a bird; more likely a chimaera of several vertebrates. | |
Protopteryx [169] | Zhang & Zhou | 2000 | Valid | Early Cretaceous | China | ||
Pterodactylus [170] | Rafinesque | 1815 | Misidentification | N/A | N/A | Sometimes thought to be a bird before being described as a pterosaur. | |
Pterygornis [171] | Wang et al. | 2015 | Valid | Early Cretaceous | China | ||
Qiliania [172] | Ji et al. | 2011 | Valid | Early Cretaceous | China | ||
Rahona [173] | Forster et al. | 1998a | Preoccupied | N/A | N/A | Name preoccupied by the moth Rahona Griveaud 1975 and replaced by Rahonavis. | |
Rahonavis [174] | Forster et al. | 1998b | Valid | Late Cretaceous | Madagascar | Considered a non-avialan theropod by some researchers. | |
Rapaxavis [175] | Morschhauser et al. | 2009 | Valid | Early Cretaceous | China | ||
Rhamphorhynchus [176] | Meyer | 1846 | Misidentification | N/A | N/A | Originally thought to be a bird before being described as a pterosaur. | |
Samrukia [177] | Naish et al. | 2012 | Misidentification | N/A | N/A | Originally described as a bird; actually a pterosaur. | |
Sapeornis [178] | Zhou & Zhang | 2002 | Valid | Early Cretaceous | China | ||
Sazavis [179] | Nesov | 1989 | Valid | Late Cretaceous | Uzbekistan | ||
Scansoriopteryx [180] | Czerkas & Yuan | 2002 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Schizooura [181] | Zhou et al. | 2012 | Valid | Early Cretaceous | China | ||
Serikornis [182] | Lefèvre et al. | 2017 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Shangyang | Wang & Zhou | 2019 | Valid | Early Cretaceous | China | ||
Shanweiniao [183] | O'Connor et al. | 2009 | Valid | Early Cretaceous | China | ||
Shengjingornis [184] | Li et al. | 2012 | Valid | Early Cretaceous | China | ||
Shenqiornis [185] | Wang et al. | 2010 | Valid | Early Cretaceous | China | ||
Shenshiornis [186] | Hu | 2010 | Junior synonym | Early Cretaceous | China | Junior synonym of Sapeornis. | |
Shenzhouraptor [187] | Ji et al. | 2002 | Disputed | Early Cretaceous | China | Possible synonym of Jeholornis. | |
Shuilingornis [188] | Wang et al. | 2024 | Valid | Early Cretaceous | China | ||
Similiyanornis [7] | Wang et al. | 2020 | Valid | Early Cretaceous | China | ||
Sinornis [189] | Sereno & Rao | 1992 | Valid | Early Cretaceous | China | ||
Songlingornis [53] | Hou | 1997 | Valid | Early Cretaceous | China | ||
Soroavisaurus [190] | Chiappe | 1993 | Valid | Late Cretaceous | Argentina | ||
Struthiosaurus [191] | Bunzel | 1871 | Misidentification | N/A | N/A | Originally thought to be a bird; actually an ankylosaurian dinosaur. | |
"Styginetta" | Stidham | 2001 | Nomen nudum | N/A | N/A | Obscure; named in a thesis. | |
Sulcavis [192] | O'Connor et al. | 2013 | Valid | Early Cretaceous | China | ||
Telmatornis | Marsh | 1870 | Valid | Late Cretaceous | United States | ||
Teviornis [193] | Kurochkin et al. | 2002 | Valid | Late Cretaceous | Mongolia | ||
Tianyuornis [194] | Zheng et al. | 2014 | Valid | Early Cretaceous | China | ||
Tingmiatornis [195] | Bono et al. | 2016 | Valid | Late Cretaceous | Canada | ||
Torotix [37] | Brodkorb | 1963 | Valid | Late Cretaceous | United States | ||
Tytthostonyx [196] | Olson & Parris | 1987 | Valid | Late Cretaceous | United States | ||
Vegavis [197] | Clarke et al. | 2005 | Valid | Late Cretaceous | Antarctica | ||
Vescornis [198] | Zhang et al. | 2004 | Disputed | Early Cretaceous | China | Possible junior synonym of Jibeinia. | |
Volgavis [179] | Nessov & Yarkov | 1989 | Valid | Late Cretaceous | Russia | ||
Vorona [199] | Forster et al. | 1996 | Valid | Late Cretaceous | Madagascar | ||
Wasaibpanchi | Malkani | 2021 | Disputed | Late Cretaceous | Pakistan | Publication status uncertain. | |
Wellnhoferia [200] | Elżanowski | 2001 | Disputed | Late Jurassic | Germany | Possible junior synonym of Archaeopteryx. | |
Wyleyia [201] | Harrison & Walker | 1973 | Valid | Early Cretaceous | England | ||
Xiangornis [202] | Hu et al. | 2012 | Valid | Early Cretaceous | China | ||
Xinghaiornis [203] | Wang et al. | 2013 | Valid | Early Cretaceous | China | ||
