"Sue", AMNH 5027, "Stan", and "Jane", to scale with a human
Tyrannosaurus is one of the most iconic dinosaurs and is known from numerous specimens, some of which have individually acquired notability due to their scientific significance and media coverage.
Described as Gorgosaurus lancensis and later given its own genus, Nanotyrannus. Now considered a juvenile specimen of Tyrannosaurus rex by most researchers.
One of the best-known specimens, with casts present in numerous institutions. Sold to a private owner in 2020 for US$31 million, formerly in the Black Hills Institute as BHI 3033.
Proposed to be largest known specimen, scientific consensus not yet reached. Exceeds "Sue" in 84.6% of the published bone measurements. Has been estimated to be larger than "Sue" in two published studies.
Considered the 14th T-rex specimen to be found. The first in-situ t-rex to be known from the state of North Dakota. Discovered and collected by amateurs and shows tooth marks on bones, indicating scavenging.
Currently the most complete specimen known to science,an adolescent preserved entangled with a Triceratops. Like Jane, a minority of paleontologists consider it a different taxon.
The specimen belongs to a private collector, but 3D-printed replicas of the bones are accessioned in the Nottingham Natural History Museum collection under the number NCMG 2021-7[1]
The first-named fossil specimen which can be attributed to Tyrannosaurus rex consists of two partial vertebrae (one of which has been lost) found by Edward Drinker Cope in 1892. Cope believed that they belonged to an "agathaumid" (ceratopsid) dinosaur, and named them Manospondylus gigas, meaning "giant porous vertebra" in reference to the numerous openings for blood vessels he found in the bone.[3] The M. gigas remains were later identified as those of a theropod rather than a ceratopsid, and H.F. Osborn recognized the similarity between M. gigas and Tyrannosaurus rex as early as 1917. However, due to the fragmentary nature of the Manospondylus vertebrae, Osborn did not synonymize the two genera.[4]
The holotype of Tyrannosaurus rex, a partial skull and skeleton originally called AMNH 973 (AMNH stands for American Museum of Natural History), was discovered in the U.S. state of Montana in 1902 and excavated over the next three years. Another specimen (AMNH 5866), found in Wyoming in 1900, was described in the same paper under the name Dynamosaurus imperiosus. At the time of their initial description and naming, these specimens had not been fully prepared and the type specimen of T. rex had not even been fully recovered.[5] In 1906, after further preparation and examination, Henry Fairfield Osborn recognized both skeletons as belonging to the same species. Because the name Tyrannosaurus rex had appeared just one page earlier than Dynamosaurus in Osborn's 1905 work, it was considered the older name and has been used since. Had it not been for page order, Dynamosaurus would have become the official name.[6]
Holotype: CM 9380
Reconstructed mount of T. rex holotype CM 9380
CM 9380 is the type specimen used to describe Tyrannosaurus rex. Fragments of (then) AMNH 973 were first found in 1902 by Barnum Brown, assistant curator of the American Museum of Natural History and a famous paleontologist in his own right. He forwarded news of it to Osborn; it would be three years before they found the rest of it.[citation needed] In 1905 when the type was described by Osborn, previous knowledge of dinosaur predators at the time were based on Jurassiccarnosaurs, so the short fore-arms of the Tyrannosaurus were treated with extreme caution, with suspicion that bones of a smaller theropod had become jumbled with the remains of the bigger fossil.[5][7] Following the 1941 entry of the United States into World War II, the holotype was sold to the Carnegie Museum of Natural History in Pittsburgh for protection against possible bombing raids.[8] The specimen, now labeled CM 9380, is still mounted in Pittsburgh, at first with the tail acting as a tripod in the old-fashioned kangaroo pose. It has since received a modernization of its posture (mounted by Phil Fraley and crew) and can now be found balancing with tail outstretched. Along with a more lifelike posture, the specimen also now includes a composite reconstruction of the skull by Michael Holland.[citation needed] It has been reconstructed in recent years, it measured an estimated 11.9 meters in length and an estimated weight of 7.4–14.6 metric tonnes, 9.1 metric tonnes being the average estimate in that study, although most earlier studies have suggested lower weight figures.[9]
With a length of 12.1-12.2 meters, AMNH 5027 was discovered and excavated in 1908 by Barnum Brown in Montana, and described by Osborn in 1912 and 1916. At the time of discovery, a complete cervical (neck vertebrae) series for Tyrannosaurus was not previously known, so it was this specimen that brought the short, stocky tyrannosaur neck to light. Compared to later specimens (BMNH R7994 and FMNH PR2081, for instance) the cervical series of AMNH 5027 is much more gracile, so with later discoveries the distinction between tyrannosaurid necks and the necks of carnosaurs became more obvious.[10] This specimen also provided the first complete skull of Tyrannosaurus rex. In total, Brown found five partial Tyrannosaurus skeletons. The skeleton of this specimen was used as the iconic symbol for the Jurassic Park film series.
Scale model of the never-completed Tyrannosaurus rex exhibit Osborn planned for the American Museum of Natural History
Osborn planned to mount the similarly sized AMNH 5027 and AMNH 973 together in dynamic poses.[11] Designed by E.S. Christman, the scene was to depict a rearing Tyrannosaurus (AMNH 5027) snapping at another cowering one (AMNH 973), as they fought over the remains of a hadrosaur, described at the time as Trachodon. However, technical difficulties prevented the mount from being executed. One obvious problem was that the Cretaceous Dinosaur Hall was too small to accommodate this dramatic display, and AMNH 5027 was already mounted by itself as the central attraction of the hall. The forearms of Tyrannosaurus were not well documented and the hands were unknown, so for the sake of the display, the forearms of AMNH 5027 were given three fingers, based on the forelimbs of Allosaurus (the more allosaur-like arms were replaced several years later when better fossils of tyrannosaurid arms were found).
The mount retained a rearing pose similar to the initial proposal. By the 1980s it was generally accepted that such a pose would have been anatomically impossible in life, and the skeleton was re-mounted in a more accurate, horizontal pose during a renovation of the museum's dinosaur halls in the early 1990s. The mount can still be seen on display on the fourth floor of the American Museum. The American Museum of Natural History features AMNH 5027 in its famed Hall of Saurischian Dinosaurs to this day.[citation needed]
Nanotyrannus: CMNH 7541
Nanotyrannus holotype
A small but nearly complete skull of Nanotyrannus lancensis, frequently considered to be a juvenile T. rex, was recovered from Montana in 1942. This skull, Cleveland Museum of Natural History (CMNH) 7541, measures 60 centimeters (2.0ft) in length and was originally classified as a species of Gorgosaurus (G. lancensis) by Charles W. Gilmore in 1946.[12] In 1988, the specimen was re-described by Robert T. Bakker, Phil Currie, and Michael Williams, then the curator of paleontology at the Cleveland Museum of Natural History, where the original specimen was housed and is now on display. Their initial research indicated that the skull bones were fused, and that it therefore represented an adult specimen. In light of this, Bakker and colleagues assigned the skull to a new genus, named Nanotyrannus for its apparently small adult size. The specimen is estimated to have been around 5.2 meters (17ft) long when it died.[13] However, a detailed analysis of the specimen by Thomas Carr in 1999 showed that the specimen was, in fact, a juvenile, leading Carr and many other paleontologists to consider it a juvenile specimen of T. rex.[14] The current classification of CMNH 7541 is not universal,[15] with some research suggesting the specimen belongs to a taxon distinct from Tyrannosaurus.[16]
LACM 23844
In 1966, a crew working for the Natural History Museum of Los Angeles County under the direction of Harley Garbani discovered another T. rex (LACM 23844) which included most of the skull of a very large, mature animal. When it was put on display in Los Angeles, LACM 23844 was the largest T. rex skull on exhibit anywhere.[17]
"Black Beauty": RTMP 81.6.1
"Black Beauty"
"Black Beauty" (specimen number RTMP 81.6.1) is a well-preserved fossil of Tyrannosaurus rex. The nickname stems from the apparent shiny dark color of the fossil bones, which occurred during fossilization by the presence of minerals in the surrounding rock;[citation needed] it was the first Tyrannosaurus rex specimen to receive a nickname, beginning a trend that continues with most major T. rex finds.[18] Black Beauty was found in 1980 by a high school student, Jeff Baker, while on a fishing trip with a friend in the region of the Crowsnest Pass, Alberta. A large bone was found in the riverbank and shown to their teacher. Soon afterward, the Royal Tyrrell Museum was contacted,[19] and excavation of the sandstone matrix surrounding the fossils began in 1982.[20] The dig site where the fossil was found is located at approximately near the confluence of the Crowsnest and Willow Rivers, and consisted of rock belonging to the Willow Creek Formation.[18] The specimen is housed in the Royal Tyrrell Museum in Drumheller, Alberta, Canada.[21]
In 2009, a paper by Jack Horner and colleagues illustrated the concept of parasitic infections in dinosaurs by analysing the lesions found on the cranial bones of Black Beauty.[22] The specimen has been used to study comparative morphology between tyrannosaurids and Tyrannosaurus individuals.[citation needed]
Replicas of Black Beauty have been shown in some exhibitions and museums, including both simple skull montages and complete skeletons. Casts are on display in museums around the world.[citation needed]
Stan is the nickname given to a fossil about 11.78 m (38 ft) long found in Hell Creek Formation, South Dakota, close to Buffalo in 1987 by Stan Sacrison, who also discovered the Tyrannosaurus specimen nicknamed "Duffy". The original fossils are now housed at Black Hills Institute of Geological Research, Inc. center. It is a well known specimen, and one of the most complete,[23] with 199 bones recovered.[24] About 30 casts of the original fossil have been sold worldwide, each for a price of about $100,000.[25][26] Stan's skeleton was auctioned for $31.8 million in a 2020 Christie's New York sale, making it a record-breaking dinosaur sale,[27] with the buyer eventually being revealed as the under construction Natural History Museum Abu Dhabi in the Saadiyat Cultural District.[28]
Like many other fossils of Tyrannosaurus rex, the skeleton of Stan shows many broken and healed bones. These include broken ribs and damages in the skull. One of the most prominent injuries are in the neck and the skull. A piece of bone is missing at the rear, and the skull also bears a hole 1 inch wide, probably made by another Tyrannosaurus. Also, two of the cervical vertebrae are fused, and another has additional bone growth. This could have been caused by another Tyrannosaurus bite. The bite marks are healed, indicating that Stan survived the wounds.[29] Stan could also have been infected by Trichomonas-like parasites.[30]
"Wankel Rex": MOR 555
"The nation's T. rex" mount in the Smithsonian Museum
In 1988, local rancher Kathy Wankel discovered another Tyrannosaurus rex in Hell Creek sediments on an island in the Charles M. Russell National Wildlife Refuge of Montana. This specimen was excavated by a team from the Museum of the Rockies led by paleontologist Jack Horner, with assistance from the U.S. Army Corps of Engineers. The specimen, given the number MOR 555 but informally called the "Wankel rex," includes approximately 80-85 percent[31] of the skeleton, including the skull, as well as what at the time was the first complete T. rex forelimb. It has an estimated length of around 11.6 meters (38ft) and a weight between 5.8 metric tons (6.4 short tons) and 10.8 metric tons (11.9 short tons) in newer figures.[32] It is estimated that the "Wankel rex" was 18 years old when it died, an adult but not completely grown. The "Wankel rex" was also one of the first fossil dinosaur skeletons studied to see if biological molecules still existed within the fossilized bones. Doctoral candidate Mary Schweitzer found heme, a biological form of iron that makes up hemoglobin (the red pigment in blood).[33]
It was long on exhibit at the Museum of the Rockies in Bozeman, Montana.[17] In June 2013, the Corps loaned the specimen to the National Museum of Natural History, Smithsonian Institution museum in Washington, D.C., for 50 years. (The Museum of the Rockies continues to display a cast reconstruction of the skull by Michael Holland). The specimen went on temporary display on National Fossil Day, 16 October 2013, and was exhibited until the museum's dinosaur hall exhibit closed for renovation in the spring of 2014. The skeleton, named "The Nation's T. rex"[34] became the centerpiece of the dinosaur hall when it reopened in 2019.[35] Casts of MOR 555 are on display at the National Museum of Scotland,[36] the Australian Fossil and Mineral Museum, and the University of California Museum of Paleontology. A bronze cast of the specimen, known as "Big Mike", stands outside the Museum of the Rockies.[33]