Xiaotingia [204] | Xu et al. | 2011 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Yandangornis [205] | Cai & Zhao | 1999 | Valid | Late Cretaceous | China | ||
Yangavis [206] | Wang et al. | 2018 | Valid | Early Cretaceous | China | ||
Yanornis [207] | Zhou & Zhang | 2001 | Valid | Early Cretaceous | China | ||
Yatenavis [208] | Herrera et al. | 2022 | Valid | Late Cretaceous | Argentina | ||
Yi [209] | Xu et al. | 2015 | Valid | Late Jurassic | China | Considered a non-avialan theropod by some researchers. | |
Yixianornis | Zhou & Zhang | 2001 | Valid | Early Cretaceous | China | ||
Yixianosaurus [210] | Xu & Wang | 2003 | Valid | Early Cretaceous | China | Considered a non-avialan theropod by some researchers. | |
Yuanchuavis [211] | Wang et al. | 2021 | Valid | Early Cretaceous | China | ||
Yuanjiawaornis [212] | Hu et al. | 2015 | Valid | Early Cretaceous | China | ||
Yumenornis [40] | Wang et al. | 2013 | Valid | Early Cretaceous | China | ||
Yungavolucris [213] | Chiappe | 1993 | Valid | Late Cretaceous | Argentina | ||
Yuornis [214] | Xu et al. | 2021 | Valid | Late Cretaceous | China | ||
Zhongjianornis [215] | Zhou et al. | 2010 | Valid | Early Cretaceous | China | ||
Zhongornis [216] | Gao et al. | 2008 | Valid | Early Cretaceous | China | ||
Zhouornis [217] | Zhang et al. | 2013 | Valid | Early Cretaceous | China | ||
Zhylgaia [218] | Nessov | 1988 | Valid | Late Cretaceous | Kazakhstan | Dating is uncertain; may be Late Cretaceous or Paleogene. | |
Zhyraornis [111] | Nessov | 1984 | Valid | Late Cretaceous | Uzbekistan |
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.
Yanornis is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: Yanornis martini, based on several fossils found in the 120-million-year-old Jiufotang Formation at Chaoyang, and Yanornis guozhangi, from the 124-million-year-old Yixian Formation.
The Xiagou Formation is the middle strata of the Xinminbao Group. It is named for its type site in Xiagou, in the Changma Basin of Gansu Province, northwestern China and is considered Early Cretaceous in age. It is known outside the specialized world of Chinese geology as the site of a Lagerstätte in which the fossils were preserved of Gansus yumenensis, the earliest true modern bird.
Longipteryx is a genus of prehistoric bird which lived during the Early Cretaceous. It contains a single species, Longipteryx chaoyangensis. Its remains have been recovered from the Jiufotang Formation at Chaoyang in Liaoning Province, China. Apart from the holotype IVPP V 12325 - a fine and nearly complete skeleton — another entire skeleton and some isolated bones are known to date.
Cathayornis is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known definitively from only one species, Cathayornis yandica, one of the first Enantiornithes found in China. Several additional species were once incorrectly classified as Cathayornis, and have since been reclassified or regarded as nomina dubia.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2002.
Ornithothoraces is a group of avialan dinosaurs that includes all enantiornithes and the euornithes, which includes modern birds and their closest ancestors. The name Ornithothoraces means "bird thoraxes". This refers to the modern, highly advanced anatomy of the thorax that gave the ornithothoracines superior flight capability compared with more primitive avialans. This anatomy includes a large, keeled breastbone, elongated coracoids and a modified glenoid joint in the shoulder, and a semi-rigid rib cage. In spite of this at least the sternum seems to have developed convergently rather than being a true homology.
Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces.
The Jiufotang Formation is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms. It is a member of the Jehol group. The exact age of the Jiufotang has been debated for years, with estimates ranging from the Late Jurassic to the Early Cretaceous. New uranium-lead dates reveal the formation is deposited in the Aptian stage of the Early Cretaceous. Fossils of Microraptor and Jeholornis are from the Jiufotang.