In 2022, Gregory S. Paul and colleagues argued that the Wankel rex was not actually a T. rex, but rather the holotype for a new species: Tyrannosaurus regina.[37] This was heavily criticized by several other leading paleontologists, including Stephen Brusatte, Thomas Carr, Thomas Holtz, David Hone, Jingmai O'Connor, and Lindsay Zanno when they were approached by various media outlets for comment.[38][39][40] Their criticism was subsequently published in a technical paper.[41]
Susan Hendrickson of the Black Hills Institute discovered the best-preserved Tyrannosaurus currently known, in the Hell Creek Formation near Faith, South Dakota, on 12 August 1990.[citation needed] About 90% of the skeleton was recovered,[42] allowing the first complete description of a Tyrannosaurus rex skeleton.[43] This specimen, named "Sue" in honor of its discoverer, soon became embroiled in a legal battle over its ownership. The owner of the land the fossil was found, Maurice Williams, as well the Sioux Tribe he belonged to, claimed ownership, the Institute had considered itself to have. In 1997, the suit was settled in favor of Williams and the fossil was returned to Williams' ownership. Williams quickly offered up "Sue" for auction by Sotheby's in New York, where it was sold to the Field Museum of Natural History in Chicago for US$8.4 million—the highest price ever paid for a fossil before being surpassed by Stan.[citation needed]
Sue has a length of 12.3–12.4 meters (40.4–40.7ft), stands 3.66–3.96m (12–13ft) tall at the hips, and according to the most recent studies estimated to have weighed between 8.4 and 14 metric tons when alive.[32][44][45][46][47][48][excessive citations] It has been hypothesised that Sue's impressive size may have been achieved due to a prolonged ontogenic development, since it is the third oldest Tyrannosaurus known. Sue's age at the time of death was estimated by Peter Mackovicky and the University of Florida to be 28 years old, over 6–10 years older than most big Tyrannosaurus specimens, like MOR 555, AMNH 5027 or BHI 3033.[49] The only known specimen of T. rex that is older than Sue is Trix.[50][51]
Preparation of "Sue" (FMNH PR2081) was completed at the Field Museum and the skeleton was placed on exhibit on 17 May 2000.[52][53]
"Montana's T. rex": MOR980
Montana's T. rex (formerly nicknamed "Peck's Rex) real fossils displayed at the Museum of the Rockies
Montana's T. rex (also known as "Peck's rex", "Peckrex", "Rigby's rex" and Tyrannosaurus "imperator") is the nickname given to a fossil specimen found in Montana in 1997.[54] The discovery was made by Louis E. Tremblay on 4 July 1997 working under the supervision of J. Keith Rigby Jr. who led the excavation and bone preparation.
The skeleton of Montana's T. rex includes a relatively complete skull with jaws, multiple vertebrae of the back and tail, a well preserved gastralium, and hipbone with complete ischium and pubis. The left hindleg is relatively complete with a 1.2-meter-long (3.9ft) femur, missing only some toe bones. The forelimbs include the scapula and furcula, both humeri and right hand phalanges, as well as metacarpal III. Montana's T. rex has been the subject of research regarding parasitic infections in dinosaurs.[55] The forelimbs of Montana's T. rex have also been studied as they show evidence of use. This evidence includes the construction of metacarpal III, as well as repeated fractures in the furcula—possibly caused by heavy loads or pressure (Carpenter and Lipkin, 2005).[56]
The fossils of Montana's T. rex are exhibited at Museum of the Rockies as part of a full skeletal mount completed with cast elements replacing the missing bones.[57] This mount was installed after the Wankel Rex (now nicknamed The Nation's T. rex) was loaned to the Smithsonian to occupy a central part in the museum's dinosaur hall, featuring a dynamic mount showing the apex predator devouring a Triceratops carcass.[58][59] It has been stated by Pete Makovicky, the Chicago museum's lead curator of dinosaurs, that this specimen is in the same size range as "Sue" and "Scotty".[60]
"Bucky": TCM 2001.90.1
Cast of the "Bucky" specimen
Bucky is a fossil of a juvenile specimen on display at The Children's Museum of Indianapolis in Indianapolis, Indiana. It is the first juvenile Tyrannosaurus ever placed on permanent exhibit in a museum.[61] The dinosaur remains were found in 1998 in the Hell Creek Formation near the town of Faith, South Dakota. The skeleton, transported by water, ended up in a low shallow valley along with bones from an Edmontosaurus and Triceratops. It was discovered by rancher and cowboy Bucky Derflinger.[62] Bucky was well preserved and easily prepared by the Black Hills Institute in South Dakota.[63] Part of The Children's Museum of Indianapolis Dinosphere exhibit,[64] Bucky is displayed along with Stan, an adult Tyrannosaurus, in a hunting scene. Both dinosaurs are attacking a Triceratops specimen known as "Kelsey".[65]
Bucky is one of the few dinosaur fossils found with a furcula;[66] Bucky's furcula was the first one found for the genus Tyrannosaurus.[67] Bucky also has a nearly complete set of gastralia, or belly ribs, and an ulna, or lower arm bone. As of now, 101 bones, or about 34% of Bucky's skeleton, has been discovered and verified.[62] Bucky is the sixth-most complete Tyrannosaurus rex out of more than 40 that have been discovered.[61]
"E. D. Cope": BHI 6248
E. D. Cope (named after the paleontologist of the same name) is a Tyrannosaurus specimen discovered in South Dakota by Bucky Derflinger in 1999 at the same site as AMNH 3982. Excavations of this 10% complete skeleton began in 2000. The known material includes a partial skull, several vertebrae, and ribs.[18] A very wide femur with a length of 1300 mm and a circumference of 630 mm is also known.[37]
Jane is a fossil specimen of small tyrannosauriddinosaur, officially known as BMRP 2002.4.1, discovered in the Hell Creek Formation in southern Montana in 2001.[68] Despite having a typically female name, Jane's sex is unknown—the specimen was named after Burpee Museum benefactor Jane Solem. The specimen was found in the summer of 2001 by Carol Tuck and Bill Harrison on an expedition led by Burpee Museum curator Michael Henderson.[69] After four years of preparation, Jane was put on display at Rockford, Illinois' Burpee Museum of Natural History as the centerpiece of an exhibit called "Jane: Diary of a Dinosaur."[citation needed]
The Jane specimen has been central to the debate regarding the validity of the proposed tyrannosaurid genus Nanotyrannus. However, the Jane material has yet to be properly studied and described by scientists. Although Larson (2013) saw Jane as more identical to CMNH 7541 and LACM 28471 than to adult T. rex in having a higher tooth count, large pneumatic foramen on the center of the quadratojugal, T-shaped postorbital, and fused shoulder blade and pelvis,[70] Yun (2015) concurred with the opinion of most workers that Nanotyrannus is a juvenile T. rex, noting that a juvenile specimen of Tarbosaurus described by Tsuihiji et al. (2011) also has a T-shaped postorbital.[71] Paleontologists who support the theory that Jane represents a juvenile believe the tyrannosaur was approximately 11 years old at its time of death, and its fully restored skeleton measured 6.5 meters (21ft) long, a bit more than half as long as the largest-known complete T. rex specimen, nicknamed "Sue," which measures 12.4m (41ft) long. According to Hutchinson et al. (2011), the weight of the Jane specimen in life was probably between 639kg (1,409lb) and 1,269kg (2,798lb), 954kg (2,103lb) being the average estimate.[32]
"B-rex": MOR 1125
Reconstructed skull of B-rex
This specimen was found in the lower portion of the Hell Creek Formation near Fort Peck Lake in the Charles M. Russell National Wildlife Refuge in Garfield County, Montana. Its discoverer was Bob Harmon, a fossil preparator for the Museum of the Rockies, and was nicknamed the "B-rex" (or "Bob-rex") in honor of Harmon. The specimen was discovered in 2000, and excavated by MOR from 2001 to 2003.[62] Although only 37 percent of the skeleton was present, this included almost all of the skull (although the skull was nearly completely disarticulated). The specimen also includes several cervical, dorsal, sacral, and caudal vertebrae; several chevrons; some cervical and dorsal ribs; left scapula and coracoid; the furcula; the left ulna; both femora, tibiae, and ulnae; the right calcaneum; right astragalus; and a number of pesphalanges.[62]
Femur of MOR 1125 from which demineralized matrix and peptides (insets) were obtained
In the March 2005 Science magazine, Mary Higby Schweitzer of North Carolina State University and colleagues announced the recovery of soft tissue from the marrow cavity of a fossilized femur belong to B-Rex. Flexible, bifurcating blood vessels and fibrous but elastic bone matrix tissue were recognized. In addition, microstructures resembling blood cells were found inside the matrix and vessels. The structures bear resemblance to ostrich blood cells and vessels. However, since an unknown process distinct from normal fossilization seems to have preserved the material, the researchers are being careful not to claim that it is original material from the dinosaur.[72] Paleontologist Thomas Kaye of the University of Washington in Seattle hypothesized that the soft-tissue is permineralized biofilm created by bacteria while digesting and breaking down the original specimen. He has discovered this to be true in many specimens from the same area.[73] In 2016, it was finally confirmed by Mary Higby Schweitzer and Lindsay Zanno et al that the soft tissue was medullary bone tissue, like that in modern birds when they are readying to lay eggs. This confirmed the identity of the Tyrannosaurus MOR 1125 as a female.[74]
"Samson"
Samson
A T. rex specimen was discovered on private land in Harding County, South Dakota, once in 1981 by Michael and Dee Zimmerschied, and again on 4 October 1992 (Alan and Robert Detrich re-discovered Samson after it was originally found and deemed by paleontologists that several bones had washed in and there was nothing left). It was shortly after that when Fred Nuss and Candace Nuss of Nuss Fossils with the Detrich brothers found the most complete and undistorted Tyrannosaurus rex skull ever discovered.[75] Following the sale of "Sue," another Tyrannosaurus rex skeleton was, the specimen was put up for auction on eBay in 2000 under the name of "Z-rex", with an asking price of over US$8 million. It failed to sell online but was purchased for an undisclosed price in 2001 by British millionaire Graham Ferguson Lacey, who renamed the skeleton "Samson" after the Biblicalfigure of the same name.[citation needed] It was prepared by the Carnegie Museum starting in May 2004.[75] After preparation was complete in March 2006, the specimen was returned to Lacey.[76] It, along with some other dinosaur skeletons, was sold again at auction on 3 October 2009.[citation needed]
Samson measured 11.9m (39ft), only slightly shorter than Sue.[77]
"Baby Bob"
On 7 July 2013, fossil hunter Robert Detrich of Wichita, Kansas, unearthed the remains of what is believed to be a 4-year-old Tyrannosaurus rex. Detrich unearthed the fossil dubbed "Baby Bob" in a fossil-rich area near the Eastern Montana town of Jordan.[78][self-published source?] Its femur measures about 25 inches, and if all the preliminary data pans out, that would make it among the smallest T. rex specimens ever found. Baby Bob has been fully excavated, although it will take another year to clean. Detrich said the skull, which is about 75 percent complete, and most of the major skeletal elements were found strewn across a flood plain, although very few vertebra and ribs were found.
Scotty is currently thought by many to be the largest Tyrannosaurus specimen ever discovered.
"Scotty", cataloged as RSM P2523.8, was discovered in Saskatchewan, Canada in 1991. Since its discovery and extensive subsequent study, "Scotty" has been referred to as the largest T. rex ever discovered in the world, the largest of any dinosaur discovered in Canada, and as one of the oldest and most complete fossils of its kind at more than 70% bulk.[79] "Scotty" resides at the Royal Saskatchewan Museum's T. rex Discovery Centre in Eastend, SK, Canada. In May 2019, a second mount was erected at the Royal Saskatchewan Museum in Regina, where the exhibit reflects the recent discoveries about the fossil.[80]
"Scotty" was discovered by Robert Gebhardt, a high school principal from Eastend, SK who accompanied palaeontologists from the Royal Saskatchewan Museum on a prospective expedition into the Frenchman Formation in southwestern Saskatchewan on 16 August 1991.[79] It wasn't until June 1994 that the Royal Saskatchewan Museum was able to begin the excavation, which was led and overseen by the Museum's Ron Borden, as well as resident paleontologists Tim Tokaryk and John Storer who were with Gebhardt when he uncovered the first fossils.[80][81] The bones were deeply packed in dense, iron-laden sandstone, which took more than twenty years for the team to fully remove, excavate, and assemble the majority of the skeleton, with additional trips being made to the site to retrieve smaller bones and teeth.[81][82][83][84] The entire process of excavating the skeleton was also slowed down by its considerable size.[79]
"Scotty" is reported to be 13.1 metres (43 feet) long and weighed an estimated 8.8 tonnes (8,800kg).[83][85][86] Despite it not being a complete fossil, paleontologists were able to create the estimation for the weight and length through measurements of important weight bearing bones such as the femur, hip, and shoulder bones that have all been measured to be larger and thicker with "Scotty" than the corresponding bones with "Sue". Going from the latest study "Scotty" exceeds "Sue" in 84.6% of the published measurements.[87] While the reported measurements and weight for "Scotty" are larger than those of "Sue", some scientists posit that the two fossils are too close in size to officially declare "Scotty" the largest.