Protopteryx is an extinct bird and possibly the basalmost enantiornithean, from the Cretaceous period. The type species is P. fengningensis. It was first discovered in the Sichakou Member of the Yixian Formation or Huajiying Formation of Hebei Province, northern China, dating from 131 Ma ago. Protopteryx has been found in the Daibeigou formation, as well. The name Protopteryx means "primitive feather": "proto-" meaning "the first of" and "-pteryx" meaning "feather" or "wing." The name comes from the fact that Protopteryx feathers are more primitive than those of modern birds, such as the two elongated tail feathers that lack barbs and rami.
Shanweiniao is a genus of long-snouted enantiornithean birds from Early Cretaceous China. One species is known, Shanweiniao cooperorum. There is one known fossil, a slab and counterslab. The fossil is in the collection of the Dalian Natural History Museum, and has accession number DNHM D1878/1 and DNHM1878/2. It was collected from the Lower Cretaceous Dawangzhengzi Beds, middle Yixian Formation, from Lingyuan in the Liaoning Province, China.
The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans about 1.6 million years during the early Cretaceous period. It is known for its fossils, listed below.
Hongshanornithidae is an extinct group of early ornithuromorph birds from the early Cretaceous period of China. It includes the genera Hongshanornis and Tianyuornis from the Yixian Formation of Inner Mongolia, Longicrusavis from the Yixian Formation of Liaoning Province, Parahongshanornis from the Jiufotang Formation of Liaoning Province, and Archaeornithura, the oldest known member, from the Huajiying Formation of Hebei Province.
Longipterygidae is a family of early enantiornithean avialans from the Early Cretaceous epoch of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125-120 million years ago.
Bohaiornis is a genus of enantiornithean dinosaurs. Fossils have been found from the Lower Cretaceous Jiufotang Formation of western Liaoning, China. The only known species, Bohaiornis guoi, was named by Dongyu Hu, Li Li, Lianhaim Hou and Xing Xu in 2011 on the basis of a fully articulated and well-preserved skeleton of a sub-adult. This specimen, LPM B00167, preserved two long, ribbon-like feathers attached to the tail rather than a fan of shorter pennaceous feathers. It was similar to the slightly older Eoenantiornis, but much larger in size. Bohaiornis is the type species of Bohaiornithidae, a family of large predatory enantiornitheans from the Early Cretaceous.
Bohaiornithidae is a group of early predatory enantiornithean dinosaurs from the early Cretaceous Period of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125–120 million years ago. Bohaiornithidae was first coined by Wang and colleagues in 2014. They defined it as the natural group formed by all descendants of the common ancestor of the type species, Bohaiornis guoi, and Shenqiornis mengi.
Luis María Chiappe is an Argentine paleontologist born in Buenos Aires who is best known for his discovery of the first sauropod nesting sites in the badlands of Patagonia in 1997 and for his work on the origin and early evolution of Mesozoic birds. He is currently the Vice President of Research and Collections at the Natural History Museum of Los Angeles County and director of the museum's Dinosaur Institute. He was a postdoctoral researcher at the American Museum of Natural History, New York after immigrating from Argentina. Chiappe is currently the curator of the award winning Dinosaur Hall at the Natural History Museum of Los Angeles County, an adjunct professor at the University of Southern California, BBC advisor and author of scientific and popular books.
Pengornithidae is a group of early enantiornithines from the early Cretaceous Period of China, with the putative member Falcatakely possibly extending this clade's range into the Late Cretaceous of Madagascar, and several putative pengornithids also hail from this formation. Specimens of these animals have been found both in the Huajiying Formation and Jiufotang Formation of Liaoning and Hebei provinces, dating from the Hauterivian age to the Aptian age.
Parapengornis is an extinct genus of enantiornithine bird from the Lower Cretaceous of what is now China. The holotype specimen was discovered in the Jiufotang Formation near Lingyuan, western Liaoning province, and was catalogued as IVPP V18687. The nearly complete, articulated specimen is preserved on a slab and has impressions of pennaceous feathers. Only parts of the sternum, the left hand, and right foot are missing. In 2015, it became the basis of the new genus and species Parapengornis eurycaudatus, named by the Chinese palaeontologists Han Hu, Jingmai K. O’Connor, and Zhonghe Zhou. The generic name consists of the Latin word para and the name of the related genus Pengornis, indicating their close relationship. The name Pengornis is itself derived from "Peng", a mythological bird from Chinese folklore, and ornis, which means bird in Greek. The specific name is derived from the Latin words eury, meaning broad, and caudatus, meaning tail, in reference to the broad and expanded pygostyle. A nearly complete specimen formerly assigned to Pengornis was also reassigned to Parapengornis by these authors.
Jingmai Kathleen O'Connor is a paleontologist who works as a curator at the Field Museum.
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