Like other T. rex fossils, "Scotty" shows signs of trichomoniasis, a parasitic infection in the jaw that left visible holes in the bone and was unique to this specific species of dinosaur.[85][88] Additionally, a broken and healed rib on its right side, broken tail vertebra, as well as a hole near the eye socket are possibly the result of another T. rex attack.[79][85] Other abnormalities, such as impacted teeth, suggest that "Scotty" was not only bitten, but also bit other animals.[83]
"Tristan"
Tristan in Berlin
Commercial paleontologist Craig Pfister discovered the specimen in the lower Hell Creek Formation in Carter County, Montana, in 2010.[89][90] Its excavation and preservation lasted four years.[90] It was later sold to Danish-born investment banker Niels Nielsen, who loaned the specimen to the Museum für Naturkunde in Berlin, Germany, for research and exhibition.[91] It has been on display at Museum für Naturkunde between 2015 and 2020,[90] moving to the Natural History Museum of Denmark for one year, and expected back in 2021.[92][93] Nielsen and his friend Jens Jensen named the specimen Tristan-Otto (short: Tristan) for their sons.[94] The Museum für Naturkunde Berlin lists it under specimen number MB.R.91216.[95] Several European museums have Tyrannosaurus casts (replicas) or parts, but Tristan is one of only two original skeletons on display in the continent (the other is "Trix" in the Netherlands).[89] The matte-black fossilised skeleton is about 12m (39ft) long and 3.4m (11ft) tall at the hips.[96][97] Tristan is among the most complete known Tyrannosaurus skeletons: It was re-assembled from about 300 separate parts, 170 of which are original (including 98% of the skull and all the teeth), the rest reproductions.[90][91][96] It is estimated to have died when about 20 years old and it was in poor health, having several bone fractures, bite marks to the skull and signs of disease in the jaw.[89][96] The disease present in Tristan's jaw was suggested to be a case of tumefactive osteomyelitis.[98]
"Thomas"
Thomas, mounted with two younger specimens
From 2003 to 2005, Thomas was excavated by NHMLA paleontologists in southeastern Montana. At 17 years old, 34ft (10m) long and nearly 7,000lb (3,200kg), it is estimated to be a 70% complete specimen. Thomas is mounted in a "growth series" with the youngest-known Tyrannosaurus rex fossil, a two-year-old, 11-foot-long (3.4m) specimen, and a 13-year-old, 20-foot-long (6.1m) juvenile specimen.[99]
Victoria is a specimen found near Faith, South Dakota in 2013.[101] Victoria is estimated to be around 12ft tall and 40ft long, and she is thought to have died in her subadult stage, between 15 and 25 years of age.[102] Victoria has also been the subject of a traveling exhibition being displayed in places such as the Arizona Science Center.[103][104] Her cause of death is unknown; however, she was believed to have been bitten in the lower jaw by another Tyrannosaurus. The bite may have become infected, spreading and leading to sepsis.[101]
"Ivan"
Ivan the T. rex, during installation
Ivan is a 65% complete T. rex displayed at the Museum of World Treasures in Wichita, Kansas. The specimen has the most complete tail of any T. rex, only missing around 3 vertebrae. Ivan is around 40ft long and 12 feet high.[105]
In 2013, a team of paleontologists from the Naturalis Biodiversity Center (Leiden, Netherlands) traveled to Montana where they discovered and unearthed a large and remarkably complete Tyrannosaurus rex specimen that lived 67 million years ago.[106]Black Hills institute collaborated with the team in the excavation. The bones were cleaned and assembled in a mount at Black Hills Institute's installations, with the help of both Chicago's Field Museum of Natural History and the Naturalis Museum in Leiden. Chicago's Field museum sent digital models of their famous specimen, FMNH PR 2081 (Sue) to complete the cast and Naturalis museum replicated the bones using 3D-printing technology.
The specimen was named Trix after former Netherlands' Queen Beatrix. At arrival in the Netherlands, it started touring on public display in an itinerant exhibition titled T. rex in Town. The first exhibit spanned from 10 September 2016 to 5 June 2017 and was set at the only room of the Naturalis museum open to public at the time (the 17th-century building known as Pesthuis), due to the fact that the museum was undergoing restoration. When the Netherlands exhibition ended, it continued travelling through other European countries in 2017, 2018 and 2019. As of August 2019, Trix was returned to display at the Naturalis museum where it is installed in a special room that was under construction during Trix's European tour.[107]
According to Peter Larson (director of Black Hills Institute), Trix is among the most complete Tyrannosaurus found. Between 75% and 80% of its skeletal volume was recovered.[108][109] They are thought to have been at least 30 years old at death.[50][51]
Cast of T.rex skeleton incorporating authentic fossil elements from "Titus", exhibited at Wollaton Hall, Nottingham
"Titus" is the name given to an obsidian-black skeleton of a Tyrannosaurus rex discovered in Montana's Hell Creek Formation in 2014 and excavated in 2018. It is 20% complete, and was named after the protagonist in Shakespeare's Titus Andronicus. Exhibited in the Nottingham Natural History Museum for 13 months beginning July 2021, it was during that time only the second specimen of Tyrannosaurus to be on exhibit in United Kingdom, the other being the type of the junior synonym Dynamosaurus imperosus which has the jaw on display in the Natural History Museum, London.[110] External bone inspection has revealed injuries to Titus' right tibia (possibly a claw or bite wound); a deformed toe on the right foot; and a bitten and healed tail. The bite wound near the end of the tail indicates a possible attack by another Tyrannosaurus rex.[111]
The remains of "Titus" were discovered in September 2014 by commercial paleontologist Craig Pfister near Ekalaka, Carter County, Montana.[110] Excavation of the specimen began in 2018, and took 18 months.[112] The bones of "Titus" were shipped to conservationist Nigel Larkin in the UK, who constructed the mount using a cast of the Tyrannosaurus specimen Stan to supplement the known bones of "Titus", after scanning the bones using photogrammetry to create digital models that were 3D printed for use in the exhibition, alongside the display of the real fossil skeleton, and which remained at the museum after the end of the exhibition.[113][114] For the exhibit at the Nottingham Natural History Museum at Wollaton Hall, Titus was reconstructed in a walking pose.[110]
"Tufts-Love": UWBM 99000
Tufts-Love T. rex
In 2016 Greg Wilson, David DeMar, and a paleontology team from the Burke Museum of Natural History and Culture, the University of Washington, and the Dig Field school excavated the partial remains of a Tyrannosaurus rex from Montana. The partial skeleton was found by two Burke Museum volunteers, Jason Love and Luke Tufts, and was named the "Tufts-Love" rex.[citation needed] Paleontologists at the Burke Museum believe that the Tufts-Love rex was around 15 years old when it died. The skull is of average size for an adult T. rex. The specimen was found in Late Cretaceous deposits and it is estimated to be 66.3 million years old.[115] The Tufts-Love rex is undergoing preparation by Michael Holland and his team at the Burke Museum. The skeleton is estimated to be 30% complete, but it includes a complete (all of the bones of the skull and jaws are preserved) and mostly articulated skull. Holland describes the skull as minimally distorted and in an "exquisite" state of preservation.[116]
"Peter": AWMM-IL 2022.9
Peter (L) & Barbara (R) side by side 2022
Peter is the nickname given to a specimen on loan to the Auckland War Memorial Museum by an anonymous owner, currently on display alongside “Barbara” until the end of 2023.[117][118] The specimen is estimated to be 66.8 million years old and almost of adult size. He was recovered from Niobrara County, Wyoming.
“Peter” is one of only four incredibly rare and visually stunning obisidian black colored tyrannosaurus rex.
He was likely killed by exocannibalism[119] as entire sections of bone were damaged, and some were split open by huge crushing bite-forces. The nature of the crushing on the femur and tibia, along with size of the bite marks, indicates that these bones were bitten through by another Tyrannosaurus rex. There is also a set of smaller, parallel tooth marks nearby on the shaft that are not attributable to an adult T. rex. Explanations for this behavior range from response to over-crowded populations, limited food supply, sexual dominance, or even play.[120]
"Barbara": AWMM-IL 2022.21
Peter (Black, Left) & Barbara (Right, Beige) Pose at Auckland Museum 2022
Barbara is the nickname given to a specimen on loan to the Auckland War Memorial Museum by the same anonymous owner as "Peter", currently on display alongside him until the end of 2023.[121] The pair will be the first adult male and female T. rex to be displayed together. She is one of a few specimens believed to be pregnant.[122]
"Barbara's" circumstances are particularly rare, more so when taken into account that she suffered and survived long after a debilitating foot injury. While she was no longer able to capture her prey, it is suggested that she got by with the help of a mate or cohort feeding her, as the injury would have rendered her immobile for upwards of 6 months. This is vaguely supported by trackway evidence that has been used to imply tyrannosaur group hunting. It is doubtful that "Barbara" ever successfully hunted again as a predator unless its prey was nearby.[122]
"Bloody Mary" (as named by Pete Larson)[123] is a nickname for a nearly complete tyrannosaurid (generally considered an adolescent T. rex or a "Nanotyrannus") at the North Carolina Museum of Natural Sciences. This specimen, considered the most complete of any Tyrannosaurus specimen at more than 98% preservation,[124] was discovered in Montana in 2006. Following years of unsuccessful attempts to sell it to museums or auction it off, the NCMNS started negotiations in 2016, which were prolonged due to a legal battle over the rights to the fossil, which was resolved in 2020. The fossils were officially displayed in 2024.[125]
The Dueling Dinosaurs T. rex is an adolescent that was preserved entangled with a Triceratops; their combined fossil is referred to as the "Dueling Dinosaurs". Given the injuries present on both fossils, it has been theorized that both died while fighting one another. The specimen is presently undergoing study. Important biological data is likely preserved within the specimen, including body outlines, skin impressions, soft tissues, injuries, stomach contents, and even original proteins.[126][127]
Tyrannosaurus mcraeensis: NMMNH P-3698
Reconstructed skull of T. mcraeensis indicating the holotype material in white
The remains of a tyrannosaur were discovered in 1983 in the Campanian-early Maastrichtian Hall Lake Formation in New Mexico by Donald Staton and Joe LaPoint. Reposited at the New Mexico Museum of Natural History and Science, the fossil material (NMMNH P-3698) consists of skull and lower jaw bones, in addition to isolated teeth and chevrons. Some of the bones were briefly mentioned in 1984 as belonging to T. rex,[128] and were described in 1986.[129] In 2024, Sebastian G. Dalman and colleagues described this specimen as the holotype of a new Tyrannosaurus species, T. mcraeensis. This species differs from T. rex in having smaller postorbital crests, a proportionately longer and shallower lower jaw with a less prominent chin suggestive of a weaker bite, and more laterally compressed teeth.[2]
Albertosaurus is a genus of large tyrannosaurid theropod dinosaur that lived in northwestern North America during the early to middle Maastrichtian age of the Late Cretaceous period, about 71 million years ago. The type species, A. sarcophagus, was apparently restricted in range to the modern-day Canadian province of Alberta, after which the genus is named, although an indeterminate species has been discovered in the Corral de Enmedio and Packard Formations of Mexico. Scientists disagree on the content of the genus and some recognize Gorgosaurus libratus as a second species.
Tyrannosaurus is a genus of large theropod dinosaur. The type species Tyrannosaurus rex, often shortened to T. rex or colloquially T-Rex, is one of the best represented theropods. It lived throughout what is now western North America, on what was then an island continent known as Laramidia. Tyrannosaurus had a much wider range than other tyrannosaurids. Fossils are found in a variety of rock formations dating to the latest Campanian-Maastrichtian ages of the late Cretaceous period, 72.7 to 66 million years ago. It was the last known member of the tyrannosaurids and among the last non-avian dinosaurs to exist before the Cretaceous–Paleogene extinction event.
Tyrannosauridae is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to fifteen genera, including the eponymous Tyrannosaurus. The exact number of genera is controversial, with some experts recognizing as few as three. All of these animals lived near the end of the Cretaceous Period and their fossils have been found only in North America and Asia.
Tarbosaurus is a genus of large tyrannosaurid dinosaur that lived in Asia during the Late Cretaceous epoch, about 70 million years ago. It contains the single type species: Tarbosaurus bataar, which is known from the Nemegt Formation of Mongolia, with more fragmentary remains found further afield in the Subashi Formation of China. Tarbosaurus is represented by dozens of fossil specimens, including several complete skulls and skeletons. These remains have allowed studies focusing on its phylogeny, skull mechanics, and brain structure. Further fossil remains have been reported from other geologic formations of Asia, however, these remains are fragmentary and can not be confidentially assigned to Tarbosaurus or the type species.
Daspletosaurus is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 78 and 74.4 million years ago, during the Late Cretaceous Period. The genus Daspletosaurus contains three named species. Fossils of the earlier type species, D. torosus, have been found in Alberta, while fossils of a later species, D. horneri, have been found only in Montana. D. wilsoni has been suggested as an intermediate species between D. torosus and D. horneri that evolved through anagenesis, but this theory has been disputed by other researchers.
Gorgosaurus is a genus of tyrannosaurid theropod dinosaur that lived in western North America during the Late Cretaceous Period (Campanian), between about 76.5 and 75 million years ago. Fossil remains have been found in the Canadian province of Alberta and the U.S. state of Montana. Paleontologists recognize only the type species, G. libratus, although other species have been erroneously referred to the genus.
Aublysodon is a genus of carnivorous dinosaurs known only from the Judith River Formation in Montana, which has been dated to the late Campanian age of the late Cretaceous period. The only currently recognized species, Aublysodon mirandus, was named by paleontologist Joseph Leidy in 1868. It is sometimes considered dubious now, because the type specimen consists only of an isolated premaxillary (front) tooth. Although this specimen is now lost, similar teeth have been found in many US states, western Canada, and Asia. These teeth almost certainly belong to juvenile tyrannosaurine tyrannosaurids, but most have not been identified to species level. However, it is likely that the type tooth belongs to one of the species in the genus Daspletosaurus, which was present in contemporary formations, and which matches specific details of the original tooth. The synapomorphies alleged to distinguish the Aublysodontinae, especially lack of serrations on premaxillary teeth could have been caused by tooth wear in life, postmortem abrasion, or digestion. Most other "aublysodontine"-type teeth may be from ontogenetic stages or sexual morphs of other tyrannosaurids.
Alioramus is a genus of tyrannosaurid theropod dinosaurs from the Late Cretaceous period of Asia. It currently contains two species. The type species, A. remotus is known from a partial skull and three foot bones recovered from the Mongolian Nemegt Formation, which was deposited in a humid floodplain about 70 million years ago. These remains were named and described by Soviet paleontologist Sergei Kurzanov in 1976. A second species, A. altai, known from a much more complete skeleton also from the Nemegt Formation, was named and described by Stephen L. Brusatte and colleagues in 2009. Its relationships to other tyrannosaurid genera were at first unclear, with some evidence supporting a hypothesis that Alioramus was closely related to the contemporary species Tarbosaurus bataar. However, the discovery of Qianzhousaurus indicates that it belongs to a distinct branch of tyrannosaurs, namely the tribe Alioramini.
The Burpee Museum of Natural History is located along the Rock River in downtown Rockford, Illinois, United States, at 737 North Main Street.
Charles Whitney Gilmore was an American paleontologist who gained renown in the early 20th century for his work on vertebrate fossils during his career at the United States National Museum. Gilmore named many dinosaurs in North America and Mongolia, including the Cretaceous sauropod Alamosaurus, Alectrosaurus, Archaeornithomimus, Bactrosaurus, Brachyceratops, Chirostenotes, Mongolosaurus, Parrosaurus, Pinacosaurus, Styracosaurus ovatus and Thescelosaurus.
Tyrannosaurinae is one of the two extinct subfamilies of Tyrannosauridae, a family of coelurosaurian theropods that consists of at least three tribes and several genera. All fossils of these genera have been found in the Late Cretaceous deposits of western North America and east Asia. Compared to the related subfamily Albertosaurinae, tyrannosaurines overall are more robust and larger though the alioramins were gracile by comparison. This subfamily also includes Lythronax, one of the oldest known tyrannosaurid genera, as well as the youngest and most famous member of the group, Tyrannosaurus rex.
Sue is the nickname given to FMNH PR 2081, which is one of the largest, most extensive, and best preserved Tyrannosaurus rex specimens ever found, at over 90 percent recovered by bulk. FMNH PR 2081 was discovered on August 12, 1990, by American explorer and fossil collector Sue Hendrickson, and was named after her.
"Stan", also known by its inventory number BHI 3033, is a Tyrannosaurus rex fossil found in the Hell Creek Formation in South Dakota, just outside of Buffalo in 1987, and excavated in 1992. It is the fifth most complete T. rex fossil discovered to date, at more than 70% bulk. In October 2020, the fossil was sold for $31.8 million at auction, making it at the time the most expensive dinosaur specimen and fossil ever sold. This record stood until July 2024, when the Stegosaurus fossil Apex sold at auction for $44.6 million. In March 2022, Abu Dhabi's Department of Culture and Tourism stated that they had acquired Stan and were planning on displaying the fossil at a new museum of natural history scheduled to open in 2025.
Lythronax is a genus of tyrannosaurid dinosaur that lived in North America around 81.9-81.5 million years ago during the Late Cretaceous period. The only known specimen was discovered in Utah in the Wahweap Formation of the Grand Staircase–Escalante National Monument in 2009, and it consists of a partial skull and skeleton. In 2013, it became the basis of the new genus and species Lythronax argestes; the generic name Lythronax means "gore king", and the specific name argestes originates from the Greek poet Homer's name for the wind from the southwest, in reference to the specimen's geographic provenance in North America.
This timeline of tyrannosaur research is a chronological listing of events in the history of paleontology focused on the tyrannosaurs, a group of predatory theropod dinosaurs that began as small, long-armed bird-like creatures with elaborate cranial ornamentation but achieved apex predator status during the Late Cretaceous as their arms shrank and body size expanded. Although formally trained scientists did not begin to study tyrannosaur fossils until the mid-19th century, these remains may have been discovered by Native Americans and interpreted through a mythological lens. The Montana Crow tradition about thunder birds with two claws on their feet may have been inspired by isolated tyrannosaurid forelimbs found locally. Other legends possibly inspired by tyrannosaur remains include Cheyenne stories about a mythical creature called the Ahke, and Delaware stories about smoking the bones of ancient monsters to have wishes granted.
Trix is a Tyrannosaurus rex specimen excavated in 2013 in Montana, United States by a team of paleontologists from the Naturalis Biodiversity Center in Leiden, the Netherlands and Black Hills institute of Geological Research in South Dakota. This Tyrannosaurus, over thirty years old – the oldest known Tyrannosaurus specimen – lived about 67 million years ago. It is considered to be the third most complete Tyrannosaurus found, with between 78% and 80% of its bone volume recovered. The specimen was named Trix after the former Queen Beatrix of the Netherlands. It is one of only two Tyrannosaurus specimens on permanent exhibit in mainland Europe. The other one is a specimen named Tristan on exhibit at the Natural History Museum of Berlin.
The Dueling Dinosaurs or Montana Dueling Dinosaurs is a fossil specimen originating from the Hell Creek Formation of Montana. It consists of the fossilized skeletons of a tyrannosaur and a Triceratops horridus entangled with one another and entombed in sandstone. This is identical to the Fighting Dinosaurs, an 80-million-year-old specimen found in 1971 Mongolia where a Velociraptor and Protoceratops were locked in battle and preserved. The "dueling" inference comes from the numerous injuries sustained by both dinosaurs, including a tooth from the tyrannosaur embedded within the Triceratops, although it is not known whether they were actually buried fighting one another. Tyrannosaurus rex was a 40 ft. long carnivore weighing 6 tons and ruled the Cretaceous period with strong jaws and sharp teeth to tear up to 500 pounds of flesh and bone in a single bite. Triceratops was bigger than an elephant and was notified by a rounded skull with horns and a bony frill to intimidate and defend against predators. Despite the scientific importance of the specimen, it remained obscure for decades due to a lengthy legal dispute over property rights to the specimen, which has since been resolved. The fossil is in the possession of and being studied by the North Carolina Museum of Natural Sciences, where it went on display in 2024.
Titus is an obsidian black skeleton of a Tyrannosaurus rex discovered in the Hell Creek Formation, Montana, United States in 2014 and excavated in 2018.
Scotty is the nickname for the Tyrannosaurus rex fossil, catalogued as RSM P2523.8, that was discovered in Saskatchewan, Canada in 1991. The fossilised remains were painstakingly removed, almost completely by hand, over two decades from the rock in which they were embedded. When the preparation was complete in 2011, a ~65% complete T. rex skeleton was revealed.
Victoria is a specimen of the species Tyrannosaurus rex recognized for its well-preserved and nearly complete skeleton, making it the second most complete T. rex finds in recent history. Victoria got her name in the lab in Victoria, British Columbia, where it was studied and restored.
References
↑ Larkin, N.R., Dey, S., Smith, A.S. and Evans, R. 2022. 21st Century Rex: maximising access to a privately owned Tyrannosaurus rex skeleton in the digital age. Geological Curator, 11(6), 341–354.
1 2 Osborn, H.F. (1905). "Tyrannosaurus and other Cretaceous carnivorous dinosaurs". Bulletin of the American Museum of Natural History. 21 (3): 259–265. doi:10.5281/zenodo.1038151. hdl:2246/1464.
↑ Osborn, Henry Fairfield; Brown, Barnum (1906). "Tyrannosaurus, Upper Cretaceous carnivorous dinosaur". Bulletin of the American Museum of Natural History. 22. hdl:2246/1473.
↑ Brown, B. (1915). "Tyrannosaurus, a Cretaceous carnivorous dinosaur, the largest flesh-eater that ever lived". Scientific American. 63 (15): 322–323. doi:10.1038/scientificamerican10091915-322.
↑ Osborn, H.F. (1913). "Tyrannosaurus: restoration and model of the skeleton". Bulletin of the American Museum of Natural History. 32: 91–92. hdl:2246/1735.
↑ Gilmore, C. W. (1946). "A new carnivorous dinosaur from the Lance Formation of Montana". Smithsonian Miscellaneous Collections. 106: 1–19.
↑ Bakker, R.T.; Williams, M.; Currie, P.J. (1988). "Nanotyrannus, a new genus of pygmy tyrannosaur, from the latest Cretaceous of Montana". Hunteria. 1: 1–30.
1 2 Horner, J.R. & Lessem, D. 1993. The Complete T. rex New York: Simon & Schuster. 238pp.
1 2 3 Larson, Peter H. (2008). "One Hundred Years of Tyrannosaurus Rex: The Skeletons.". In Larson, Neal L. (ed.). Tyrannosaurus rex the Tyrant King. pp.16–17.
↑ Brochu, C.R. (2003). "Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull". Memoirs of the Society of Vertebrate Paleontology. 7: 1–138. doi:10.2307/3889334. JSTOR3889334.
1 2 Crosslin, Rick; Mary Fortney; Dinosphere Exhibit Development Team (2004). "Resource Materials: Bucky-Tyrannosaurus rex". Dinosphere: A 3–5 Unit of Study.
↑ Larson, Neal L.; Peter Larson; Kenneth Carpenter (2008). "One Hundred Years of Tyrannosaurus Rex: The Skeletons". Tyrannosaurus Rex, the Tyrant King: 43–44.
↑ Crosslin, Rick; Mary Fortney; Dinosphere Exhibit Development Team (2004). "Tyrannosaurus rex Attack Scene-What will be the outcome?". Dinosphere: A 3–5 Unit of Study: 88.
↑ Larson, P. L. 2013. The case for Nanotyrannus in J. M. Parrish, R. A. Molnar, P. J. Currie., and E. B. Koppelhus (eds.) Tyrannosaurid Paleobiology, University of Indiana Press, Bloomington and Indianapolis, pp. 15–53.
↑ Burnham, M.; Mclain, M.; Nelson, D.R.; Snyder, K. (2018). "Tyrannosaur cannibalism: a case of a tooth-traced tyrannosaurid bone in the Lance Formation (Maastrichtian), Wyoming". PALAIOS. 33 (4): 164–173. Bibcode:2018Palai..33..164M. doi:10.2110/palo.2017.076. S2CID134871802.
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